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HARVARD UNIVERSITY 

Library of the 

Museum of 

Comparative Zoology 



Bulletin of the Museum of Comparative Zoology 

AT HARVARD COLLEGE 

Vol. LXXXIII 




A CHECKLIST OF AFRICAN MAMMALS 



By 

Glover M, Allen 



CAMBRIDGE, MASS., U. S. A. 

PRINTED FOR THE MUSEUM 

February, 1939 



PUBLICATIONS 
OF THE 

MUSEUM OF COMPARATIVE ZOOLOGY 
AT HARVARD COLLEGE 



There have been published of the Bulletin, Vols. I to 
LXXXIII, and LXXXV, No. 1 & 2; of the Memoirs, Vol. I to 
LIV, No. 1, 2 & 3, and Vol. LV. 

The Bulletin and Memoirs are devoted to the publication of 
original work by the Officers of the Museum, of investigations 
carried on by students arid others in the different Laboratories 
of Natural History, and of work by specialists based upon the 
Museum Collections and Exploration. 

These publications are issued in numbers at irregular intervals. 
Each number of the Bulletin and of the Memoirs is sold separately. 
A price list of the publications of the Museum will be sent on 
application to the Director of the Museum of Comparative 
Zoology, Cambridge, Massachusetts. 



Bulletin of the Museum of Comparative Zoology 

AT HARVARD COLLEGE 

Vol. LXXXIII 



A CHECKLIST OF AFRICAN MAMMALS 



By 

Glover M. Allen 



CAMBRIDGE, MASS., U.S.A. 

PRINTED FOR THE MUSEUM 

February, 1939 




To the memory of 
JOHN CHARLES PHILLIPS, M.D. 

Sportsman, Naturalist, Author, Conservationist 

His far-sighted work for the protection of 
game has greatly helped to es- 
tablish a wiser use of 
Wildlife Resources 



A CHECKLIST OF AFRICAN MAMMALS 

By Glover M. Allen 

Since the publication of Trouessart's Catalogus Mammalium in 
1904-05, over thirty years have elapsed. In the meantime the conti- 
nent of Africa has been increasingly explored and intensive collecting 
has been carried on in almost every considerable area. This has re- 
sulted in a vast increase in our knowledge of its mammalian fauna and 
often, unfortunately, in a multiplicity of names applied to supposedly 
distinct forms. 

At the present time it may be said that the mammalian fauna of 
Africa is fairly well ascertained, and that the likelihood of many more 
really new genera or species being found is small. Nevertheless much 
revisionary work is yet to be done as adequate collections accumulate 
in the larger museums. As a first necessity in indicating what has been 
done and where further study is urgent, a complete checklist of the 
names applied to African mammals is here presented, intended to in- 
clude all those currently regarded as valid as well as all synonyms 
through July 1938, so far as possible. It is hardly to be hoped that 
no names have been omitted from this catalogue, yet the compiler 
believes that what omissions may be detected are unimportant. So 
far as library resources have been available, the great majority of the 
references have been carefully checked with the originals and the cita- 
tions of names are as originally given, with their type localities follow- 
ing each. 

The area covered is the continent of Africa west of the Suez Canal, 
with the inclusion of Madagascar and the Mascarene islands, and the 
seas adjacent to the continent. Domesticated animals, unless feral, 
and the human races are purposely omitted, but with these excep- 
tions the list includes all species known to have lived within historic 
times. 

In citing the references, the title of the work is first given, followed 
by the volume, then a colon, and after that the page numbers and the 
date. The "type locality" follows the reference. Where there is no 
volume number the page reference is preceded by "p." Genera are 
listed alphabetically under their respective families, and the species 
alphabetically under their genera. Where the range is ascertained, 
it is briefly stated. 



4 bulletin: museum of comparative zoology 

Considerable pains have been given to stating as exactly as possible 
the actual dates of publication. Where, as in the case of the Proceed- 
ings of the Zoological Society of London, the various "parts" for a 
given year are not issued in the same calendar year, the year for which 
the part is published is first given, and the actual date of issue con- 
cludes the reference. For guidance in ascertaining the dates of publica- 
tion of certain works, use has been made of Sherborn's "Index" and the 
following special papers : — 

Duncan, F. Martin 

1937. On the dates of publication of the Society's "Proceedings," 1859- 
1926. With an appendix containing the dates of pubhcation of 
"Proceedings," 1830-1858, compiled by the late F. H. Waterhouse, 
and of the "Transactions," 1833-1869, by the late Henry Peavot, 
originally published in P. Z. S. 1893, 1913. Proc. Zool. Soc. London, 
107: ser. A, 71-84, Apl. 

Sherborn, C. Davies 

1891. On the dates of the parts, plates, and text of Schreber's "Saug- 
thiere." Proc. Zool. Soc. London, for 1891, p. 587-592. 

Sherborn, C. Davies 

1897. On the dates of the natural history portion of Savigny's "Descrip. 
tion de I'Egypte." Proc. Zool. Soc. London, for 1897, p. 285-288. 

Sherborn, C. Davies 

1926. Dates of publication of early catalogues of natural history issued 
by the British Museum. Ann. Mag. Nat. Hist. (9) 17: 271-272, 
Feb. 

Waterhouse, F. H. 

1880. On the dates of publication of the parts of Sir Andrew Smith's 
"Illustrations of the Zoology of South Africa." Proc. Zool. Soc. 
London, for 1880, p. 489-491. 
See also Jentink, F. A., Notes Leyden Mus., 15: 182, 1893. 

I would express my thanks to Miss Barbara Lawrence (now Mrs. 
William E. Schevill), Assistant Curator of Mammals, for much help 
in various ways. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS O 

Class MAMMALIA 

Order INSECTIVORA. Insectivores 
Family MACROSCELIDIDAE. Elephant Shrews 

Elephantulus Thomas and Schwann 

Macroscelides A. Smith, Proc. Zool. Soc. London, for 1830, p. 11, 6 Jan. 1831, 

in part. 
Elephantulus Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 33, p. 

10, 5 June 1906; Proc. Zool. Soc. London, p. 577, 10 Oct. 1906. Genotype 

Macroscelides rupestris A. Smith. 

Elephantulus capensis Roberts 

Elephantulus capensis Roberts, Ann. Transvaal Mus., 10: 62, 31 Jan. 1924. 
Cape Province, Klaver. 

Elephantulus dundasi Dollman 

Elephantulus dundasi Dollman, Ann. Mag. Nat. Hist. (8) 5: 95, Jan. 1910. 
Harich near Lake Baringo, Kenya Colony. 

Elephantulus edwardsii (A. Smith) 

Macroscelides edwardsii A. Smith, Illustr. Zool. South Africa, Mamm., pi. 14, 
1839. Near Oliphants River, Cape Colony. 

Elephantulus fuscipes (Thomas) 

Macroscelides fusdpes Thomas, Ann. Mag. Nat. Hist. (6) 13: 68, Jan. 1894. 
N'doruma, Niam-Niam country, 5° N., 27 30' E. 
Range: Central Africa, N'doruma. 

Elephantulus intufi intufi (A. Smith) 

Macroscelides intufi A. Smith, Report of the Expedition for Exploring Central 

Africa, p. 42, 1836; Illustr. Zool. South Africa, Mamm., part 6, pi. 12, 

1839. Kurrichane, Marico district, Transvaal. 

Elephantulus intufi alexandri (Ogilby) 

Macroscelides alexandri Ogilby, Proc. Zool. Soc. London, p. 5, July 1838. 

Damaraland, Southwest Africa. 
Elephantulus intufi alexandri Shortridge, Mamm. South West Africa, 1 : 22, 

1934. 
Range: Southwest Africa, Damaraland. 

Elephantulus intufi kalaharicus Roberts 

Elephantulus intufi kalaharicus Roberts, Ann. Transvaal Mus., 15: 17, Oct. 
1932. Central Kalahari, Damara Pan, Bechuanaland. 



6 bulletin: museum of comparative zoology 

Elephantulus intufi mossamedensis Hill and Carter 

Elephantulus intufi mossamedensis Hill and Carter, Amer. Mus. Novitates, 
no. 937, p. 1, 13 July 1937. Angola, 101 km. east of Mossamedes. 

Elephantulus ocularis Kershaw 

Elephantulus ocularis Kershaw, Ann. Mag. Nat. Hist. (9) 8: 563, Nov. 1921. 
Dodoma, Tanganyika Territory. 
Range: Tanganyika Territory, Upper Ruaha River. 

Elephantulus peasei (Thomas) 

Macroscelides peasei Thomas, Ann. Mag. Nat. Hist. (7) 8: 154, Aug. 1901. 
Hoolul, Abyssinia. 

Elephantulus pulcher pulcher (Thomas) 

Macroscelides pulcher Thomas, Ann. Mag. Nat. Hist. (6) 13: 69, Jan. 1894. 
Usambiro, south of Victoria Nyanza. 

Elephantulus pulcher rendilis Lonnberg 

Elephantulus pulcher rendilis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 
(2) 48: no. 5, p. 49, 26 June 1912. Kenya Colony, north of Guaso Nyiro. 

Elephantulus renatus Kershaw 

Elephantulus renatus Kershaw, Ann. Mag. Nat. Hist. (9) 11: 588, May 1923. 

Gwao's near Itigi, Singida, about 85 miles northwest of Kilimatinde, 

Tanganyika Territory. 

Elephantulus revoilii (Huet) 

Macroscelides revoilii B.uet, Bull. Soc. Philom. Paris, (7) 5:96, 1881. Somali- 
land. 

Elephantulus rozeti rozeti (Duvernoy) 

Macroscelides rozeti Duvernoy, Mem. Soc. d'Hist. Nat. Strasbourg, 1 : pt. 2, 
art. 4, p. 1, pis. 1, 2, 1830. Near Algiers, Algeria. 
Range: Northern Algeria, Oran, northern Rif. 

Elephantulus rozeti atlantis Thomas 

Elephantulus rozeti atlantis Thomas, Novitates Zool., 20: 587, 21 Oct. 1913. 
Northern slope of Great Atlas of Morocco, south of Seksawa, Am Moussa. 

Elephantulus rozeti clivorum Thomas 

Elephantulus deserti clivorum Thomas, Novitates Zool., 20: 588, 21 Oct. 

1913. Plateau of eastern Algeria, Guelt-es-Stel. 
Range: As above, also Matmata, southern Tunis, and Maafa, eastern 

Algeria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS / 

Elephantulus rozeti deserti (Thomas) 

Macroscelides rozeti deserti Thomas, Ann. Mag. Nat. Hist. (7) 8: 155, Aug. 1901. 
Near Djebel Bourzel, Biskra, Algeria. 

Elephanttjlus rozeti moratus Thomas 

Elephantulus rozeti moratus Thomas, Novitates ZooL, 20: 587, 21 Oct. 1913. 

Djebel Chedar, about 80 kilometers southeast of Mazagan, southwestern 

Morocco. 
Range: Southwest Morocco, also desert of Zragna. 

Elephantulus rufescens rufescens (Peters) 

Macroscelides rufescens Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 198, 
pi. 1, f. 3, 1878. Ndi, Kenya Colony. 
Range: East Africa, Kibwezi, Uataita, Mozambique. 

Elephantulus rufescens boranus (Thomas) 

Macroscelides boranus Thomas, Proc. Zool. Soc. London, for 1900, p. 802, 1 Apl. 
1901. Mega, western Boran Galla, southeast of Lake Rudolf, Kenya 
Colony. 
Range: Lake Rudolf to Guaso Nyiro, Kenya Colony. 

Elephantulus rufescens delicatus DoUman 

Elephantulus delicatus Dollman, Ann. Mag. Nat. Hist. (8) 8: 652, Nov. 1911. 
Mt. Nyiro, Orr Valley, Kenya Colony. 

Elephantulus rufescens mariakan^ Heller 

Elephantulus rufescens mariakanse Heller, Smithsonian Misc. Coll., 60: no. 12, 
p. 10, 4 Nov. 1912. Mariakani, Kenya Colony. 

Elephantulus rufescens ph^eus Heller 

Elephantulus phseus Heller, Smithsonian Misc. Coll., 56: no. 15, p. 8, 23 Dec. 
1910. Sotik district, Njoro O Solali, Kenya Colony. 

Elephantulus rupestris rupestris (A. Smith) 

Macroscelides rupestris A. Smith, Proc. Zool. Soc. London, for 1830, p. 11, 

6 Jan. 1831. Mountains north of Orange River, South Africa. 
Macroscelides typus Lesson, Cent. Zool., p. 51, pi. 12, 1830 (not Smith). 

Elephantulus rupestris jamesoni Chubb 

Elephantulus rupestris jainesoni Chubb, Ann. Transvaal Mus., 1: 181, Jan. 
1909. Johannesburg, Transvaal. 

Elephantulus rupestris mapogonensis Roberts 

Elephantulus rupestris mapogonensis Roberts, Ann. Transvaal Mus., 5: 277, 
16 May 1917. Ngelelle River, Zoutpansberg, Transvaal. 



8 bulletin: museum of comparative zoology 

Elephantulxjs rtjpestris myurtjs Thomas and Schwann 
Elephantulus rupestris myurus Thomas and Schwann, Proc. Zool. Soc. London, 
p. 586, 10 Oct. 1906. Woodbush, northeastern Transvaal. 

Elephantulus somalicus (Thomas) 

Macroscelides somalicus Thomas, Ann. Mag. Nat. Hist. (7) 8: 255, Sep. 1901. 
Arabsiyo, Somaliland. 

Elephantulus van d ami Roberts 

Elephantulus vandami Roberts, Ann. Transvaal Mus., 10: 62, 31 Jan. 1924. 
Cradock, Cape Province. 



Macroscelides A. Smith 

Sorex Shaw, General Zoology, Mamm., 1 : part 2, p. 536, 1800 (in part, not 
Linnaeus. 

Macroscelides A. Smith, Zool. Journ., 4: 435, 1829. Genotype Macroscelides 
typus A. Smith = Sorex proboscideus Shaw. 

Eumerus L Geoffroy, Ann. des Sci. Nat., Zool., (1) 18: 172, Oct. 1829; publ. 
as Eumere, for latinization of name see Table M^thodique des Matieres, 
of same vol., p. 470. Name pubhshed and put into sjmonymy at same time. 

Macroscelis J. B. Fischer, Synopsis Mamm., Addenda, p. 657, 1830. Genotype 
not mentioned. 

Rhinomys Lichtenstein, Darstell. neuer oder wenig bekannt. Saugeth. Zool. 
Mus. Berhn, pi. 38 and text, 1827-34. Genotype Rhinomys jaculus Lich- 
tenstein = Macroscelides proboscideus (Shaw). 

Macroscelides melanotis Ogilby 

Macroscelides melanotis Ogilby, Proc. Zool. Soc. London, for 1838, p. 5, pi. 3, 
f. 3, July 1838. Damaraland, South Africa. 

Macroscelides proboscideus proboscideus (Shaw) 

Sorex proboscideus Shaw, General Zoology, Mamm., 1: part 2, 536, 1800. Cape 

of Good Hope. 
Rhinomys jaculus Lichtenstein, Darstell. neuer oder wenig bekannt. Saugeth. 

Zool. Mus. Berlin, pi. 38 and text, 1827-34. East coast of South Africa. 
Macroscelides typus A. Smith, Zool. Journ., 4: 436, May 1829. Interior of 

South Africa. 
Macroscelides typicus A. Smith, Illustr. Zool. South Africa, Mamm., part 6, 

pi. 10, 1839. 

Macroscelides proboscideus chiversi Roberts 

Macroscelides proboscideus chiversi Roberts, Ann. Transvaal Mus., 15: 265, 
1 Aug. 1933. 76 miles north of Upington district, Bechuanaland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 9 

Macroscelides proboscideus hewitti Roberts 

Macroscelides proboscideus heivitti Roberts, Ann. Transvaal Mus., 13: 86, 6 
July 1929. Cradock, Cape Colony. 

Macroscelides proboscideus langi Roberts 

Macroscelides proboscideus langi Roberts, Ann. Transvaal Mus., 15: 265, 
1 Aug. 1933. Vlermuisklip, van Rhynsdorp district, Cape Colony. 

Nasilio Thomas and Schwann 

Nasilio Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 33, p. 10, 
5 June 1906; Proc. Zool. Soc. London, p. 578, 10 Oct. 1906. Genotype 
Macroscelides brachyrhynchus A. Smith. 

Macroscelides A. Smith (in part). Report of the Expedition for Exploring Cen- 
tral Africa, p. 42, 1836; lUustr. Zool. South Africa, Mamm., part 6, pi. 13, 
1839. 

Nasilio brachyrhynchus brachyrhynchus (A. Smith) 

Macroscelides brachyrhynchus A. Smith, Report of the Expedition for Exploring 

Central Africa, p. 42, 1836; lUustr. Zool. South Africa, Mamm., part 6, 

pi. 13, 1839. "The country between Lake Latakoo and the Tropic." 
Macroscelides brevirostris Schinz, Syst. Verz. Siiugethiere, 1 : 284, 1844. South 

Africa. 
Macroscelides fuscus Peters, Reise nach Mossambique, Zool., 1: Saugethiere, 

p. 87, pis. 19, 22, 1852. Eastern Africa. 
Range: Bechuanaland, Transvaal, Rhodesia, Nyasaland, Mozambique. 

Nasilio brachyrhynchus albiventer Osgood 

Nasilio brachyrhynchus albiventer Osgood, Publ. Field Mus. Nat. Hist., zool. 
ser., 10: 13, 16 Feb. 1910. Lake Elmenteita, Kenya Colony. 
Range: Interior of Kenya Colony, Meru, Naivasha. 

Nasilio brachyrhynchus delamerei (Thomas) 

Macroscelides delamerei Thomas, Ann. Mag. Nat. Hist. (7) 8: 155, 1 Aug. 1901. 
Athi River, Kenya Colony. 
Range: Kenya Colony, coastal region, Athi, Kapiti, and Loita Plains. 

Nasilio brachyrhynchus langi Roberts 

Nasilio brachyrhyncha langi Roberts, Ann. Transvaal Mus., 13: 85, 6 July 1929. 
Portuguese Southeast Africa, lower Limpopo River, Mazambo. 

Nasilio brachyrhynchus lulu^ Matschie 

Nasilio brachyrhynchus Muse Matschie, Zeitschr. f. Saugethierkunde, 1: 110, 
24 Dec. 1926. Belgian Congo, Lulua River, near Luluaburg, Lutaba. 



10 bulletin: museum of comparative zoology 

Nasilio brachyrhynchus mababiensis Roberts 

Nasilio brachyrhynchus mababiensis Roberts, Ann. Transvaal Mus., 15: 18, 
Oct. 1932. Northern Bechuanaland, Tsotsoroga Pan. 

Nasilio brachyrhynchus selindensis Roberts 

Nasilio brachyrhyncha selindensis Roberts, Ann. Transvaal Mus., 19: 99, 
8 Nov. 1937. Mt. Selinda, Southern Rhodesia. 

Nasilio brachyrhynchus shortridgei Roberts 

Nasilio brachyrhyncha shortridgei Roberts, Ann. Transvaal Mus., 13: 86, 6 July 
1929. Ndola, Northern Rhodesia. 

Nasilio brachyrhynchus tzaneenensis Roberts 

Nasilio brachyrhyncha tzaneenensis Roberts, Ann. Transvaal Mus., 13: 85, 
6 July 1929. Northeastern Transvaal, Tzaneen Estate. 

Nasilio brachyurus Bocage 

Macroscelides brachyura Bocage, Jorn. Sci. Math., Phys.eNat., Lisboa, (1) 9: 
27, 1882. Caconda, Angola. 
Range: Angola to Rhodesia. 

Nasilio malos^e (Thomas) 

Macroscelides brachyrhynchus malosae Thomas, Proc. Zool. Soc. London, for 
1897, p. 928, Apl. 1898. Mt. Malosa, Nyasaland, 1800 m. 

Nasilio schinzi (Noack) 

Macroscelides brachyrhynchus, var. schinzi Noack, Zool. Jahrb., Syst., 4: 198, 
28 Feb. 1889. Ondongastamm, Ovamboland. 
Range: Western Africa, Ovambo, Benguela. 

Petrodromus Peters 

Petrodromus Peters, Verh. K. Preuss. Akad. Wiss., Berlin, p. 258, 1846. Geno- 
type Petrodromus tetradactylus Peters. 



Subgenus Petrodromus Peters 

Petrodromus beir^ Roberts 

Petrodromus beirse Roberts, Ann. Transvaal Mus., 4: 69, 20 Oct. 1913. Zimbiti, 
Beira, Mozambique. 

Petrodromus matschiei Neumann 

Petrodromus matschiei Neumann, Zool. Jahrb., Syst., 13: 541, 10 Oct. 1900. 
Burunge and Mkigwa near Tabora, Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 11 

Petrodromus robustus Thomas 

Petrodromus robustus Thomas, Ann. Mag. Nat. Hist. (9) 1: 367, May 1918. 
Upper Lufua River, Katanga, southern Congo Beige. 

Petrodromus tetradactylus tetradactylus Peters 

Petrodromus tetradactylus Peters, Verh. K. Preuss. Akad. Wiss., Berlin, p. 
258, pi. 3, fig. 4, 1846. Tette, southern Mozambique. 

Petrodromus tetradactylus swynnertoni Thomas 

Petrodromus tetradactylus swynnertoni Thomas, Ann. Mag. Nat. Hist. (9) 1: 
368, May 1918. Chirinda Forest, Melsetter, Southern Rhodesia. 

Petrodromus tetradactylus warreni Thomas 

Petrodromus tetradactylus warreni Thomas, Ann. Mag. Nat. Hist. (9) 1: 364, 
May 1918. Manguzi, northern Zululand, Natal. 

Petrodromus tordayi Thomas 

Petrodromus tordayi Thomas, Ann. Mag. Nat. Hist. (8) 5: 83, Jan. 1910. Mis- 

umba, Sankuru River, south-central Congo Beige. 
Petrodromus tordayi tumbanus Kershaw, Revue Zool. Africaine, 11 : 360, 27 Dec. 

1923. Bikoro, Lake Tumba, Congo Beige. 

Petrodromus venustus Thomas 

Petrodromus venustus Thomas, Ann. Mag. Nat. Hist. (7) 12: 339, Sep. 1903. 

Namurive, northern Nyasaland. 
Petrodromus occidentalis Roberts, Ann. Transvaal Mus., 4: 69, 20 Oct. 1913. 
Northwestern Rhodesia. 
Range: Northwestern Rhodesia, southern Nyasaland. 

Subgenus Cercoctenus Hollister 

Cercoctenus Hollister, Smithsonian Misc. Coll., 66: 1, 10 Feb. 1916. Genotype 
Petrodromus sultani Thomas. 

Petrodromus schwanni Thomas and Wroughton 

Petrodromus schwanni Thomas and Wroughton, Proc. ^ool. Soc. London, p. 
289, 1 Aug. 1907. Coguno, Inhambane, Mozambique. 

Petrodromus sultani sultani Thomas 

Petrodromus sultan Thomas, Proc. Zool. Soc. London, p. 435, Oct. 1897. Mom- 
basa, Kenya Colony. Misprint of specific name corrected to sultani in 
ibid., p. 928, Apl. 1898. 

Petrodromus sultani sangi Heller 

Petrodromus sultani sangi Heller, Smithsonian Misc. Coll., 60: no. 12, p. 12, 
4 Nov. 1912. Mt. Mbololo, Taita Hills, Kenya Colony. 



12 bulletin: museum of comparative zoology 

Subgenus Mesoctenus Thomas 

Mesociemis Thomas, Ann. Mag. Nat. Hist. (9) 1 : 366, May 1918. As a genus, 
type Petrodromus rovumse Thomas. 

Petrodromus nigriseta Neumann 

Petrodromus nigriseta Neumann, Zool. Jahrb., Syst., 13: 541, footnote, 10 Oct. 
1900. Marui and Morogoro, Usaramo, Tanganyika Territory. 

Petrodromus rovum^ Thomas 

Petrodromus rovwnx Thomas, Proc. Zool. Soc. London, p. 434, Oct. 1897. 

Rovuma River, 100 miles inland, Tanganyika Territory. 
Petrodromus (Mesoctenus) mossambicus Thomas, Ann. Mag. Nat. Hist. (9) 1 : 

369, May 1918. Cabaceira, Mozambique. 

Rhinonax Thomas. Black-and-rufous Elephant-shrews 

Rhinonax Thomas, Ann. Mag. Nat. Hist. (9) 1: 370, May 1918. As a subgenus 
of Rhynchocyon; type Rhynchocyon chrysopygus Gunther. 

Rhinonax adersi (Dollman) 

Rhynchocyon adersi Dollman, Ann. Mag. Nat. Hist. (8) 10: 130, July 1912. 
Zanzibar. 

Rhinonax chrysopygus (Gunther) 

Rhynchocyon chrysopygus Gunther, Proc. Zool. Soc. London, p. 164, pi. 14, 
June 1881. River Mombasa, Kenya Colony. 
Range: East coast of Kenya Colony. 

Rhinonax petersi petersi (Bocage) 

Rhynchocyon petersi Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (1) 7: 

159, pi. 4, f. 2, Feb. 1880. "Envoys de Zanzibar," but actually from the 

opposite mainland of Tanganyika Territory (see Dollman, G., Ann. Mag. 

Nat. Hist. (8) 10: 131, July 1912). 
Rhynchocyon petersi usambaras Neumann, Zool. Jahrb., Syst., 13: 542, 10 Oct. 

1900. Usambara. 
Range: Tanganyika Territory. 

Rhinonax petersi fischeri (Neumann) 

Rhynchocyon petersi fischeri Neumann, Zool. Jahrb., Syst., 13: 543, 10 Oct. 
1900. LTsegua, Tanganyika Territory. 

Rhinonax petersi melanurus (Neumann) 

Rhynchocyon petersi melanurus Neumann, Zool. Jahrb., Syst., iS: 542, 10 Oct. 
1900. Lindi, Uluguru Mts., Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 13 

Rhynchocyon Peters. Checkered Elephant-shrews 

Rhynchocyon Peters, Ber. iiber Verhandl. K. Preuss. Akad. Wiss., Berlin, p. 36, 
1847. Genotype Rhynchocyon cirnei Peters. 

Rhynchocyon cirnei Peters 

Rhynchocyon cirnei Peters, Ber. iiber Verhandl. K. Preuss. Akad. Wiss., Berlin, 
p. 37, pi. 4, f. 1, 1847. Quellimane, Bororo district, Mozambique. 

Rhynchocyon claudi Thomas and Wroughton 

Rhynchocyon claudi Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 370, 
May 1907. Beritio, Uele River, Congo Beige. 

Rhynchocyon hendersoni Thomas 

Rhynchocyon hendersoni Thomas, Ann. Mag. Nat. Hist. (7) 10: 403, Nov. 1902. 
Plateau west of Lake Naivasha, Kenya Colony. 

Rhynchocyon macrtjrus Giinther 

Rhynchocyon macrurus Giinther, Proc. Zool. Soc. London, p. 163, June 1881. 
Rovuma River, 8° S., Tanganyika Territory. 
Range: Rovuma River to east of Lake Nyasa. 

Rhynchocyon reichardi Reichenow 

Rhynchocyon reichardi Reichenow, Zool. Anzeiger, 9: 316, 17 May 1886. Ma- 
rungu, southeastern Congo Beige. 

Rhynchocyon stuhlmanni stuhlmanni Matschie 

Rhynchocyon stuhlmanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 66, 
1893. Ruanda, Semliki River, northeastern Congo Beige. 

Rhynchocyon stuhlmanni nudicatjdatus Lydekker 

Rhynchocyon stuhlmanni nudicaudata Lydekker, Proc. Zool. Soc. London, for 
1906, p. 995, Apr. 1907. Ituri Forest, Congo Beige. 

Rhynchocyon swynnertoni Kershaw 

Rhynchocyon swynnertoni Kershaw, Ann. Mag. Nat. Hist. (9) 11: 587, May 
1923. Kipera, Kilosa, Tanganyika Territory. 



14 bulletin: museum of comparative zoology 



Family CHRYSOCHLORID^. Golden Moles 

Broom (1916) believes the golden moles are not Insectivora, but should 
stand as a separate order, Chrysochloroidea Dobson. 



Amblysomus Pomel 
Subgenus Amblysomus Pomel 

Amblysomus Pomel, Arch, des Sci. Phys. et Nat., Geneve, 9: 244, 1848. Geno- 
type Chrysochloris hottentotus A. Smith. 

Calcochloris Mivart, Journ. Anat. and Phys., 2 (= ser. 2, vol. 1): 133, 1868. 
Genotype Chrysochloris obtusirostris Peters. 

Amblysomus corri^ coRRiiE Thomas 

Amblysomus corrise Thomas, Abstr. Proc. Zool. Soc. London, no. 20, p. 5, 

23 May 1905; Proc. Zool. Soc. London, 2: 57, 7 Oct. 1905. Knysna, 

southern Cape Colony. 

Amblysomus corri^ septentrionalis Roberts 

Amblysomus corriae septentrionalis Roberts, Ann. Transvaal Mus., 4; 73, 20 
Oct. 1913. Wakkerstroom, Transvaal. 

Amblysomus hottentotus hottentotus (A. Smith) 

Talpa rufa Shaw, General Zool., 1: pt. 2, p. 522, 1800. Said to be "America," 

but perhaps refers to this species. 
Chrysochloris hottentotus A. Smith, Zool. Journ., 4: 436, May 1829. "Interior 

parts of South Africa." 
Chrysochloris holosericea Lichtenstein, Darstellung neue oder wenig bek. 

Saugeth., pi. 41, f. 2, and text, 1834. Interior of Cape Colony, on the 

borders of "Kafferland." 
Chrysochloris affinis Wagner, Schreber's Saugthiere, Suppl., 2: 123, footnote, 

1844. No locality. 
Chrysochloris rutilans Wagner, Schreber's Saugthiere, Suppl., 2: 125, 1844. 

"Kafferlande." 

Amblysomus hottentotus albifrons (Broom) 

Chrysochloris hottentota albifrons Broom, Trans. South African Philos. Soc, 18: 
302, 24 Dec. 1907. Howick, Natal. 

Amblysomus hottentotus albirostris (Wagner) 

Chrysochloris albirostris Wagner, Schreber's Saugthiere, Suppl., 2: 124, 1841. 

"Kaffernland." "May prove to be distinct" (Thomas and Schwann). 

A subsp. of hottentota according to Broom, Ann. Transvaal Mus., 1: 15, 

1908. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 15 

Amblysomus hottentottus pondolise Thomas and Schwann, Proc. Zool. Soc. 
London, 1: 260, 10 Aug. 1905. Notinsila, western Pondoland, Cape 
Colony. 

Amblysomus hottentotus garneri Roberts 

Amblysomus hottentottus garneri Roberts, Ann. Transvaal Mus., 5: 278, 16 May 
1917. Commissioner's residence, Piggs Peak, Swaziland, Transvaal. 

Amblysomus hottentotus iris Thomas and Schwann 

Amblysomus iris Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 18, 

p. 23, 25 Apl. 1905; Proc. Zool. Soc. London, 1: 259, pi. 16, f. 1, 10 

Aug. 1905. Umvolosi Station, Zululand, Natal. 

Amblysomus hottentotus longiceps (Broom) 

Chrysochloris hottentota longiceps Broom, Trans. South African Philos. Soc, 
18: 299, 24 Dec. 1907. Near Pietermaritzburg, Natal. 

Amblysomus marleyi Roberts 

Amblysomus marleyi Roberts, Ann. Transvaal Mus., 14: 225, 28 July 1931. 
Ubombo, Zululand, Natal. 



Subgenus Chrysotricha Broom 

Chrysotricha Broom, Trans. South African Philos. Soc, 18: 303, 24 Dec. 1907. 
As a subgenus of Chrysochloris, type Chrysochloris obtusirostris Peters; 
raised to generic rank by Roberts, Ann. Transvaal Mus., 18: 225, 21 Feb. 
1936. 

Amblysomus obtusirostris obtusirostris (Peters) 

Chrysochloris obtusirostris Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 
p. 467, 1851; Reise nach Mossambique, Saugeth., p. 70, pi. 18, f. 1; pi. 22, 
f. 18-23, 1852. Inhambane, Portuguese East Africa, lat. 24° S. 

Amblysomus obtusirostris chrysillus Thomas and Schwann 
Amblysomus chrysillus Thomas and Schwann, Proc. Zool. Soc London, 1: 261, 

pi. 16, f. 2, 10 Aug. 1905. Delagoa Bay, Portuguese East Africa. 
Chrysotricha obtusirostris chrysilla Roberts, Ann. Transvaal Mus., 18: 225, 

21 Feb. 1936. 

Bematiscus Cope 

Bematiscus Cope, Amer. Naturalist, 26: 127, footnote, Feb. 1892. Genotype, 
Chrysochloris villosus A. Smith, see Thomas, Proc. Zool. Soc. London, 1 : 
259, 1905. 



16 bulletin: museum of comparative zoology 

Bematisctjs rufopallidus Roberts • 

Bematiscus rufopallidus Roberts, Ann. Transvaal Mus., 10: 65, 31 Jan. 1924. 
Wakkerstroom, Transvaal. 

Bematiscus transvaalensis transvaalensis Broom 

Bematiscus transvaalensis Broom, Abstr. Proc. Zool. Soc. London, no. 121, p. 25, 

13 May 1913; Proc. Zool. Soc. London, p. 546, 1 Sep. 1913. Springs, 

Transvaal. 

Bematiscus transvaalensis pratensis (Roberts) 

Chrysospalax pratensis Roberts, Ann. Transvaal Mns., 4: 74, 20 Oct. 1913, 

Pretoria, Transvaal. 

A subspecies of B. transvaalensis ?, see Roberts, Ann. Transvaal Mus., 
10: 65, 31 Jan. 1924. 

Bematiscus villosus (A. Smith) 

Chrysochloris villosus A. Smith, South African Quart. Journ., 2: 81, 1833; 
Illustr. Zool. South Air., Mamm., pi. 9 and text, 1839. "Towards Natal." 



Chlorotalpa Roberts 

Chlorotalpa Roberts, Ann. Transvaal Mus., 10: 64, 31 Jan. 1924. Genotype 
Chrysochloris duthiex Broom. 

Chlorotalpa duthiex (Broom) 

Chrysochloris duthiex Broom, Trans. South African Philos. Soc, 18: 292, 
24 Dec. 1907. Knysna, Cape Colony. 

Chlorotalpa fosteri St. Leger 

Chlorotalpa fosteri St. Leger, Ann. Mag. Nat. Hist. (10) 8: 605, Dec. 1931. 
Mt. Elgon, Uganda, 9300 feet. 

Chlorotalpa guillarmodi Roberts 

Chlorotalpa guillarmodi Roberts, Ann. Transvaal Mus., 18: 253, 21 Feb. 1936. 
Mamathes, northwestern Basutoland. 

Chlorotalpa Montana Roberts 

Chlorotalpa montana Roberts, Ann. Transvaal Mus., 10: 64, 31 Jan. 1924. 
Kastrol Nek, east of Wakkerstroom, Transvaal, 6500 feet. 

Chlorotalpa stuhlmanni (Matschie) 

Chrysochloris stuhlmanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 123, 
1894. Ukonjo and Kinyawanga, Ruwenzori region, Uganda. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 17 

Chlorotalpa tropicalis G. M. Allen and Loveridge 

Chlorotalpa tropicalis G. M. Allen and Loveridge, Proc. Boston Soc. Nat. Hist., 

38: 418, Dec. 1927. Bagilo, Uluguru Mountains, Tanganyika Territory, 

6000 feet. 

Chrysochloris Lacepede 
Subgenus Chrysochloris Lacepede 

Chrysochloris Lacepede, Tabl. des Mamm., p. 7, 1799; Buffon's Hist. Nat., 
Quadrupedes, Didot ed., 14: 158, 1799. Genotype: included C. capensis 
and C. rubra, hence is Talpa asiatica Linnseus. 

Talpa, Sorex, in part, of older authors. 

Chrysochloris asiatica (Linnaeus) 

Talpa asiatica Linnaeus, Syst. Nat., ed. 10, 1 : 53, 1758. "In Sibiria." 

Talpa aurea Pallas, in Schreber's Saugthiere, 3: 562, 1777. Cape of Good Hope; 

a substitute name for asiatica since the animal is not Asiatic. 
Talpa inaurata Pallas, in Schreber's Saugthiere, pi. 157, 1777. Cape of Good 

Hope. In the text (3: 562) this plate is referred to Talpa aurea, of which 

the name becomes therefore a synonym. 
Sorex auratus Vosmaer, Beschreibung Groeng. MoL, fig., 1787. 
Chrysochloris rubra Lacepede and Daudin, in Lacepede's Tabl. des Mamm., p. 

7, 1799. Cape of Good Hope. 
Chrysochloris capensis Lacepede and Daudin, in Lacepede's Tabl. des Mamm., 

p. 7, 1799. Cape of Good Hope. 

Chrysochloris bayoni de Beaux 

Chrysochloris bayoni de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. 
Nat. Milano, 60: 236, Sep. 1921. Robben Island, Cape Colony. 

Chrysochloris congicus Thomas 

Chrysochloris congicus Thomas, Ann. Mag. Nat. Hist. (8) 5: 84, Jan. 1910. 
Lusambo, upper Sankuru River, south-central Congo Beige. 

Chrysochloris dam.\rensis Ogilby 

Chrysochloris damarensis Ogilby, Proc. Zool. Soc. London, p. 5, July 1838. 
Damaraland, Southwest Africa. 

Chrysochloris leucorhina leucorhina Huet 

Chrysochloris leucorhina Huet, Nouv. Arch. Mus. d'Hist. Nat., Paris, (2) 
8: 8, pi. 1, 1885. Coast of the Gulf of Guinea. 

Chrysochloris leucorhina cahni Schwarz and Mertens 
Chrysochloris leucorhina cahni Schwarz and Mertens, Senckenbergiana, 4: 
151, 30 Nov. 1922. Molundu, Ja River, southeastern Cameroons. 



18 bulletin: museum of comparative zoology 

Chrysochloris minor Roberts 

Chrysochloris minor Roberts, Ann. Transvaal Mus., 6: 113, Apl. 1919. Klaver, 
on Olifants River, Cape Province. 

Chrysochloris namaqtjensis Broom 

Chrysochloris namaquensis Broom, Ann. Mag. Nat. Hist. (7) 19: 266, Mar. 
1907. Caries, Namaqualand, Southwest Africa. 

Chrysochloris sclateri Broom 

Chrysochloris sclateri Broom, Ann. Mag. Nat. Hist. (7) 19; 263, Mar. 1907. 
Beaufort West, Cape Colony. 

Chrysochloris tenuis Broom 

Chrysochloris tenuis Broom, Ann. Mag. Nat. Hist. (7) 19: 267, Mar. 1907. 
Garies, Namaqualand, Southwest Africa. 

Chrysochloris vermiculus Thomas 

Chrysochloris vermiculus Thomas, Ann. Mag. Nat. Hist. (8) 5: 84, Jan. 1910. 
Yambuya, Aruwimi River, Congo Beige. 

Chrysochloris wintoni Broom 

Chrysochloris wintoni Broom, Ann. Mag. Nat. Hist. (7) 19: 264, Mar. 1907. 
Port Nolloth, Cape Colony. 

Chrysospalax Gill 

Chrysospalax Gill, Standard Nat. Hist., 5 (Mammaha): 137, 26 Dec. 1883. 
As a subgenus; type Chrysochloris trevelyani Giinther. See Roberts, 
Ann. Transvaal Mus., 10: 64, 31 Jan. 1924. 

Chrysospalax dobsoni (Broom) 

Bematiscus dobsoni Broom, Proc. Zool. Soc. London, p. 190, 18 Aug. 1918. 

Maritzburg, Natal. 

Chrysospalax lesch^ (Broom) 

Bematiscus leschse Broom, Proc. Zool. Soc. London, p. 189, 18 Aug. 1918. 
Isolo, Cape Colony. 

Chrysospalax trevelyani (Giinther) 

Chrysochloris trevelyani Giinther, Proc. Zool. Soc. London, p. 311, Aug. 1875. 
Pirie forest, near King William's Town, Cape Province. 

Eremitalpa Roberts 

Eremitalpa Roberts, Ann. Transvaal Mus., 10: 63, 31 Jan. 1924. Genotype 
Chrysochloris granti Broom. 



ALLEN: CHECKLIST OF AFRICAN MAIMMALS 19 

Eremitalpa granti (Broom) 

Chrysochloris granti Broom, Ann. Mag. Nat. Hist. (7) 19: 265, Mar. 1907. 
Garies, Namaqualand, Southwest Africa. 
For Claud H. B. Grant. 

Neamblysomus Roberts 

Neamblysomus Roberts, Ann. Transvaal Mus., 10: 64, 31 Jan. 1924. Genotype 
Chrysochloris gunningi Broom. 

Neamblysomus gunningi (Broom) 

Chrysochloris gunningi Broom, Ann. Transvaal Mus., 1: 14, Apl. 1908. Wood- 
brush Hill, Zoutpansberg, Transvaal. 
For Dr. J. W. B. Gunning, Director of Transvaal Mus. 



Family ERINACEID.E. Hedgehogs 

See Thomas, O. The generic division of the Hedgehogs. Ann. Mag. Nat* 
Hist. (9) 1: 193-196, Feb. 1918. Cabrera, A. Genera Mammalium. Insecti- 
vora, Galeopithecia, 1925. 



.^thechinus Thomas. Four-toed Hedgehogs 

Mhechinus Thomas, Ann. Mag. Nat. Hist. (9) 1: 194, Feb. 1918. Genotype 

Erinaceus algirus Duvernoy and Lereboullet. 
Erinaceus A. Smith, South African Quart. Journ., p. 10, 1831 (in part). 

^THECHiNUS ALGIRUS ALGIRUS (Duvernoy and Lereboullet) 

Erinaceus algirus Duvernoy and Lereboullet, Mem. Soc. Mus. d'Hist. Nat. 

Strasbourg, 3: art. QQ, 4, 1840. Oran, Algeria. 
Erinaceus Jallax Dobson, Monogr. Insectivora, 1: 9, 1882. 
Range: Mogador to Tripoli. 

.^THECHINUS ALGIRUS CANICULUS (Thomas) 

Erinaceus algirus caniculus Thomas, Ann. Mag. Nat. Hist. (8) 16: 152, Aug. 
1915. Toston, Fuerte-ventura, Canary Islands. 

Erinaceus krugi Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 78, 1877. 
Locality unknown, probably West Africa or southern Europe (type speci- 
men killed in Mayaguez, Porto Rico). 

.(Ethechinus ALGIRUS LAVAUDENi Cabrera 

.Eihechinus algirus lavaudeni Cabrera, Bol. Real Soc. Espan. Hist. Nat., 
Madrid, 28: 454, 30 Nov. 1928. Mogador, Morocco. 



20 bulletin: museum of comparative zoology 

Erinaceus diadematus Dobson, Monogr. Insectivora, 1: 10, 1882 (part, not 
Fitzinger). 
Range: Southern Morocco. 

Named for Louis Lavauden, chief of the Forest Service of Madagascar, in 
recognition of his interest in mammals of No. Africa. 

.^THECHINUS FRONTALIS FRONTALIS (A. Smith) 

Erinaceus frontalis A. Smith, South African Quart. Journ., 2: 10, 29, 1831. 

Cape Colony. 
Erinaceus capensis A. Smith, Philos. Mag., 9: 61, 1831 (nomen nudum). 
Erinaceus fractilis Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 79, 1877 

(lapsus for frontalis) . 

tEthechinus frontalls angola Thomas 

/Ethechinus angolse Thomas, Ann. Mag. Nat. Hist. (9) 1: 230, Mar. 1918. 

Benguella, Angola. 
.Ethechinus frontalis angolse Thomas, Proc. Zool. Soc. London, p. 290, 29 Apl. 

1926. 

^Ethechinus sclateri (Anderson) 

Erinaceus sclateri Anderson, Proc. Zool. Soc. London, p. 415, fig., Oct. 1895. 
Somaliland. 

Atelerix Pomel 

Atelerix Pomel, Arch, des Sci. Phys. et Nat. Geneve, 9: 251, 1848. Genotype 

Erinaceus albiventris Wagner; type fixed by Thomas (Ann. Mag. Nat. 

Hist. (9) 1: 195, Feb. 1918). 
Erinaceus Wagner, Schreber's Saugthiere, Suppl., 2: 22, 1841 (in part, not of 

Linnaeus). 
Peroechinus Fitzinger, Sitzb. K. Akad. Wiss. Wien, 54: sect. 1, p. 565, 1866; 

56: sect. 1, p. 856, 1867. Genotype Erinaceus pruneri Wagner. 

Atelerix albiventris (Wagner) 

E[rinaceus] albiventris Wagner, Schreber's Saugthiere, Suppl., 2: 22, 1841. 

Locality unknown, but doubtless Senegambia. Regarded as unidentifiable 

by J. A. Allen (Bull. Amer. Mus. Nat. Hist., 47: 8, 1922) but see Allen, 

G.M., Journ. Mammalogy, 17: 62, Feb. 1936. 
Erinaceus adansoni Rochebrune, Bull. Soc. Philom. Paris, (7) 7: 7, 1883. 

"Saint-Louis, Pointe de Barbarie, Cap Vert, Joalles, etc." 
Range: Senegambia. 

Atelerix pruneri pruneri (Wagner) 

E[rinaceus] pruneri Wagner, Schreber's Saugthiere, Suppl., 2: 23, 1841. Upper 
Nile, taken as Sennaar, where Pruner travelled. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 21 

Erinaceus heterodactylus Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stock- 
holm, for 1841, p. 227, 1842. Bahr-el-Abiad, Sennaar. 

Erinaceus diadematus Fitzinger, Sitzb. K. Akad. Wiss. Wien, 56: sect. 1, 
p. 853, 1867. Sennaar and Kordofan. 

Atelerix pruneri atratus (Rhoads) 

Erinaceus albiventris atratus Rhoads, Proc. Acad. Nat. Sci., Philadelphia, p. 544, 
8 Dec. 1896. Lake Rudolf, Ngare Nocbor, Abyssinia. 

Atelerix pruneri hindei (Thomas) 

Erinaceus hindei Thomas, Ann. Mag. Nat. Hist. (8) 5, 193, Feb. 1910. Kitui, 

Kenya Colony, 3500 feet. 
Erinaceus sotikie Heller, Smithsonian Misc. Coll., 56: no. 15, p. 1, 23 Dec. 1910. 

Southern Guaso Nyiro, Sotik District, Kenya Colony. 

Atelerix pruneri kilimanus Thomas 

Atelerix kilimanus Thomas, Ann. Mag. Nat. Hist. (9) 1: 232, Mar. 1918. 
Rombo, Mt. Kilimanjaro, Tanganyika Territory, 5300 feet. 

Atelerix faradjius J. A. Allen 

Atelerix faradjius J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 13, 20 July 1922. 
Faradje, northeastern Congo Beige. 

Atelerix langi J. A. Allen 

Atelerix langi J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 15, 20 July 1922. 
Faradje, northeastern Congo Beige. 

Atelerix spiculus (Thomas and Wroughton) 

Erinaceus spiculus Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 371, 
May 1907. Maifoni, near Lake Chad, Nigeria. 

Atelerix spinifex Thomas 

Atelerix spinifex Thomas, Ann. Mag. Nat. Hist. (9) 1: 231, Mar. 1918. Illela, 
Nigeria, 95 miles north of Sokoto. 

Hemiechinus Fitzinger. Long-eared Hedgehogs 

Hemiechinus Fitzinger, Sitzb. K. Akad. Wiss. Wien, 54: sect. 1, 565, 1866. 
Genotype Erinaceus platyotis Sundevall = E. libycus Hemprich and Ehren- 
berg. Included five species, without selection of genotype. 

Ericius Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 1841, 
p. 223, 1842 (not Tilesius, 1813). Included seven species; no type desig- 
nated. 

Erinaceus (in part), Pallas, Nov. Comment. Acad. Petrop., 14: 593, 1770 (not 
Linnaeus). 



22 bulletin: museum of comparative zoology 

Hemiechinus libycus (Hemprich and Ehrenberg) 

Erinaceus auritus Geoffrey and Audouin, Descrip. de I'Egypte, 2: 737, pi. 5, 

f. 3, 1829 (not of Pallas). Borders of the Egyptian desert. 
Erinaceus libycus Hemprich and Ehrenberg, Symbolse Physicse, Mamm., dec. 

2, sheet k, footnote, Sep. 1832. Libyan desert, near Alexandria, Egypt. 
Ericius 'platyotis Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 

for 1841, p. 232, 1842. Egypt. 
Erinaceus segyptius Riippell, Mus. Senckenberg., 3: 159, 1842. Egypt. (Name 

based on Geoffroy, Descr. de I'Egypte, Mamm.). 



Paraechinus Trouessart 

Paraechinus Trouessart, Rev. et Mag. de Zool., (3) 7: 242, 1879. Genotype 

Erinaceus micropus Blyth. 
Macroechinus Satunin, Annuaire Mus. Zool. Acad. Imp. Sci., St. Petersbourg, 

11: 189, Mar. 1907. Genotypes Erinaceus hypomelanus Brandt and E. 

macracanthus Blanford, of Asia. 

Paraechinus ^thiopicus (Hemprich and Ehrenberg) 

Erinaceus sethiopicus Hemprich and Ehrenberg, Symbolae Physicse, Mamm., 

dec. 2, sheet k, footnote, Sep. 1832. Deserts of Dongola, Anglo-Egyptian 

Sudan. 
Erinaceus senaariensis Hedenborg, Oken's Isis, p. 8, 1839. 
[Erinaceus] brachydactytus Wagner, Schreber's Saugthiere, Suppl., 2: 24, 1841. 

Egypt. 
Paraechinus sethiopicus Cabrera, Genera Mamm., Insectivora, Galeopithecia, 

p. 70, pi. 7, fig. 4, 1925. 
Hemiechinus pallidus Fitzinger, Sitzb. K. Akad. Wiss. Wien, 54: sect. 1, p. 565, 

1866 (nomen nudun); 56: sect. 1, p. 866, 1867. Sennaar. 

Paraechinus deserti deserti (Loche) 

Erinaceus deserti Loche, Cat. Mamm. et Oiseaux Algeria, p. 20, 1858. Explora- 
tion Sci. de I'Algerie, Zool., Mammiferes, p. 93, 1867. Southern Sahara, 
in the oasis of the Beni-Mzab, Ouargla, and Tuggurt. 

Paraechinus deserti blancalis Thomas 

Paraechinus deserti blancalis Thomas, Ann. Mag. Nat. Hist. (9) 8: 570, Nov. 
1921. Island of Djerba, southeastern Tunis. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 23 

Family TENRECID.E. Tenrecs 

Cryptogale G. Grandidier 

Cryptogale G. Grandidier, Bull. Mus. d'Hist. Nat. Paris, 34: 64, 1928. Geno- 
type Cryptogale australis G. Grandidier. 

Cryptogale australis G. Grandidier 

Cryptogale australis G. Grandidier, Bull. Mus. d'Hist. Nat. Paris, 34: 64, 
1928. Grotte d'Andrahomana, south of Fort Dauphin, Madagascar. 

Dasogale G. Grandidier 

Dasogale G. Grandidier, Bull. Acad. Malgache, for 1928, p. 85, 1930. Genotype 
Dasogale fontoynonti G. Grandidier. 

Dasogale fontoynonti G. Grandidier. 

Dasogale fontoynonti G. Grandidier, Bull. Acad. Malgache, for 1928, p. 85, 
2 pis., 5 text-f., 1930. Forests of eastern Madagascar. 

EcHiNOPS Martin 

Echinops Martin, Proc. Zool. Soc. London, p. 17, July 1838. Genotype Echi- 

nops telfairi Martin. 
Echinogale Wagner, Schreber's Saugth., Suppl., 2: iii, 29, 1841 (misprinted 

Eghinogale on p. 29). Genotype Echinops telfairi Martin. 

Echinops telfairi telfairi Martin 

Echinops telfairi Martin, Proc. Zool. Soc. London, p. 17, July 1838. Mada- 
gascar. 

Setiger setosus Jentink, Notes Leyden Mas., 1: 142, July 1879 (in part, not 
Schreber) . 

Echinops miwarti A. Grandidier, Rev. et Mag. de Zool. (2) 21: 338, Sep. 1869. 
Tulear, Madagascar. 

Echinops telfairi nigrescens (Petit) 

Ericulus (Echinops) telfairi nigrescens Petit, Faune d. Colonies Fran^aises, 

Paris, 4: no. 5, pp. 570, 575, 1931. Southwest coast of Madagascar, 

Morombe to Androka. 

Echinops telfairi pallescens Thomas 

Echinops telfairi pallescens Thomas, Proc. Zool. Soc. London, p. 501, Oct. 1892. 
Manumbu, southern Madagascar. 



24 bulletin: museuim of comparative zoology 

Hemicentetes Mivart 

Hemicentetes Mivart, Proc. Zool. Soc. London, p. 72, June 1871. Genotype 

Erinaceus madagascariensis Shaw. 
Erinaceus Schreber, Saugth., 3: pi. 166, 1777 (in part). 
Setiger E. Geoffrey, Catal. Mamm. Mus. d'Hist. Nat., Paris, p. 72, 1803 (in 

part). 
Centenes G. Cuvier, Regne Anim., ed. 1, 1: 136, 1817 (in part). 
Centetes Schinz and Brodtmann (not Illiger), Naturg. Abbild. Saugth., pi. 26, 

1827. 
Ericius Giebel, Zeitschr. Ges. Naturw., 37: 57, 1871 (not Tilesius 1813; not 

Sundevall 1841). 
Echinodes Pomel, Arch, des Sci. Phys. et Nat., Geneve, 9: 251, 1848 (nomen 

nudum). 

Hemicentetes nigriceps Giinther 

Hemicenteles nigriceps Giinther, Ann. Mag. Nat. Hist. (4) 16: 125, Aug. 1875. 

Fienerentova, Madagascar. 
Hemicenteles variegatus, var. huffoni Jentink, Notes Leyden Mus., 1: 150, July 

1879. Madagascar. 

Hemicentetes semispinosus (G. Cuvier) 

Ericulus semispinosus Cuvier, Tabl. Elem. d'Hist. Nat., p. 108, 1798.. Mada- 
gascar. 

Erinaceus ecaudatus Schreber, Saugth., 3: 584, 590, pi. 165*, 1777. Mada- 
gascar. 

Erinaceus madagascariensis Shaw, General Zool., Mamm., 1: pt. 2, p. 548, 
1800 (not Zimmermann). Madagascar. 

Hemicentetes semispinosus variegatus E. tieoffroy, Cat. Mamm. Mus. National 
d'Hist. Nat., Paris, p. 72, 1803. 

Leptogale Thomas 

Leptogale Thomas, Ann. Mag. Nat. Hist. (9) 1: 306, Apr. 1918. Genotype 

Oryzoryctes gracilis Major. 
Oryzoryctes Major, Ann. Mag. Nat. Hist. (6) 18, 321, Oct. 1896 (in part, not 

Grandidier). 

Leptogale gracilis (Major) 

Oryzoryctes gracilis Major, Ann. Mag. Nat. Hist. (6) 18: 321, Oct. 1896. Mada- 
gascar, Ambohimitombo Forest. 

LiMNOGALE Major 

Limnogale Major, Ann. Mag. Nat. Hist. (6) 18: 318, Oct. 1896. Genotype 
Limnogale mergulus Major. 



ALLEN: CHECKLIST OF AFRICAN 1MAM1\L\LS 25 

LiMNOGALE MERGULUS MajOF 

Limnogale mergulus Major, Ann. Mag. Nat. Hist. (6) 18: 319, Oct. 1896. 
Imasindrary, Betsileo, Madagascar. 



MiCROGALE Thomas 

Microgale Thomas, Journ. Linn. Soc. London, zool., 16: 319, 1882. Genotype 
Microgale longicaudala Thomas. 
On the grouping of the described forms of the genus, see Grandidier, G., 
Bull. Mus. National d'Hist. Nat. Paris (2) 6: 477, Nov. 1934. 

Microgale brevicaudata G. Grandidier 

Microgale brevicaudata G. Grandidier, Bull. Mus. d'Hist. Nat., Paris, 5: 349, 
1899. Mahanara, northeastern Madagascar. 

Microgale cowani Thomas 

Microgale cowani Thomas, Journ. Linn. Soc. London, zool., 16: 320, 1882. 
East Betsileo, Ankdfana Forest, Madagascar. 

Microgale crassipes Milne-Edwards 

Microgale crassipes Milne-Edwards, Ann. des Sci. Nat., zool. (7) 15: 98, 1893. 
Tananarive, Madagascar. 

Microgale drotjhardi G. Grandidier 

Microgale drouhardi G. Grandidier, Bull. Mus. d'Hist. Nat., Paris, (2) 6: 474, 
1934. Diego-Suarez, northern Madagascar. 

Microgale longicatjdata Thomas 

Microgale longicaudala Thomas, Journ. Linn. Soc. London, zool., 16: 320, 1882. 
East Betsileo, Ankafana Forest, Madagascar. 

Microgale longirostris Major 

Microgale longirostris Major, Ann. Mag. Nat. Hist. (6) 18: 321, Oct. 1896. 

Madagascar. 
Microgale cowani nigrescens Elliot, Proc. Biol. Soc. Washington, 18: 237, 9 Dec. 

1905. Madagascar. 

Microgale majori Thomas 

Microgale majori Thomas, Ann. Mag. Nat. Hist. (9) 1: 305, Apr. 1918. East 
Betsileo, Ankafana Forest, Madagascar. 

Microgale parvula G. Grandidier 

Microgale parvula G. Grandidier, Bull. Mus. d'Hist. Nat., Paris, (2) 6: 476, 
1934. Diego-Suarez, northern Madagascar. 



26 bulletin: museum of comparative zoology 

MicROGALE PRiNCiPULA Thomas 

Mirrogale principula Thomas, Ann. Mag. Nat. Hist. (9) 17: 251, Feb. 1926. 
Midongy-du-Sud, southeastern Madagascar. 

MiCROGALE PROLiXACAUDATA G. Grandidier 

Microgale prolixacaudata G. Grandidier, Bull. Mus. d'Hist. Nat., Paris, (2) 9: 
348, ff. 2, 4, 6, Nov. 1937. Diego-Suarez region, extreme northern Mada- 
gascar. 

Microgale pusilla Forsyth Major 

Microgale pusilla Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 462, Dec. 1896. 

Forest of the Independent Tanala of Ikongo, near Vinanitelo, Madagascar; 

also subfossil in Children's Cave, near Sirabe. 

Microgale sorella Thomas 

Microgale sorella Thomas, Ann. Mag. Nat. Hist. (9) 17: 252, Feb. 1926. Be- 
forona Forest, inland of Andevorante, eastern Madagascar. 

Microgale taiva Forsyth Major 

Microgale taiva Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 461, Dec. 1896. 
Ambohimitombo forest, Tanala country, Madagascar. 

Microgale thomasi Major 

Microgale thornasi Major, Ann. Mag. Nat. Hist. (6) 18: 320, Oct. 1896. Ampi- 
tambc Forest, northeastern Betsileo, Madagascar. 

Nesogale Thomas 

Nesogale Thomas, Ann. Mag. Nat. Hist. (9) 1: 303, Apr. 1918. Genotype 

Microgale dobsoni Thomas. 
Microgale Thomas, Ann. Mag. Nat. Hist. (5) 14: 337, Nov. 1884 (in part). 

Nesogale dobsoni (Thomas) 

Microgale dobsoni Thomas, Ann. Mag. Nat. Hist. (5) 14: 337, Nov. 1884. Nan- 
desen Forest, central Betsileo, Madagascar. 

Nesogale talazaci (Major) 

Microgale talazaci Major, Ann. Mag. Nat. Hist. (6) 18: 320, Oct. 1896. Forest 

of the Independent Tanala of Ikongo, near Vinanitelo, one day's journey 

south of Fianarantsoa, southern Betsileo, Madagascar. 

Nesoryctes Thomas 

Oryzorictes Milne-Edwards and A. Grandidier, Le Naturaliste, 4: 55, 1 Apl. 

1882 (not Grandidier). 
Nesoryctes Thomas, Ann. Mag. Nat. Hist. (9) 1: 307, Apr. 1918. Genotype 

Oryzoryctes tetradactylus Milne-Edwards and A. Grandidier. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS Zi 

Nesoryctes tetradactylus (Milne-Edwards and A. Grandidier) 
Oryzorictes tetradactylus Milne-Edwards and A. Grandidier, Le Naturaliste, 4: 

55, 1 Apl. 1882. Ampitambc, Sirabe, Imerina, Madagascar. 
Oryzorydes niger Major, Ann. Mag. Nat. Hist. (6) 18: 322, Oct. 1896. Marshes 

near Sirabe, Ampitambe, Madagascar. 

Oryzorictes A. Grandidier 

Oryzorictes A. Grandidier, Rev. et Mag. de Zool. (2) 22: 50, 1870. Genotype 

Oryzorictes hova A. Grandidier. 
Oryzoryctes Trouessart, Rev. et Mag. de Zool. (3) 7: 275, 1879. 

Oryzorictes hova A. Grandidier 

Oryzorictes hova A. Grandidier, Rev. et Mag. de Zool. (2) 22: 50, 1870. Ankaye 
and Antsianak, Madagascar. 

Oryzorictes talpoides G. Grandidier and Petit 

Oryzoryctes talpoides G. Grandidier and Petit, Bull. Mus. d'Hist. Nat. Paris, (2) 
2: 498, June 1930. Rizieres de Marovoay, Province de Majunga, Mada- 
gascar. 

Paramicrogale G. Grandidier and Petit 

Paramicrogale G. Grandidier and Petit, Bull. Soc. Zool. de France, 56: 127, 
15 June 1931. Genotype Paramicrogale occidentalis G. Grandidier and 
Petit. 

Paramicrogale decaryi (G. Grandidier) 

Microgale decaryi G. Grandidier, Bull. Mus. d'Hist. Nat. Paris, 34: 69, 1928. 

Grotte d'Andrahomana, south of Fort-Dauphin, Madagascar. 
Paramicrogale decaryi G. Grandidier and Petit, Bull. Soc. Zool. de France, 56: 

138, 15 June 1931. 

Paramicrogale occidentalis G. Grandidier and Petit 

Paramicrogale occidentalis G. Grandidier and Petit, Bull. Soc. Zool. de France, 
56: 129, 3 text-f., 15 June 1931. Region of Maintirano, Madagascar. 

Setifer Froriep. Madagascan Hedgehog 

Erinaceus Schreber, Saugth., 3: 583,1777 (in part). 

Setiger E. Geoffroy, Catal. Mamm. Mus. National d'Hist. Nat., Paris, p. 72, 

1803 (in part). 
Setifer Froriep, in Dumeril, Analit. Zool. mit Zusatzen, p. 15, 1806. Genotype 

Erinaceus setosus Schreber. 
Centetes Illiger, Prodr. Syst. Mamm. et Avium, p. 124, 1811 (in part). 
Centenes Cuvier, Regne Anim., ed. 1, 1: 136, 1817 (in part). 
Ericulus I. Geoffroy, Ann. des Sci. Nat., zool. (2) 8: 60, 1837. No type designated. 



28 bulletin: museum of comparative zoology 

Setifer setosus (Schreber) 

Erinaceus setosus Schreber, Siiugth., 3: 583, 590, pi. 164, 1777. Madagascar. 
Erinaceus acanthurus Boddsert, Elenchus Anim., p. 129, 1785. Madagascar. 
Centenes spinosus Desmarest, Encyclop. Methodique, Mammalogie, 1: 162, 

1820. Madagascar. 
Ericulus nigrescens I. Geoffroy, Mag. de Zool., Mammiferes, (2) 1: 29, 32, pi. 

3, 4, 1839. Madagascar. 
Ericulus nigricans Bartlett, Proc. Zool. Soc. London, p. 64, June 1875 (nomen 

nudum). 
Setifer madagascariensis Zimmermann, Specimen Zool., p. 503, 1777 (in part). 



Tenrec Lacepede. The Tenrec 

Tenrec Lacepede, Tabl. Mamm., p. 7, 1799. Genotype Erinaceus ecaudalus 

Schreber. 
Tenrecus Desmarest, Encyclop. Methodique, Mammalogie, 1: 161, 1820 (in 

synonymy). 

Tenrec ecaudatus (Schreber) 

Erinaceus ecaudatus Schreber, Saugth., 3: 584, pi. 165, 1777. Madagascar. 

Tenrec madagascariensis Zimmermann, Specimen Zool., p. 503, 1777 (in part). 

Erinaceus tendrec Borowski, Gemeinniizzige Naturgesch. d. Thierreichs, (Ber- 
lin), 1: pt. 2, 98, 1780. East Indies and Madagascar. "It does not seem 
possible to identify the animal" (Kellogg, R., Proc. Biol. Soc. Washington, 
45: 147, 9 Sep. 1932). 

Erinaceus tanrec Boddsert, Elenchus Anim., p. 129, 1785. 

Centetes armatus I. Geoffroy, Ann. des Sci. Nat., zool., (2) 8: 60, 1837. Mada- 
gascar. 
Range: Comoros, Nosi Be, Madagascar. 



Family POTAMOGALID^ 

Potamogale DuChaillu 

Potamogale DuChaillu, Proc. Boston Soc. Nat. Hist., 7: 363, Nov. 1860. 
Genotype Cynogale velox DuChaillu. 

Cynogale DuChaillu, Proc. Boston Soc. Nat. Hist., 7: 361, 1860 (in part, not 
Gray). 

Mythomys Gray, Proc. Zool. Soc. London, for 1861, p. 275, Apl. 1862. Geno- 
type Cynogale velox DuChaillu. 

Bayonia Bocage, Mem. Acad. Sci., Lisbon, 1 class, 4: 1, Mem. 2, p. 3, 1865; 
Proc. Zool. Soc. London, p. 402, 2 text-f. (skull), Oct. 1865. Genotype 
Potamogale velox DuChaillu. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 29 

POTAMOGALE VELOX DuChaillu 

Potamogale velox DuChaillu, Proc. Boston Soc. Nat. Hist., 7: 361, 1860. 

Equatorial Africa = Gaboon. 
P[otamogale] allmani Jentink, Notes Leyden Mus., 16: 236, Mar. 1895. Old 

Calabar. Indistinguishable from typical velox (Allen, J. A., Bull. Amer. 

Mus. Nat. Hist., 47: 8, 20 July 1922). 
Potamogale velox argens Thomas, Ann. Mag. Nat. Hist. (8) 16: 470, Dec. 1915. 

Medje, Congo Beige (see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 

6, 20 July 1922, by whom this is shown to be identical with the typical 

form). 
Range: Cameroons to Angola and Congo Beige. 



Geogale Milne-Edwards and A. Grandidier 

Geogale Milne-Edwards and A. Grandidier, Ann. des Sci. Nat., zool. (5) 15: 
1, 1872. Genotype Geogale aiirita Milne-Edwards and A. Grandidier. 

Geogale aurita aurita Milne-Edwards and A. Grandidier 
Geogale aurita Milne-Edwards and A. Grandidier, Ann. des Sci. Nat., zool. (5) 
15: 1, 1872. Tulear and Murundava, western Madagascar. 

Geogale aurita orientalis G. Grandidier and Petit 

Geogale aurita orientalis G. Grandidier and Petit, Faune des Colonies Fran^aises, 
4: no. 4, 446, 1930. Fenerive, Madagascar. 



Family SORICJD.E. Shrews 

Crocidura Wagler. White-toothed Slirews 

Crocidura Wagler, Oken's Isis, p. 275, 1832. Genotype Sorex leucodon Her- 
mann. 

Rhinomus Murray, Proc. Roy. Phys. Soc. Edinburgh, 2: 159, 1860. Genotype 
Rhinornus soricoides Murray = ? 

Leucodon Fatio, Faune Verteb. Suisse, 1: 132, 1869. Genotype Leucodon 
microurus Fatio = Sorex leucodon Hermann. 

Paurodus Schulze, Helios, Abh. u. Vortrage aus Gesammtgebiete Naturwiss., 
14: 90, 1897. As a subgenus of Crocidura; included Sorex leucodon Her- 
mann and S. araneus Schreber ( = Crocidura russula fide Miller) of which 
the former is here designated the type. 

Heliosorex Heller, Smithsonian Misc. Coll., 56: no. 15, p. 6, pi. 1, 23 Dec. 1910. 
Genotype Heliosorex roosevelti Heller. A synonym of Crocidura, fide Hol- 
hster, Bull. U. S. Nat. Mus., no. 99, pt. 1, p. 68, 1918. 



30 bulletin: museum of comparative zoology 

Review and keys: Thomas, O. On shrews from British East Africa. Ann. 
Mag. Nat. Hist. (7) 14: 236-241, Sep. 1904. 

Dollman, Guy. On the African shrews belonging to the genus Croddura. 
Ann. Mag. Nat. Hist. (8) 15: 507-527, May 1915; 562-575, June 1915; (8) 
16: 66-80, July 1915; 124-146, Aug. 1915; 357-380, Oct. 1915; 506-514, Dec. 
1915; (8) 17: 188-209, Feb. 1916. 

Note — Crossopus fodiens is credited to Algeria by Loche (Explor. Sci. de 
I'Algerie, ZooL, Mammiferes, p. 88, 1867), but no doubt erroneously. Sorex 
araneus, S. ie.tr agonurus, Croddura leucodon, listed by Loche (Explor. Sci. 
de I'Algerie, pp. 84, 86, 87, 88, 1867) are probably C. r. agilis or other 
species of Croddura. 

Crocidura albicatjda Peters 

Croddura albicauda Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 1866: 
885, ?1867. Angasilia, Comoro Islands. 

Crocidura allex allex Osgood 

Croddura allex Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: no. 3, p. 20, 
7 Apl. 1910. Naivasha, Kenya Colony. 

Crocidura allex alpina Heller 

Croddura alpina Heller, Smithsonian Misc. Coll., 56: no. 9, p. 5, 22 July 1910. 

West slope of Mt. Kenya, 10,000 feet, Kenya Colony. 
Croddura allex alpina Hollister, Bull. U. S. Nat. Mus., no. 99: pt. 1, p. 68, 1918. 

Crocidxjra anchiet^ Bocage 

Croddura anchietse Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 1: 26, 
Mar. 1889. Caconda, Angola. 

Crocidura ansorgei Dollman 

Croddura ansorgei TioWmsin, Ann. Mag. Nat. Hist. (8) 15: 511, May; 16: 138, 
Aug. 1915. Duque de Braganga, Angola. 

Crocidura arethusa Dollman 

Croddura arethusa Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 16: 144, 
Aug. 1915. Kabwir, Bauchi Province, northern Nigeria. 

Crocidura argentata (Sundevall) 

Sorex argentaius Sundevall, in Grill, Kongl. Svenska Vet.-Akad. Handl., Stock- 
holm, (2) 2: no. 10, p. 16, footnote, 1860. Roodeval, Karroo, South Africa. 

Crocidura aridula Thomas and Hinton 

Croddura aridula Thomas and Hinton, Proc. Zool. Soc. London, p. 252, July 6, 
1923. Kulme, Wadi Aribo, Darfur, 3300 feet. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 31 

Crocidtjra baileyi Osgood 

Crocidura baileyi Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 225, 28 Dec. 
1936. Ras Dashan (Mt. Geech), Simien Mts., Ethiopia. 
For Alfred M. Bailey, coll. 

Crocidura batesi Dollman 

Crocidura batesi Dollman, Ann. Mag. Nat. Hist. (8) 15: 510, May; 16: 143, 
Aug. 1915. Corao River, Gaboon. 

Crocidura beirje BEiRiE Dollman 

Crocidura beirse Dollman, Ann. Mag. Nat. Hist. (8) 15: 511, May; 16: 69, July 
1915. Beira, Portuguese East Africa. 

Crocidura beir^ nyik2e Dollman 

Crocidura beirse nyikse Dollman, Ann. Mag. Nat. Hist. (8) 15: 512, May; 16: 70, 
July 1915. Nyika plateau, northern Nyasaland, 6-7000 feet. 

Crocidura beta Dollman 

Crocidura beta Dollman, Ann. Mag. Nat. Hist. (8) 15: 513, May; 16: 78, July 

1915. "Charnia" River, Kenya Colony; = Chania River, near Nyeri and 

Fort Hall. 

Crocidura bicolor bicolor Bocage 

Crocidura bicolor Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 1 : 29, Mar. 
1889. "Gambos, dans I'interieur de Mossamedes", Angola. 

Crocidura bicolor cuninghamei Thomas 

Crocidura cuninghamei Thomas, Ann. Mag. Nat. Hist. (7) 14: 240, Sep. 1904. 
Island one mile north of Sajitu Id., Victoria Nyanza. 

Crocidura bicolor elgonius Osgood 

Crocidura bicolor elgonius Osgood, Ann. Mag. Nat. Hist. (8) 5: 369, Apl. 1910. 
Earui, Mt. Elgon, Kenya Colony. 

Crocidura bicolor hendersoni Dollman 

Crocidura bicolor hendersoni Dollman, Ann. Mag. Nat. Hist. (8) 15: 517, May 
1915; 17: 189, Feb. 1916. Nyasaland. 

Crocidura bicolor nana Dobson 

Crocidura nana Dobson, Ann. Mag. Nat. Hist. (6) 5: 225, Mar. 1890. Dollo, 

Somaliland. 
Crocidura bicolor nana Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 229, 28 

Dec. 1936. 



32 bulletin: museum of comparative zoology 

Crocidura bicolor nanilla Thomas 

Crocidura nanilla Thomas, Ann. Mag. Nat. Hist. (8) 4: 99, Aug. 1909. Prob- 
ably Entebbe, Uganda. 

According to Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 230, 28 Dec. 
1936) this is probably a race of C. bicolor. 

Crocidura nancilla St. Leger, Ann. Mag. Nat. Hist. (10) 9: 240, Mar. 1932 
(lapsus cal. for nanilla). 

Crocidura bicolor planiceps Heller 

Crocidura planiceps Heller, Smithsonian Misc. Coll., 56: no. 15, p. 5, 23 Dec. 

1910. Rhino Camp, Lado Enclave. 
Crocidura bicolor planiceps Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 1, p. 67, 

1918. 

Crocidura bicolor rudolfi St. Leger 

Crocidura denti St. Leger, Ann. Mag. Nat. Hist. (10) 9: 240, Mar. 1932. Koliok- 

well River, Lake Rudolf, Uganda (not C. jacksoni denti DoUman, 1915). 
Crocidura rudolfi St. Leger, Ann. Mag. Nat. Hist. (10) 10: 487, Nov. 1932 (in 

place of C. denti St. Leger, not C. jacksoni denti DoUman, 1915). Koliok- 

well River, Lake Rudolf, ITganda. 

According to Osgood (Zool Ser. Field Mus. Nat. Hist., 20: 230, 28 Dec. 

1936) this is probably a race of C. bicolor. 

Crocidura bicolor woosnami Dollman 

Crocidura bicolor woosnami Dollman, Ann. Mag. Nat. Hist. (8) 15: 516, May 
1915; 17: 189, Feb. 1916. Lake Ngami, Bechuanaland, South Africa. 

Crocidura bolivari Morales Agacino 

Crocidura bolivari Morales Agacino, Bol. Soc. Espan. Hist. Nat., Madrid, 34: 
93, f. 1, 1 Apl. 1934. Villa Cisneros, Rio de Ore. 

Crocidura bottegi Thomas 

Crocidura {Cr.) bottegi Thomas, Ann. Mus. Civ. Stor. Nat. Genova, (2): 18, 

677, 21 Feb. 1898. Northeast of Lake Rudolf, Abyssinia, between Badditu 

and Dime. 

Crocidura bovei Dobson 

Crocidura bovei Dobson, Ann. Mus. Civ. Stor. Nat. Genova, (2) 5: 425, fig., 25 
Oct. 1887. Vivi, Lower Congo, Congo Beige. 

Crocidura boydi Dollman 

Crocidura boydi Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 16: 357, 
Oct. 1915. Titebbi, Uele River. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 33 

Crocidura buttikoferi Jentink 

Crocidura buttikoferi Jentink, Notes Leyden Mus., 10: 47, Apl. 1888. Roberts- 
port, Liberia. 

Crocidura butleri Thomas 

Crocidura butleri Thomas, Ann. Mag. Nat. Hist. (8) 8: 375, Sep. 1911. Be- 
tween Chakchak and Dem Zubeir, Bahr-el-Ghazal, Sudan. 

Crocidura caliginea HoUister 

Crocidura caliginea HoUister, Bull. Amer. Mus. Nat. Hist., 35: 664, pi. 7, f. 1; 
pi. 8, ff. 1, la, 21 Oct. 1916. Medje, Congo Beige. 

Crocidura capensoides (A. Smith) 

Sorex capensoides A. Smith, South African Quart. Journ., 2: 62, 1833. Near 
Cape Town, South Africa. 

Crocidura chitauensis Hill and Carter 

Crocidura chitauensis Hill and Carter, Amer. Mus. Novitates, no. 937, p. 3, 
13 July 1937. Chitau, Angola, 4930 feet, 

Crocidura Cinderella Thomas 

Crocidura Cinderella Thomas, Ann. Mag. Nat. Hist. (8) 8: 119, July 1911; 
GemenjuUa, French Gambia. 

Crocidura congobelgica HoUister 

Crocidura congobelgica HoUister, Bull. Amer. Mus. Nat. Hist., 35: 670, pi. 7, f. 
6; pi. 9, f. 3, 3a, 21 Oct. 1916. LubUa, Bafwasende, Congo Beige. 

Crocidura crossei Thomas 

Crocidura (Cr.) crossei Thomas, Ann. Mag. Nat. Hist. (6) 16: 53, July 1895. 
Asaba, 150 miles up the Niger. 

Crocidura cuanzensis HiU and Carter 

Crocidura cuanzensis HiU and Carter, Amer. Mus. Novitates, no. 937, p. 2, 13 
July 1937. Chitau, Angola, 4930 feet. Near to C. katharina. 

Crocidura cyanea (Duvernoy) 

Sorex cyaneus Duvernoy, Mem. Soc. d'Hist. Nat. Strasbourg, 2: pt. 3, paper 
LL, p. 2, 1838; Mag. de Zool., 1842, p. 21, pi. 40, 41. "La riviere des Ele- 
phants, au sud de TAfrique." 

Sorex infumatus Wagner, Schreber's Saugthiere, Suppl., 2: 76, 1841; 5: 562, 
1855. Cape of Good Hope. 



34 bulletin: museum of comparative zoology 

Crocidura darfurea Thomas and Hinton 

Croddura darfurea Thomas and Hinton, Proc. Zool. Soc. London, p. 251, 
6 July 1923. Zalingei, Darfur. 

Crocidura deserti Schwann 

Crocidura deserti Schwann, Proc. Zool. Soc. London, p. 103, 7 June 1906. 
Molopo River, west of Morokwen, Bechuanaland. 

Crocidura dolichura Peters 

Crocidura (Croc.) dolichura Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 
p. 475, pi. 2, f. 1, 1876. Bonjongo, Cameroons. 

Crocidura electa Dollman 

Crocidura electa Dollman, Ann. Mag. Nat. Hist. (8) 5: 175, Feb. 1910. Kam- 
toby, south of Lake Tanganyika, Rhodesia. 

Crocidura erica Dollman 

Crocidura erica Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 16: 145, 
Aug. 1915. Pungo Andongo, Angola. 

Crocidura ferruginea Heuglin 

Crocidura ferruginea Heuglin, Leopoldina, 5: 36, June 1865, in Nova Acta 

Acad. Cses. Leop.-Carol., Dresden, 1865. "Lande der Ridj-neger", 

Bahr-el-Ghazal, Anglo-Egyptian Sudan. 
Probably indeterminable (Dollman). 

Crocidura fischeri Pagenstecher 

Crocidura fischeri Pagenstecher, Jahrb. Wiss. Anst., Hamburg, 2: 34, pi. 1, f. 
1-3, 1885. Nguruman, Tanganyika Territory. 

Crocidura flavescens (I. Geoff roy) 

Sorex flavescens I. Geoffroy, Dictionnaire Class., 11: 324, 1827; Mem. Mus. 
d'Hist. Nat. Paris, 15: 126, 1827. "La Cafrerie et le pays des Hotten- 
tots." 

Sorex capensis A. Smith, South African Quart. Journ., 2: 62, 1833 (not Smuts; 
not Geoffroy); 111. Zool. South Africa, Mamm., pi. 45, fig. 1, Mar. 1845. 

Sorex cinnamomeus Lichtenstein, Verb. Ges. Naturf. Freunde, Berlin, 2: 381, 
1829. 

Sorex rutilus Sundevall, Ofversigt af Kongl. Vet.-Akad. Forhandl., Stockholm, 
3: no. 4, p. 119, note, 1846. Port Natal. 

(New name for S. flavescens Smith, 111. Zool. So. Afr., descr. to PI. 45, but 
not S. flavescens I. Geoff., which has much shorter tail.) 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 35 

Crocidura floweri DoUman 

Crocidura floweri Dollman, Ann. Mag. Nat. Hist. (8) 15: 515, May 1915; 17: 
192, Feb. 1916. Gizeh, Egypt. 

Crocidura foxi Dollman 

Crocidura foxi Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 16: 143, Aug. 
1915. Panyam, northern Nigeria, 4000 feet. 

Crocidura fulvastra (Sundevall) 

Sorex fulvaster Smidevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 

1842, p. 172, 1843. Bahr-el-Abiad, Sudan. 
Crocidura strauchii Dobson, Ann. Mag. Nat. Hist. (6) 5: 225, Mar. 1890. 

Sudan. According to Dollman and to Osgood (Zool. Ser. Field Mus. Nat. 

Hist., 20: 227, 28 Dec. 1936) this is probably a synonym of C. fulvastra. 

Crocidura fumosa fumosa Thomas 

Crocidura Jumosa Thomas, Ann. Mag. Nat. Hist. (7) 14: 238, Sep. 1904. 

Western slope of Mt. Kenya, Kenya Colony, 2600 m. 
Crocidura alchemillx Heller, Roosevelt's African Game Trails, Amer. ed., p. 

480; London ed., p. 491, 1910. Summit of Aberdare range, Kenya Colony. 
Range: Central Kenya Colony. 

Crocidura fumosa johnstoni Dollman 

Crocidura fumosa johnstoni Dollman, Ann. Mag. Nat. Hist. (8) 15: 510, May; 
16: 372, Oct. 1915. Chiromo, Nyasaland. 

Crocidura fumosa macmillani Dollman 

Crocidura luna macmillani Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 

16: 360, Oct. 1915. Kotelee, Walamo, Abyssinia. 
Crocidura fumosa macmillani Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 227^ 

28 Dec. 1936. 

Crocidura fumosa montis Thomas 

Crocidura fumosa montis Thomas, Ann. Mag. Nat. Hist. (7) 18: 138, Aug. 1906. 
Ruwenzori East, Uganda, 12,500 feet. 

Crocidura fumosa schistacea Osgood 

Crocidura fumosa schistacea Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 
no. 3, p. 20, 7 Apl. 1910. Lukenya Mountain, Kenya Colony. 

Crocidura luna umbrosa Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 16: 
360, Oct. 1915. Machakos, Kenya Colony, 5400 feet. According to Os- 
good (Zool. Ser. Field Mus. Nat. Hist., 20: 228, 28 Dec. 1936) this is a 
synonym of C. f. schistacea. 



36 bulletin: museum of comparative zoology 

Crocidura fumosa selina Dollman 

Crocidura fumosa selina Dollman, Ann. Mag. Nat. Hist. (8) 15: 510, May; 16: 
370, Oct. 1915. Mabira Forest, Chagwe, Uganda. 

Crocidura fusco-murina (Heuglin) 

S[orex\ fusco murinus Heuglin, Leopoldina, 5: 36, June 1865, in Nova Acta 
Acad. Cses. Leop.-Carol., Dresden, 1865. Meshra-el-Req, Bahr-el-Ghazal, 
Anglo-Egyptian Sudan. Probably indeterminable (Dollman). 

Crocidura fuscosa Thomas 

Crocidura doriana fuscosa Thomas, Ann. Mag. Nat. Hist. (8) 12: 90, July 1913. 
Kaka, White Nile. 

Crocidura geoffroyi (J. B. Fischer) 

Sorex geoffroyi J. B. Fischer, Synopsis Mamm., Addenda, p. 664, 1830. Mauri- 
tius ("in insula Francige"). 

Crocidura giffardi de Winton 

Crocidura (Cr.) giffardi de Winton, Ann. Mag. Nat. Hist. (7) 2: 484, Dec. 1898. 

Moshi (or Mossi) near Wagadugu, 500 miles northeast of Kumasi, Gold 

Coast. 

Crocidura glebula Dollman 

Crocidura glebula Dollman, Ann. Mag. Nat. Hist. (8) 15: 517, May 1915; 17: 
196, Feb. 1916. Zungeru, northern Nigeria. 

Crocidura gracilipes Peters 

Crocidura (Cr.) gracilipes Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 
p. 590, 1870. Kilimanjaro, Tanganyika Territory. 

Crocidura hedenborgiana (Sundevall) 

Sorex hedenhorgianus Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stock- 
holm, for 1842, p. 171, 1843. Sennaar. 

Sorex hedenborgi Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 
for 1842, p. 177, 1843 (lapsus), also on p. 199. 

Crocidura hildegarde^ hildegarde^ Thomas 

Crocidura hildegardese Thomas, Ann. Mag. Nat. Hist. (7) 14: 240, Sep. 1904. 

Fort Hall, Kenya Colony. 
Crocidura maanjai Heller, Smithsonian Misc. Coll., 56: no. 15, p. 4, 23 Dec. 

1910. Kabula Muliro, Uganda. 
Crocidura lutreola Heller, Smithsonian Misc. Coll., 60: no. 12, p. 8, 4 Nov. 1912. 

Mt. Mbololo, Taita Hills, Kenya Colony. 
Crocidura hildegardese procera Heller, Smithsonian Misc. Cbll., 60: no. 12, p. 10, 

4 Nov. 1912. Mt. Lololokui, northern Guaso Nyiro, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 37 

Crocidura macowi Dollman, Ann. Mag. Nat. Hist. (8) 15: 515, May; 16: 378, 
Oct. 1915. Mt. Nyiro, south of Lake Rudolf, Kenya Colony. According 
to Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 229, 28 Dec. 1936) this 
is a synonym of C. h. hildegardese. 

Crocidura ibeana Dollman, Ann. Mag. Nat. Hist. (8) 15: 514, May; 16: 362, 
Oct. 1915. Olgerei River, Kenya Colony. According to Osgood (Zool, 
Ser. Field Mus. Nat. Hist., 20: 229, 28 Dec. 1936) this is a synonym of 
C. h. hildegardex. 

Crocidura hildegarde^e alt^ Heller 

Crocidura hildegardese altas Heller, Smithsonian Misc. Coll., 60: no. 12, p. 9, 
4 Nov. 1912. Mt. Garguez, Mathews Range, Kenya Colony, 6000 feet. 

Crocidura hildegarde^ ph^ura Osgood 

Crocidura hildegardeas phseura Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 228, 

28 Dec. 1936. West base of Mt. Guramba, northeast of AUata, Sidamo, 

Ethiopia. 

Crocidura hildegarde^ rubecula Dollman 

Crocidura hildegardese rubecula Dollman, Ann. Mag. Nat. Hist. (8) 15: 515, 
May; 16: 509, Dec. 1915. Kigezi, Uganda, 6000 feet. 

Crocidura hindei hindei Thomas 

Crocidura hindei Thomas, Ann. Mag. Nat. Hist. (7) 14: 237, Sep. 1904. Mach- 
akos, Kenya Colony. 

Crocidura hindei diana Dollman 

Crocidura hindei diana Dollman, Ann. Mag. Nat. Hist. (8) 15: 512, May; 16: 
76, July 1915. Lake Chad. 

Crocidura hindei marrensis Thomas and Hinton 

Crocidura hindei marrensis Thomas and Hinton, Proc. Zool. Soc. London, p. 
252, 6 July 1923. Wadi Konger, east-central Jebel Marra, Darfur. 

Crocidura hirta hirta Peters 

Crocidura hirta Peters, Reise nach Mossambique, Saugeth., p. 78, pi. 18, f. 2, 

1852. Tette, Mozambique, 17° S. 
Crocidura canescens Peters, Reise nach Mossambique, Saugeth., p. 83, pi. 18, 

f. 4, 1852. Tette, Mozambique, 17° S. 
Crocidura annellata Peters, Reise nach Mossambique, Saugeth., p. 85, pi. 18, 

f. 5, 1852. Tette, Mozambique, 17° S. 

Crocidura hirta flavidula Thomas and Schwann 

Crocidura flavescens flavidula Thomas and Schwann, Proc. Zool. Soc. London, 
1 : 264, 10 Aug. 1905. Umvolosi, Zululand, Natal. 



38 bulletin: museum of comparative zoology 

Crocidura hirta mutest Heller 

CrodduramutesaeHeWeT, Smithsonian Misc. Coll., 56: no. 15, p. 3, 23 Dec. 1910. 

Kampala, Uganda. 
Crocidura hirta muiesse Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 225, 28 

Dec. 1936. 

Crocidura hirta pondoensis Roberts 

Crocidura pondoensis Roberts, Ann. Transvaal Mus., 4: 71, 20 Oct. 1913. 
Ngqeleni district, Pondoland, South Africa. 

Crocidura hirta suahel^ Heller 

Crocidura suahelx Heller, Smithsonian Misc. Coll., 60: no. 12, p. 6, 4 Nov. 1912. 

Mazeras, Kenya Colony. 
Crocidura hirta suahelx Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 225, 28 

Dec. 1936. 

Crocidura holobrunneus Roberts 

Crocidura holobrunneus Roberts, Ann. Transvaal Mus., 14: 225, 28 July 1931. 
Mariepskop, Pilgrimsrest district, Transvaal. 
Range: Transvaal to Zululand. 

Crocidura jacksoni jacksoni Thomas 

Crocidura jacksoni Thomas, Ann. Mag. Nat. Hist. (7) 14: 238, Sep. 1904. 
Ravine Station, Kenya Colony. 

Crocidura jacksoni amal^ DoUman 

Crocidura jacksoni amalse Dollman, Ann. Mag. Nat. Hist. (8) 15: 516, May; 
16: 376, Oct. 1915. Amala River, Nyanza Province, Kenya Colony. 

Crocidura jacksoni denti Dollman 

Crocidura jacksoni denti Dollman, Ann. Mag. Nat. Hist. (8) 15: 516, May; 16: 

377, Oct. 1915. Between Mawambi and Avakubi, Ituri Forest, Congo 

Beige. 

Crocidura katharina Kershaw 

Crocidura katharina Kershaw, Ann. Mag. Nat. Hist. (9) 10: 101, July 1922. 
N'dola, Northern Rhodesia, 12° 50' S., 28° 40' E. 

Crocidura langi Cabrera 

Crocidura langi Cabrera, Bol. Real Soc. Espanola Hist. Nat. Madrid, 25: 21, 
30 Jan. 1925. St. Joseph de Luluaberg, Kasai, Congo Beige. 

Crocidura latona HoUister 

Crocidura latona Hollister, Bull. Amer. Mus. Nat. Hist., 35: 667, pi. 7, f. 3; pi. 
8, f. 3, 3a, 21 Oct. 1916. Medje, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 39 

Crocidura littoralis Heller 

Crocidura littoralis Heller, Smithsonian Misc. Coll., 56: no. 15, p. 5, 23 Dec. 
1910. Butiaba, east shore of Albert Nyanza, Uganda. 

Crocidura ltjdia Hollister 

Crocidura India Hollister, BuU. Amer. Mus. Nat. Hist., 35: 668, pi. 7, f. 4; pi. 
9, f. 1, la, 21 Oct. 1916. Medje, Congo Beige. 

Crocidura luimbalensis Hill and Carter 

Crocidura luimbalensis Hill and Carter, Amer. Mus. Novitates, no. 937, p. 1, 
13 July 1937. Luimbale, Angola. 

Crocidura lulu^ Matschie 

Crocidura luluse Matschie, Zeitschr. f. Saugetierk., 1: 111, 24 Dec. 1926. 
Kapelekese, near Luluaberg, Lulua River, Congo Beige. 

Crocidura luna Dollman 

Crocidura luna Dollman, Ann. Mag. Nat. Hist. (8) 5: 175, Feb. 1910. Bunkeya 
River, Katanga, Congo Beige. 

Crocidura lusitania Dollman 

Crocidura lusitania Dollman, Ann. Mag. Nat. Hist. (8) 15: 516, May 1915; 
17: 198, Feb. 1916. Trarza country, Mauritania. 

Crocidura lutrella Heller 

Crocidura lutrella Heller, Smithsonian Misc. Coll., 56: no. 15, p. 4, 23 Dec, 1910. 
Rhino Camp, Lado Enclave. 

Crocidura macarthuri St. Leger 

Crocidura macarthuri St. Leger, Ann. Mag. Nat. Hist. (10) 13: 559, May 1934. 
Merifano, 20 miles from mouth of Tana River, Kenya Colony. 

Crocidura manni Peters 

Crocidura (Crocid.) manni Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 19, 
1878. Lagos, Nigeria. 
Range: Nigeria. 

Crociduiia mariquensis (A. Smith) 

Sorex mariquensis A. Smith, Illustr. Zool. So. Africa, Mamm., pi. 44, f. 1, Oct. 
1844. "Wood near the tropic of Capricorn," South Africa. 

Crocidura marita Thomas and Hinton 

Crocidura marita Thomas and Hinton, Proc. Zool. Soc. London, p. 253, 6 July 
1923. Southeast downs, Jebel Marra, Darfur. 



40 bulletin: museum of comparative zoology 

Crocidura martensii Dobson 

Crocidura martensii Dobson, Ann. Mag. Nat. Hist. (6) 6: 496, Dec. 1890. 
"Cape of Good Hope." 
Range: To Zululand. 

Crocidura martiensseni Neumann 

Crocidura martiensseni Neumann, Zool. Jahrb., Syst., 13: 544, 10 Oct. 1900. 
Mt. Kilimanjaro, above Marangu, Tanganyika Territory, 6000 feet. 

Crocidura maurisca maurisca Thomas 

Crocidura maurisca Thomas, Ann. Mag. Nat. Hist. (7) 14: 239, Sep. 1904. 
Entebbe, Uganda. 

Crocidura maurisca geata Allen and Loveridge 

Crocidura maurisca geata G. M. Allen and Loveridge, Proc. Boston Soc. Nat. 
Hist., 38: 417, Dec. 1927. Nyingwa, Uluguru Mts., Tanganyika Territory. 

Crocidura monax Thomas 

Crocidura monax Thomas, Ann. Mag. Nat. Hist. (8) 6: 310, Sept. 1910. Rombo, 
Mt. Kilimanjaro, Tanganyika Territory, 6000 feet. 

Crocidura muricauda (Miller) 

Myosorex muricauda Miller, Proc. Washington Acad. Sci., 2: 645, f. 42, 28 
Dec. 1900. Mount Coffee, Liberia. 

Crocidura neavei Wroughton 

Crocidura neavei Wroughton, Manchester Mem., 51: no. 5, p. 7, 13 Mar. 1907. 
Kafue River, Rhodesia, 4000 feet. 

Crocidura nigricans Bocage 

Crocidura nigricans Bocage, Jorn. Sci. Math., Phys. e Nat. Lisboa, (2) 1: 28 
Mar. 1889. Quindumbo, Angola. 

Crocidura nigrofusca Matschie 

Crocidura nigrofusca Matschie, Deutsch-Ost-Afrika, 3: Siiugethiere, p. 33, 
1895. Wukalala camp, Kinyawanga, west of Semliki, Congo Beige. 

Crocidura niobe Thomas 

Crocidura niobe Thomas, Ann. Mag. Nat. Hist. (7) 18: 138, Aug. 1906. Ruwen- 
zori East, Uganda. 

Crocidura occidentalis occidentalis (Pucheran) 

Pachyura occidentalis Pucheran, Rev. et Mag. de Zool., (2) 7: 154, 1855. 
Gaboon. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 41 

Pachyura iequatorialis Pucheran, Rev. et Mag. de Zool., (2) 7: 154, 1855. 
Gaboon. 

Croddura petersii Dobson, Ann. Mag. Nat. Hist. (6) 6: 495, Dec. 1890. Ga- 
boon. 

Crocidtjra occidentalis cara DoUman 

Croddura ocddentalis cara DoUman, Ann. Mag. Nat. Hist. (8) 15: 509, 525, 
May 1915. Jala, Sierra Leone. 

Crocidtjra occidentalis daphnia HoUister 

Croddura daphnia Hollister, Smithsonian Misc. Coll., 66: no. 8, p. 1, May 1916. 
Gondokoro, Uganda. 
According to Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 222, 28 Dec. 
1936) this is doubtfully distinct from Co. nyansae. 

Crocidura occidentalis doriana Dobson 

Croddura doriana Dobson, Ann. Mus. Civ. Stor. Nat., Genova, (2) 4: 564, 

24 May 1887. Let-Marefia, Shoa, Abyssinia. 
Croddura ocddentalis doriana Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 

220, 28 Dec. 1936 (with comment on limits of the ocddentalis group). 

Crocidura occidentalis gxjineensis Cabrera 

Croddura doriana guineensis Cabrera, Mem. Soc. Espanola Hist. Nat., Madrid, 
1: 22, pi. 1, May 1903. Cape San Juan, Rio Muni, Spanish Guinea. 

Crocidtjra occidentalis hera DoUman 

Crocidura nyansae hera DoUman, Ann. Mag. Nat. Hist. (8) 15: 509, May; 
571, June 1915. Shire highlands, Blantyre district, southern Nyasaland. 

Crocidura occidentalis herero St. Leger 

Crocidura nyanzse herero St. Leger, Ann. Mag. Nat. Hist. (10) 10: 85, July 
1932. Mbambi, Grootfontein-Caprivi border, Southwest Africa. 

Crocidura occidentalis kijab^ J. A. Allen 

Croddura kijabse J. A. Allen, Bull. Amer. Mus. Nat. Hist., 26: 173, 19 Mar. 
1909. Kijabe, Kenya Colony. 

Crocidura occidentalis kivu Osgood 

Croddura flavescens kivu Osgood, Ann. Mag. Nat. Hist. (8) 5: 370, Apl. 1910. 
Lake Kivu, eastern Congo. 

Crocidura occidentalis luluana Cabrera 

Crocidura ocddentalis luluana Cabrera, Bol. Real Soc. Espanola Hist. Nat., 

Madrid, 25: 20, 30 Jan. 1925. St. Joseph de Luluabourg, Kasai, Congo 

Beige. 



42 bulletin: museum of comparative zoology 

Crocidura occidentalis NIGERIA Dollmaii 

Crocidura occidentalis nigerise Dollman, Ann. Mag. Nat. Hist. (8) 15; 511, 524, 
May 1915. Asaba, 150 miles up the Niger. 

Crocidura occidentalis nyans^e Neumann 

Crocidura flavescens nyansse Neumann, Zool. Jahrb., Syst., 13: 544, 10 Oct. 1900. 

Fort Lubwa's in Ussoga, outlet of Victoria Nyanza, Uganda. 
Crocidura occidentalis nyansx Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 

220, 28 Dec. 1936. 

Crocidura occidentalis spurrelli Thomas 

Crocidura spurrelli Thomas, Ann. Mag. Nat. Hist. (8) 6: 427, Oct. 1910. 
Bibianaha, near Dunkwa, inland of Sekondi, Gold Coast. 

Crocidura occidentalis tatiana Dollman 

Crocidura nyansse tatiana Dollman, Ann. Mag. Nat. Hist. (8) 15: 509, May; 
570, June 1915. Mt. Gargues, north of Mweru district, Kenya Colony. 

Crocidura occidentalis zuleika Dollman 

Crocidura nyansse zuleika Dollman, Ann. Mag. Nat. Hist. (8) 15: 509, May; 
570, June 1915. Chirinda Forest, Melsetter district. Southern Rhodesia. 

Crocidura odorata (Leconte) 

Sorex odoratus Leconte, Proc. Acad. Nat. Sci. Philadelphia, p. 11, 1857. 
[Cette Cama, Gaboon.] 

Crocidura olivieri (Lesson) 

Sorex olivieri Lesson, Manuel de Mammalogie, p. 121, 1827. Sakkara, Egypt, 
as mummies from catacombs. 

Crocidura oritis Hollister 

Crocidura oritis Hollister, Bull. Amer. Mus. Nat. Hist., 35: 666, pi. 7, f. 2; 
pi. 8, f. 2, 2a, 21 Oct. 1916. Avakubi, Ituri River, Congo Beige. 

Crocidura parvipes parvipes Osgood 

Crocidura parvipes Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: no. 3, 
p. 19, 7 Apl. 1910. Voi, Kenya Colony. 

Crocidura parvipes nisa HoUister 

Crocidura parvipes nisa Hollister, Smithsonian Misc. Coll., 66: no. 8, p. 2, 
May, 1916. Kibabe, Kisumu, Kenya Colony. 

Crocidura pasha Dollman 

Crocidura pasha Dollman, Ann. Mag. Nat. Hist. (8) 15: 517, May 1915; 17: 
195, Feb. 1916. Atbara River, Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 43 

Crociduea percivali DoUman 

Crocidura -percivali DoUman, Ann. Mag. Nat. Hist. (8) 15: 513, May; 16: 126, 
Aug. 1915. Jombeni Range, Nyeri district, Kenya Colony, 3500 feet. 

Crocidura pilosa Dobson 

Crocidura pilosa Dobson, Ann. Mag. Nat. Hist. (6) 6: 496, Dec. 1890. Trans- 
vaal. 

Crocidura pitmani Barclay 

Crocidura pitmani Barclay, Ann. Mag. Nat. Hist. (10) 10: 440, Oct. 1932. 
Maluwe-Serenje district, Northern Rhodesia, 3800 feet. 

Crocidura poensis poensis (Fraser) 

Sorex (Crocidura) poensis Fraser, Proc. Zool. Soc. London, for 1842, 10: 200, 
Feb. 1843. Clarence, Fernando Po. 

Crocidura poensis attila Dollman 

Crocidura poensis attila Dollman, Ann. Mag. Nat. Hist. (8) 15: 512, May; 16: 
141, Aug. 1915. Bitye, Cameroons. 

Crocidura poensis pamela Dollman 

Crocidura poensis pamela Dollman, Ann. Mag. Nat. Hist. (8) 15: 510, May; 
16: 142, Aug. 1915. Bibianaha, Gold Coast. 
Range: Gold Coast to French Gambia. 

Crocidura poensis soricoides (Murray) 

Rhinomus soricoides Murray, Proc. Roy. Phys. Soc. Edinburgh, 2: 159, 1860. 
Old Calabar, southern Nigeria. 

Crocidura polia Hollister 

Crocidura polia Hollister, Bull. Amer. Mus. Nat. Hist., 35: 669, pi. 7, f. 5; 
pi. 9, f. 2, 2a, 21 Oct. 1916. Medje, Congo Beige. 

Crocidura pondoensis Roberts 

Crocidura pondoensis Roberts, Ann. Transvaal Mus., 4: 71, 20 Oct. 1913. 
Port St. John's, Pondoland, South Africa. 

Crocidura raineyi Heller 

Crocidura raineyi Heller, Smithsonian Misc. Coll., 60: no. 12, p. 7, 4 Nov. 1912. 
Mt. Gargues, Mathews Range, Kenya Colony, 6000 feet. 

Crocidura religiosa (I. Geoffroy) 

Sorex religiosus I. Geoffroy, Mem. Mus. d'Hist. Nat. Paris, 15: 128, 1827. 
Types mummified, Thebes, Egypt. 



44 bulletin: museum of compakative zoology 

Crocidura roosevelti (Heller) 

Heliosorex roosevelti Heller, Smithsonian Misc. Coll., 56: no. 15, p. 6, pi. 1, 

23 Dec. 1910. Rhino Camp, Lado Enclave. 
Crocidura roosevelti HoUister, Bull. U. S. Nat. Mus., no. 99, pt. 1, p. 68, 1918. 

Crocidura russula agilis (Levaillant). Algerian Pygmy Shrew 

Pachyura agilis Levaillant, in Loche, Explor. Sci. de I'Algerie, Zool., pi. 4, f . 2, 
1850. Algeria. 

(The atlas of this work was published many years before the text accord- 
ing to Cabrera.) 

Sorex mauritanicus Pomel, Compt. Rend. Acad. Sci. Paris, 42: 653, 1856. 
Algeria. Perhaps indeterminable. 

Pachyura -pigmsea Loche, Explor. Sci. de I'Algerie, Zool., p. 88, 1867. Ain-el- 
Bel, Algeria. 

Crocidura russula suaveolens Pallas (not Blasius), Trouessart, Cat. Mamm. 
Viv. Foss., Suppl., p. 142, 1904 (in part). 
Range: Tunis and eastern Morocco. 

Crocidura russula yebalensis Cabrera 

Crocidura yebalensis Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 
13: 400, Oct. 1913. Tetuan, Morocco. 

Crocidura sacralis Peters 

Crocidura sacralis Peters, Reise nach Mossambique, Saugeth., p. 82, pi. 18, 
f. 3, 1852. Cabaceira Peninsula, Mozambique. 

Crocidura sansibarica Neumann 

Crocidura bicolor sansibarica Neumann, Zool. Jahrb., Syst., 13: 544, 10 Oct. 
1900. Mojoni, Zanzibar Island. 

Crocidura schweitzeri Peters 

Crocidura (Crocidura) schweitzeri Peters, Monatsb. K. Preuss. Akad. Wiss., 

Berlin, p. 187, 1877. Liberia. 
Crocidura stampfiii Jentink, Notes Leyden Mus., 10: 47, Apl. 1888. Liberia, 

Schieffelinsville and Hill-town. 

Crocidura sericea (Sundevall) 

(Sorex sericeus Sundevall, Kongl. Svenska Vet.-Akad.Handl., Stockholm, for 
1842, pp. 173, 177, 1843. Near Bahr-el-Abiad, Sudan. 

Crocidura macrodon Dobson, Ann. Mag. Nat. Hist. (6) 5: 226, Mar. 1890. 
?Sudan. According to Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 227, 
28 Dec. 1936) this is best regarded as a synonym of C. sericea. 

Crocidura shortridgei St. Leger 

Crocidura shortridgei St. Leger, Ann. Mag. Nat. Hist. (10) 10: 84, July 1932. 
Popa Falls, West Caprivi, Damaraland, Southwest Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 45 

Crocidura silacea Thomas 

Crocidura {Cr.) silacea Thomas, Ann. Mag. Nat. Hist. (6) 16: 53, July 1895. 
Figtree Creek, De Kaap, Transvaal. 

Crocidura smithii Thomas 

Crocidura (Croc.) smithii Thomas, Ann. Mag. Nat. Hist. (6) 16: 51, July 1895. 
Webi Shebeli, Somaliland. 

Crocidura somalica Thomas 

Crocidura (Cr.) somalica Thomas, Ann. Mag. Nat. Hist. (6) 16: 52, July 1895. 
Webi Shebeli, Somahland. 

Crocidura surur^ Heller 

Crocidura sururse Heller, Smithsonian Misc. Coll., 56: no. 15, p. 2, 23 Dec. 1910. 
Rhino Camp, Lado Enclave. 
Probably a race of nyansse, apud HoUister, 1916. 

Crocidura sylvia Thomas and Schwann 

Crocidura sylvia Thomas and Schwann, Proc. Zool. Soc. London, p. 587, 10 Oct. 
1906. Woodbush, Zoutpansberg district, Transvaal. 

Crocidura thomensis (Socage) 

Sorex (Crocidura) thomensis Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, 
(1) 11: 213, Feb. 1887. Island of St. Thome. 

Crocidura turba turba Dollman 

Crocidura turba Dollman, Ann. Mag. Nat. Hist. (8) 5: 176, Feb. 1910. Chilui 
Island, Lake Bangweolo, Rhodesia. 

Crocidura turba angol^e Dollman 

Crocidura turba angolas Dollman, Ann. Mag. Nat. Hist. (8) 15: 510, May; 16: 
136, Aug. 1915. Bailundu country, Angola. 

Crocidura turba nilotica Heller 

Crocidura nilotica Heller, Smithsonian Misc. CoU., 56: no. 15, p. 3, 23 Dec. 1910. 

Rhino Camp, Lado Enclave. 
Crocidura turba nilotica Hollister, Bull. Amer. Mus. Nat. Hist., 35: 664, 21 

Oct. 1916. 
Range: Lado Enclave to 'Njabara, Ethiopia. 

Crocidura turba tarella Dollman 

Crocidura turba tarella Dollman, Ann. Mag. Nat. Hist. (8) 15: 511, May; 16: 
135, Aug. 1915. Chaya, near Ruchuru, Congo Beige. 



46 bulletin: museum of comparative zoology 

Crocidura turba zaodon Osgood 

Croddura turba zaodon Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 

no. 3, p. 21, 7 Apl. 1910. Nairobi, Kenya Colony. 
Croddura turba provocax Thomas, Ann. Mag. Nat. Hist. (8) 6: 112, July 1910. 

Aberdare Mts., Kenya Colony, 11,000 feet. 
Croddura turba lakiundse Heller, Smithsonian Misc. Coll., 60: no. 12, p. 6, 

4 Nov. 1912. Lakiundu River near junction with Northern Guaso Nyiro, 

Kenya Colony. 
Croddura turba kempi Dollman, Ann. Mag. Nat. Hist. (8) 15: 511, May; 16: 

134, Aug. 1915. Kirui, Mt. Elgon, Kenya Colony, 6000 feet. 

Crocidura ultima Dollman 

Croddura ultima Dollman, Ann. Mag. Nat. Hist. (8) 15: 517, May 1915; 17: 
204, Feb. 1916. Jombeni Range, Nyeri district, Kenya Colony, 5000 ft. 

Crocidura velutina Thomas 

Croddura velutina Thomas, Ann. Mag. Nat. Hist. (7) 14: 237, Sep. 1904. 
Usambara, Tanganyika Territory. 

Crocidura viaria (I. Geoffroy) 

Sorex viarius I. Geoffroy, Voyage aux Indes-Orientales par C. Belanger, Zool., 
p. 127, 1834. Senegal. 

Crocidura voi Osgood 

Croddura voi Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: no. 3, p. 18, 
7 Apl. 1910. Voi, Kenya Colony. 

Crocidura whitakeri De Winton 

Croddura (Cr.) whitakeri De Winton, Proc. Zool. Soc. London, for 1897, p. 954, 

Apl. 1898. Sierzet, halfway between Morocco City and Mogador, Morocco . 
Croddura whitakeri Cabrera, Trab. Mus. Nac. Cienc. Nat., Madrid, zool. ser., 

no. 57, p. 65, pi. 3 (col.), 30 Dec. 1932. 

Crocidura xantippe Osgood 

Croddura xantippe Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: no. 3, 

p. 19, 7 Apl. 1910. Voi, Kenya Colony. 
Croddura Xanthippe Dollman, Ann. Mag. Nat. Hist. (8) 15: 512; 16: 375, 1915. 

Crocidura zaphiri zaphiri Dollman 

Croddura zaphiri Dollman, Ann. Mag. Nat. Hist. (8) 15: 509, May; 16: 66, 
July, 1915. Charada Forest, Kaffa, Abyssinia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 47 

Crocidura zaphiri siMioLtrs Hollister 

Crocidura simiolus Hollister, Smithsonian Misc. Coll., 66: no. 8, p. 3, May 1916. 

Kisumu, Kenya Colony. 
Crocidura zaphiri simiolus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 224, 

28 Dec. 1936 (with note on status). 

Crocidura zena zena DoUman 

Crocidura zena DoUman, Ann. Mag. Nat. Hist. (8) 15: 511, May; 16: 137, 
Aug. 1915. Chilui Island, Lake Bangweolo, Rhodesia. 

Crocidura zena cabrerai Morales Agacino 

Crocidura zena cabrerai Morales Agacino, Bol. Soc. Espanola Hist. Nat., 

Madrid, 35: 20, fig. 1, 5 Mar. 1935. St. Joseph de Luluabourg, Kasai, 

Congo Beige. 

Crocidura zimmeri Osgood 

Crocidura zimmeri Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 223, 28 Dec. 

1936. Katobwe, near Bukoma, Lualaba River, Katanga Province, Congo 

Beige. 



Myosorex Gray 

Myosorex Gray, Proc. Zool. Soc. London, for 1837, p. 124, 14 June 1838. 

Genotype Sorex varius Smuts. 
Sorex Smuts, Enum. Mammal. Cap., p. 8, 1832 (not Linnaeus). 

Myosorex blarina Thomas 

Myosorex blarina Thomas, Ann. Mag. Nat. Hist. (7) 18: 139, Aug. 1906; 
Thomas and Wroughton, Trans. Zool. Soc. London, 19: 489, pi. 20, 
lower fig.. Mar. 1910. Mubuku Valley, Ruwenzori East, LTganda, 10,000 
feet. 

Myosorex sclateri sclateri Thomas and Schwann 

Myosorex sclateri Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 15, 

p. 10, 28 Feb. 1905; Proc. Zool. Soc. London, 1: 131, 1 June 1905. Ngoye 

Hills, Zululand, South Africa, 250 meters. 

Myosorex sclateri affinis Thomas and Schwann 

Myosorex sclateri affinis Thomas and Schwann, Proc. Zool. Soc. London, 1: 
262, 263, 10 Aug. 1905. Sibudeni, Zululand, Natal, 1700 meters. 



48 bulletin: museum of comparative zoology 

Myosorex sclateri talpinus Thomas and Schwann 

Myosorex sclateri talpinus Thomas and Schwann, Proc. Zool. Soc. London, 1: 
262, 263, 10 Aug. 1905. Umvolosi, Zululand, Natal, 60 meters. 

Myosorex swinnyi Chubb 

Myosorex swinnyi Chubb, Ann. Transvaal Mus., 1: suppl., p. 1, Oct. 1908; 
182, Jan. 1909. Port St. John's district, western Pondoland, South Africa. 

Myosorex tenuis Thomas and Schwann 

Myosorex tenuis Thomas and Schwann, Proc. Zool. Soc. London, 1: 131, 132, 
1 June 1905. Zuurbron, Wakkerstroom Highlands, Transvaal. 

Myosorex transvaalensis Roberts 

Myosorex transvaalensis Roberts, Ann. Transvaal Mus., 10: 61, 31 Jan. 1924. 
Koster, on edge of "high veldt," Transvaal. 

Myosorex varius varius (Smuts) 

Sorex varius Smuts, Enumer. Mamm. Cap., p. 108, 1832. Algoa Bay, Cape of 

Good Hope. 
Sorex capensis Smuts, Enumer. Mamm. Cap., p. 8, 1832 (nee A. Smith, nee 

I. Geoffroy). Cape of Good Hope. 
Sorex herpestes Duvernoy, Mem. Soc. du Mus. d'Hist. Nat. Strasbourg, 2: 

pt. 2-3, art. LL, p. 2, 1838. Swellendam and Winberg, Cape of Good Hope. 

Myosorex varius cafer (Sundevall) 

Sorex cafer Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. Forhandl., 
Stockholm, 3: no. 4, p. 119, 1846. "E Caffraria interiore et Port-Natal." 

PiLESOREX Thomas 

Crocidura Thomas, Ann. Mag. Nat. Hist. (7) 17: 177, Feb. 1906 (in part, 

not Wagler). 
Prsesarex Thomas, Ann. Mag. Nat. Hist. (8) 11: 320, Mar. 1913. Genotype 

Crocidura goliath Thomas. 

Pr^sorex GOLIATH (Thomas) 

Crocidura goliath Thomas, Ann. Mag. Nat. Hist. (7) 17: 177, Feb. 1906. 
Efulen, Cameroons. 

ScuTisoREX Thomas 

Sylvisorex Thomas, Ann. Mag. Nat. Hist. (8) 6: 113, July 1910 (in part, not 

Thomas, 1904). 
Scutisorex Thomas, Ann. Mag. Nat. Hist. (8) 11: 321, Mar. 1913. Genotype 

Sylvisorex somereni Thomas. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 49 

ScuTisoREX coNGicus Thomas 

Scutisorex congicus Thomas, Ann. Mag. Nat. Hist. (8) 16: 470, Dec. 1915; 

J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 769-784, pi. 89-92, 1917 

(skeleton). Medje, Upper Ituri River, Congo Beige. 

Scutisorex somereni (Thomas) 

Sylvisorex somereni Thomas, Ann. Mag. Nat. Hist. (8) 6: 113, July 1910. 
Kyetume, near Kampala, Uganda. 

SuNCUS Hemprich and Ehrenberg 

Suncus Hemprich and Ehrenberg, Symbolae Physicae, Mamm., dec. 2, k, 1832. 

Genotype Suncus sacer Hemprich and Ehrenberg, see Cabrera, Journ. 

Mamm., 5: 131, 1924. 
Sunkus Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 1842, 

p. 175, 1843 (as "Div. 1" of Sorex). 
Crocidura Wagler, Oken's Isis, p. 275, 1832 (in part); et auct. var. vet. 
Pachyura de Selys-Longchamps, Etudes de Micromamm., p. 32, 1839. As a 

subgenus of Sorex, type Sorex etruscus Savi. 
Paradoxodon Wagner, Schreber's Saugthiere, Suppl., 5: 805, 1855. As a sub- 
genus of Sorex; type Sorex melanodon Blyth, from Calcutta. 
Plerodus Schulze, Helios, Abh. u. Vortrage aus Gesammtgebiete Naturwiss., 

14: 90, 1897. Genotype Crocidura suaveolens Blasius = iSorex etruscus 

Savi. 

Suncus CjErul^eus (Kerr) 

(Sorex cseridseus Kerr, Animal Kingdom of Linn., p. 207, 1792. "Java and the 

other islands of the East Indies." 
Pachyura auriculata Fitzinger, Sitzb. K. Akad. Wiss. Wien, 31: 342, 1858; 

57: 145, 1868. Madagascar (introduced). 
Sorex mauritiana Reichenbach, Naturgesch. Raubthiere, p. 341, 1834 Mauritius 

(introduced). 

Suncus chriseos (Kershaw) 

Pachyura chriseos Kershaw, Ann. Durban Mus., 3: 31, 31 Jan. 1921. Durban, 

Natal. 

Suncus coquerelii (Trouessart) 

Crocidura (Pachyura) coquerelii Trouessart, Notes Leyden Mus., 2: 86, Apl. 
1880. Island of Mayotte, northwest coast of Madagascar. 

Suncus etruscus (Savi). Etruscan Shrew 

Sorex etruscus Savi, Nuovo Giorn. de' Letterati, 1: 60, pi. 1, 1822. Pisa, Italy. 
Range: Algeria (fide Coquerel, Ann. Sci. Nat., zool., (3) 9: 195, 1848), but 
exact records are lacking. 



50 bulletin: museum of comparative zoology 

SxJNCUS GRACILIS (BlainviUe) 

Sorex gracilis BlainviUe, Ann. des Sci. Nat., zool., (2) 10: 120, 1838. Cape of 
Good Hope, South Africa. 

SuNCUs GRATULUS (Thomas and Schwann) 

Pachyura gratula Thomas and Schwann, Proc. Zool. Soc. London, for 1906, 
p. 781, 11 Apl. 1907. Legogot, eastern Transvaal. 

SuNCUs iNFiNiTEsiMus (Heller) 

Pachyura infinitesima Heller, Smithsonian Misc. Coll., 60: no. 12, p. 5, 4 Nov. 
1912. Rumruti, Laikipia Plateau, Kenya Colony, 7000 feet. 

Stjnctjs LEtJCTJRA (Matschie) 

Pachyura leticura Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 205, 1894. 

Zanzibar and between the coast and Lake Victoria. 
Crocidura albicauda Noack, Jahrb. Hamburg. Wiss. Anst., 9: 117, 1891 (not 

Peters). Zanzibar. 

SuNCUS Lixus Lixus (Thomas) 

Crocidura (Pachyura) lixa Thomas, Proc. Zool. Soc. London, for 1897, p. 930, 
Apl. 1898. Nyika Plateau, northern Nyasaland. 

SuNcus LIXUS ^QUATORius (Heller) 

Pachyura lixa sequatoria Heller, Smithsonian Misc. Coll., 60: no. 12, p. 4, 4 
Nov. 1912. Summit of Mt. Sagalla, Taita Hills, Kenya Colony, 4000 feet. 

SuNcus MADAGASCARiENSis (Coquerel) 

Sorex madagascariensis Coquerel, Ann. des Sci. Nat., zool., (3) 9: 194, pi. 11, 
f. 1, la-Id, If, 1848. Madagascar. 

SuNcus ORANGi^ (Roberts) 

Pachyura orangise Roberts, Ann. Transvaal Mus., 10: 61, 31 Jan. 1924. Angra 
Pequina, Bothaville district, Orange Free State, South Africa. 

SuNCUS SACER Hemprich and Ehrenberg. 

Suncus sacer Hemprich and Ehrenberg, Symbote Physicse, Mamm., dec. 2, 

folio k, Sep. 1832. Suez, Egypt, "in domibus oppidi." 
Sorex crassicaudus Hemprich and Ehrenberg in Lichtenstein's Darstellung 

neuer oder wenig bekannt. Saugeth., pi. 40, f. 1 and text of pt. 7, 1834. 

Region about Suez, Egypt. 
Sorex indicus var. cinereo-senea Rtippell, Mus. Senckenberg., 3: 133, 1842. 

Coast of Shoa, Abyssinia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 51 

StJNCUS VARILLA VARILLA (Thomas) 

Crocidura (Pachyura) varilla Thomas, Ann. Mag. Nat. Hist. (6) 16: 54, July 
1895. East London, Cape of Good Hope. 

SuNCtrs VARILLA MINOR G. M. AUen and Loveridge 

Suncus varilla minor G. M. Allen and Loveridge, Bull. Mus. Comp. Zool., 75: 
57, Feb. 1933. Kitungulu, Urungu, Tanganyika Territory. 

Suncus warreni Roberts 

Suncus warreni Roberts, Ann. Transvaal Mus., 13: 84, 6 July 1929. Doom 
River, van Rhynsdorp district. Cape Colony, South Africa. 

SuRDisoREX Thomas 

Surdisorex Thomas, Ann. Mag. Nat. Hist. (7) 18: 223, Sep. 1906. Genotype 
Surdisorex norse Thomas. 

Surdisorex nor^ Thomas 

Surdisorex norse Thomas, Ann. Mag. Nat. Hist. (7) 18: 223, Sep. 1906. East 
side of Aberdare Range, near Nyeri, Kenya Colony. 

Surdisorex polulus Hollister 

Surdisorex polulus Hollister, Smithsonian Misc. Coll., 66: no. 1, p. 1, 10 Feb. 
1916. West side of Mount Kenya, Kenya Colony, 10,700 feet. 

Sylvisorex Thomas. Forest Shrews 

Sylvisorex Thomas, Abstract Proc. Zool. Soc. London, no. 10, p. 12, 22 Nov- 

1904; Proc. Zool. Soc. London, for 1904, 2: 190, 18 Apl. 1905. Genotype 

Crocidura morio Gray. 
Crocidura Gray, Proc. Zool. Soc. London, p. 180, Sep. 1862 (in part; not 

Wagler) . 
Myosorex Dobson, Proc. Zool. Soc. London, for 1887, p. 575, Apl. 1888 (in 

part; not Gray). 

Sylvisorex granti granti Thomas 

Sylvisorex granti Thomas, Ann. Mag. Nat. Hist. (7) 19: 118, Jan. 1907; Trans. 

Zool. Soc. London, 19: 492, Mar. 1910. Mubuku Valley, Ruwenzori 

East, Uganda, 10,000 feet. 

Sylvisorex granti mundus Osgood 

Sylvisorex mundus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 18, 7 

Apl. 1910. Kijabe, Kenya Colony. 
Sylvisorex granti mundus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 220, 

28 Dec. 1936. Doubtfully distinguishable. 



52 bulletin: museum of comparative zoology 

Sylvisorex johnstoni (Dobson) 

Myosorex johnstoni Dobson, Proc. Zool. Soc. London, for 1887, p. 577, text-fig. 
(skull), Apl. 1888. Rio del Rey, Cameroons. 
Range: Cameroons and Fernando Po. 

Sylvisorex lunaris Thomas 

Sylvisorex lunaris Thomas, Ann. Mag. Nat. Hist. (7) 18: 139, Aug. 1906; 

Trans. Zool. Soc. London, 19: 492, Mar. 1910. Mubuku Valley, Ruwen- 

zori East, Uganda, 12,500 feet. 

Sylvisorex morio (Gray) 

Crocidura morio Gray, Proc. Zool. Soc. London, p. 180, Sep. 1862. Cameroon 
Mountains, southern Nigeria. 

Sylvisorex ollula Thomas 

Sylvisorex ollula Thomas, Ann. Mag. Nat. Hist. (8) 11: 321, Mar. 1913. 
Bitye, Ja River, Cameroons, 2000 feet. 

Sylvisorex oriundtjs Hollister 

Sylvisorex oriundus Hollister, Bull. Amer. Mus. Nat. Hist., 35: 672, pi. 7, 

f. 7; pi. 9, f. 4, 4a; pi. 11, f. 1, 21 Oct. 1916. Medje, Nava River, Congo 

Beige. 

Sylvisorex preussi (Matschie) 

Myosorex preussi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 177, 1893. 
Buea, Cameroon Mountains area, southern Nigeria. 

Sylvisorex ruand^ Lonnberg and Gyldenstolpe 

Sylvisorex ruandse Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17B: no. 5, 
p. 1, 2 May 1925. Mount Sabinio, Birunga Volcanoes, Congo Beige. 

Sylvisorex megalura (Jentink) 

Pachyura megalura Jentink, Notes Leyden Mus., 10: 48, Apl. 1888. Schieffel- 

insville. Junk River, Liberia. 
Sylvisorex megalura Thomas, Proc. Zool. Soc. London, for 1904, 2: 190, 18 

Apl. 1905. 

Sylvisorex sorella sorella (Thomas) 

Myosorex (?) sorella Thomas, Proc. Zool. Soc. London, for 1897, p. 930, 1 Apl. 
1898. Masuku Plateau, Nyasaland. 

Sylvisorex sorella angolensis Roberts 

Sylvisorex angolensis Roberts, Ann. Transvaal Mus., 13: 84, 6 July 1929. 

Mombola, Angola. 
Sylvisorex sorella angolensis Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 

219, 28 Dec. 1936. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 53 

Sylvisorex sorella gemmeus Heller 

Sylvisorex gemmeus Heller, Smithsonian Misc. Coll., 56: no. 15, p. 7, 23 Dec. 

1910. Rhino Camp, Lado Enclave, Sudan. 
Sylvisorex sorella gemmeus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 219, 

28 Dec. 1936. 

Sylvisorex sorella infuscus Thomas 

Sylvisorex gemmeus infuscus Thomas, Ann. Mag. Nat. Hist. (8) 16: 152, Aug. 

1915. Bitye, Ja River, southern Cameroons. 
Sylvisorex sorella infuscus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 219, 

28 Dec. 1936. 

Sylvisorex sorella irene Thomas 

Sylvisorex gemmeus irene Thomas, Ann. Mag. Nat. Hist. (8) 16: 151, Aug. 1915. 

Kaganbah, Uganda, 1600 meters. 
Sylvisorex sorella irene Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 219, 28 

Dec. 1936. Doubtfully distinct from S. s. gemmeus Heller. 
Range: Southern Uganda and Upper Ituri, Uele. 

Sylvisorex sorella ph^opus Osgood 

Sylvisorex sorella pha?opus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 219, 
28 Dec. 1936. Near Dungulbar, southwest side of Lake Tana, Ethiopia. 

Sylvisorex sorella sheppardi Kershaw 

Sylvisorex gemmeus sheppardi Kershaw, Ann. Durban Mus., 3: 32, 31 Jan. 

1921. Vumba, Gorongoza Province, Portuguese East Africa. 
Sylvisorex sorella sheppardi Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 219, 

28 Dec. 1936. 

Sylvisorex sorella sorelloides Lonnberg 

Sylvisorex sorelloides Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 67, Jan. 1912; 

Kungl. Svenska Vet.-Akad. Handl., Stockholm, (2) 48: no. 5, 51, pi. 3, 

f. 1 (col.), 1912. Steppe near Itiolu River, northern Guaso Nyiro, Kenya 

Colony. 
Sylvisorex sorella sorelloides Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 219, 

28 Dec. 1936. Doubtfully distinct from S. s. gemmeus Heller. 

Sylvisorex suncoides Osgood 

Sylvisorex suncoides Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 217, 28 Dec. 
1936. Kalongi, western slope of Mt. Ruwenzori, Congo Beige. 



54 bulletin: museum of comparative zoology 

Order CHIROPTERA. Bats 

Suborder Megachiroptera 

Family PTEROPIDiE. Fruit Bats 

Subfamily Pteropin^ 

Revision: Andersen, Knud. Catalogue of the Chiroptera in the collection of 
the British Museum. Second edition. Vol. 1, Megachiroptera. 8vo, London 
(British Mus.), ci + 854 + 30 pp., 79 text-ff., 1912. 

Casintcteris Thomas 

Casinyderis Thomas, Ann. Mag. Nat. Hist. (8) 6: 111, July 1910. Genotype 
Casinyderis argynnis Thomas. 

Casintcteris argynnis Thomas 

Casinyderis qrgynnis Thomas, Ann. Mag. Nat. Hist. (8) 6: 111, July 1910. 
Bitye, Ja River, Cameroons. 

Eidolon Rafinesque. Straw-colored Fruit Bat 

Eidolon Rafinesque, Analyse de la Nature, p. 54, 1815. Genotype Eidolon 

helvum (Kerr). 
Pterocyon Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin p. 423, 1861. 

Genotype Pterocyon paleaceus Peters = Eidolon helvum (Kerr). 
Cynonyderis Dobson, Cat. Chiropt. Brit. Mus., p. 70, 1878 (in part) et auct. al. 
Leiponyx Jentink, Notes Leyden Mus., 3: 60, Apl. 1881. Genotype Leiponyx 

biittikoferi Jentink. 
Liponyx Forbes, Zool. Rec, 18: Mamm., p. 13, 1882 (emendatio), not Liponyx 

Vieillot, 1816, in Birds. 
Xantharpyia Matschie, Megachiroptera Berlin. Mus., p. 63, 1899 (in part). 

Eidolon dtjpreanum (Pollen) 

Pteropus dupreanus Pollen, in Schlegel and Pollen, Proc. Zool. Soc. London, 
for 1866, p. 419, Apl. 1867. [Northwestern Madagascar.] 
Range: Madagascar. 

Eidolon helvum helvum (Kerr) 

Vespertilio vampyrus helvus Kerr, Linnseus's Animal Kingdom, 1: pt. 1, p. 

xvii, 91, 1792. No locality. 
Vespertilio vampyrus var. C, Schreber, Saugthiere, 1: 154, 1774; var. y Gmelin, 

Linnseus's Syst. Nat., ed. 13, 1: 45, 1788. Africa occidentali, etc. 
Pteropus stramineus E. Geoffroy, Cat. Mamm. du Mus. National Hist. Nat., 

Paris, p. 48, 1803. No locality. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 55 

Vespertilio caninus, var. b, Goldfuss, Vergl. Naturbeschr., Saug., p. 98, 1809 

(not V. caninus Blumenbach). 
Pterocyon paleaceus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 423, 

1861. Africa. 
Pteropus mollipilosus H. Allen, Proc. Acad. Nat. Sci. Philadelphia, for 1861, 

p. 159, 1862. Gaboon. 
Pteropus palmarum Heuglin, Leopoldina (Halle), 5: 34, June 1865. Middle and 

Upper Nile. 
Xantharpyia leucomelas Fitzinger, Sitzb. K. Akad. Wiss. Wien, 54: sect. 1, 

pt. 10, p. 544, 1866. Sennaar, Bahr-el-Abiad, Bahr-el-Azrak, Kordofan. 
Leiponyx biittikoferi Jentink, Notes Leyden Mus., 3: 59, Apl. 1881. Millsburg, 

St. Paul's River, Liberia. 
Range: Somaliland, Sennaar and Senegambia to Nyasaland, Mashonaland 

and Namaqualand. 

Epomophorus Bennett 

Epomophorus Bennett, Proc. Zool. Soc. London, for 1835, p. 149, 12 Feb. 
1836; Trans. Zool. Soc. London, 2: 33, 2 Oct. 1836. Genotype Pteropus 
whitei vel epomophorus Bennett = gambianus Ogilby. 

Epomophorus angolensis Gray 

Epomophorus macrocephalus var. angolensis Gray, Cat. Monkeys, Lemurs and 
Fruit-eating Bats Brit. Mus., p. 125, 1870. Angola. 

Epomophorus anurus Heuglin 

Epomophorus anurus Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Dresden, 

31: no. 7, p. 12, 1864. Bongo, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 
Epomophorus doriae Matschie, Megachiroptera Berlin. Mus., p. 54, 1899. 

Bogos Country, Eritrea. 

Epomophorus crypturus Peters 

Epomophorus crypturus Peters, Reise nach Mossambique, Saugethiere, p. 26, 

pi. 5; pi. 13, f. 1-6, 1852. Tette, Portuguese East Africa, 17' S. 
Range: Transvaal, Zambesi and Limpopo valleys north to Nyasa and 

Katanga. 

Epomophorus gambianus (Ogilby) 

Pteropus gambianus Ogilby, Proc. Zool. Soc. London, p. 100, 9 Oct. 1835. 

Gambia. 
Pteropus macrocephalus Ogilby, Proc. Zool. Soc. London, p. 101, 9 Oct. 1835. 

Gambia. 
Pteropus epomophorus Bennett, Proc. Zool. Soc. London, p. 149, 12 Feb. 1836. 

Gambia. 



56 bulletin: museum of comparative zoology 

Pteropus whitei Bennett, Trans. Zool. Soc. London, 2: pt. 1, p. 37, pi. 6; pi. 7 

(structure of hair), 2 Oct. 1836. Gambia. 
Epomophorus guineensis Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 5: 

136, f. 3, June 1898. Bolama, Portuguese Guinea. 
Epomophorus zechi Matschie, Megachiroptera Berlin. Mus., p. 46, pi. 10, f. 

3 a-c, 1899. Lectotype from Kunjuruma, Togoland, = Dahomey. 
Range: Senegal to Nigeria and east to Sennaar and southern Abyssinia. 

Epomophorus labiatus labiatus (Temminck) 

Pteropus labiatus Temminck, Monogr. de Mammalogie, 2: 83, pi. 39, 1837. 
"Abyssinia," i.e. Sennaar. 

Pteropus schoensis Riippell, Mus. Senckenberg., Frankfurt a. M., 3: 131, 
1842. Shoa. 

Epomophorus schovanus Heuglin, Reise in Nordost-Afrika, 2: 18, 1877 (emenda- 
tion). 
Range: Sennaar, Abyssinia, Shoa. 

Epomophorus labiatus minor Dobson 

Epomophorus minor Dobson, Proc. Zool. Soc. London, for 1879, p. 715, Apl. 
1880. Zanzibar. 
Range: Shoa south to Tanganyika Territory. 

Epomophorus pousarguesi Trouessart 

Epomophorus pousarguesi Trouessart, Cat. Mamm. Viv. Foss., Suppl., p. 55, 

1904. Based on E. macrocephalus Pousargues (not Ogilby), in Ann. Sci. 

Nat., zool. (8) 3: 253, 1896. Shari region, eastern Congo Beige. 

Epomophorus wahlbergi wahlbergi (Sundevall) 

Pteropus wahlbergi Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. For- 

handl., Stockholm, 3: no. 4, p. 118, 1846. Near Port Natal and in interior 

of Caff r aria. 
Epomophorus macrocephalus var. unicolor Gray, Cat. Monkeys, Lemurs and 

Fruit-eating Bats Brit. Mus., p. 125, 1870. Shupanga, Zambesia, South 

Africa. 
Epomophorus stuhlmanni Matschie, Megachiroptera Berlin. Mus., p. 50, 1899 

(in part). Lectotype from Vikindo, Usaramo, Tanganyika Territory. 
Range: Kenya Colony, Mt. Kenya, south through Tanganyika Territory to 

eastern Cape Colony. 

Epomophorus wahlbergi haldemani (Halowell) 

Pteropus haldemani Halowell, Proc. Acad. Nat. Sci. Philadelphia, 3: 52, 1846. 

West Africa [? Liberia]. 
Epomophorus zenkeri Matschie, Megachiroptera Berlin. Mus., p. 46, 1899. 

Lower Guinea, lectotype from Chinchoxo, Cabinda, Angola. 



ALLEN: CHE.CKLIST OF AFRICAN MAMMALS 57 

Epomophorus neumanni Matschie, Megachiroptera Berlin. Mus., p. 50, 1899. 
Lectotype from Mombasa, Kenya Colony. 
Range: Cameroons to Benguela, east to Kenya Colony and Tanganyika 
Territory. 

Epomops Gray. Epaulet Bats 

Epomophorus Tomes, Proc. Zool. Soc. London, p. 44, Feb. -May 1860 (in part). 
Epomops Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., p. 
126, 1870. Genotype Epotnophorus franqueti Tomes. 

Epomops stJTTiKOFERi (Matschie) 

Epomophorus biittikoferi Matschie, Megachiroptera Berlin. Mus., p. 45, 1899. 
Schieffelinsville, Junk River, Liberia. 
Range: Liberia and Sierra Leone. 

Epomops dobsonii (Bocage) 

Epomophorus dobsonii Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 1: 
1, 14, fig., Mar. 1889. Quindumbo, Benguela, Angola. 
Range: Angola to Katanga. 

Epomops franqueti franqijeti (Tomes) 

Epomophorus franqueti Tomes, Proc. Zool. Soc. London, p. 54, pi. 75 (col.), 

Feb.-May 1860. Gaboon. 
Epomophorus comptus H. Allen, Proc. Acad. Nat. Sci. Philadelphia, for 1861, 

p. 158, 1862. "W. Africa," = Gaboon. 
Range: Old Calabar to Loanda, east to the great lakes. 

Epomops franqueti strepitans K. Andersen 

Epomops franqueti strepitans K. Andersen, Ann. Mag. Nat. Hist. (8) 5: 106, 3 
Jan. 1910. Asaba, southern Nigeria. 
Range: Gold Coast to southern Nigeria. 

Hypsignathus H. Allen. Hammer-headed Bats 

Hypsignathus H. Allen, Proc. Acad. Nat. Sci. Philadelphia, p. 156, July 1861. 
Genotype Hypsignathus monstrosus H. Allen. 

Sphyrocephalus A. Murray, Proc. Zooh Soc. London, p. 8, June 1862. Genotype 
Sphyrocephalus labrosus Murray = Hypsignathus monstrosus H. Allen. 

Zygsenocephalus A. Murray, Proc. Zool. Soc. London, pi. 1, June 1862. Geno- 
type Zygsenocephalus labrosus Murray = Hypsignathus monstrosus H. 
AUen. 

Hypsignathus monstrosus H. Allen 

Hypsignathus monstrosus H. Allen, Proc. Acad. Nat. Sci. Philadelphia, p. 157, 
July 1861. "West Africa," = Gaboon. 



58 bulletin: museum of comparative zoology 

Sphyrocephalus labrosus A. Murray, Proc. Zool. Soc. London, p. 8, June, 1862. 

Old Calabar River. 
Zygspnocephalus labrosus A. Murray, Proc. Zool. Soc. London, pi. 1 (not text), 

June, 1862. Old Calabar River. 
Range: Gambia, and Fernando Po, to Ogowe, and east to Monbuttu, Congo 

Beige. 

MiCROPTEROPUS Matschie. Dwarf Epaulet Bat 

Micropteropus Matschie, Megachiroptera Berlin. Mus., pp. 36, 57, 1899. 

As a subgenus of Epomophorus; type Epomophorus pusillus Peters. 
Epomophorus Tomes, Proc. Zool. Soc. London, p. 42, Feb.-May 1860 (in part), 

et auct. al. 

Micropteropus pusillus (Peters) 

Epomophorus piisillus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 

870, 1867. Yoruba, southern Nigeria. 
Range: Gambia to Loanda, east to Bahr-el-Ghazal and east side of Victoria 

Nyanza. 

Myonycteris Matschie 

Myonyderis Matschie, Megachiroptera Berlin. Mus., pp. 61, 63, 1899. As a 

subgenus of Xantharpyia; type Cynonycteris torquata Dobson. 
Cynonycteris Dobson, Cat. Chiroptera Brit. Mus., p. 71, 1878 (in part). 
Rousettus Trouessart, Cat. Mamm. Viv. Foss., Suppl., p. 59, 1904 (in part). 
Xantharpyia Matschie, Megachiroptera Berlin. Mus., p. 65, 1899 (in part). 



Subgenus Myonycteris Matschie 

Myonycteris leptodon K. Andersen 

Myonycteris leptodon K. Andersen, Ann. Mag. Nat. Hist. (8) 2: 450, Nov. 1908. 
Sierra Leone. 
Range: Sierra Leone and Liberia. 

Myonycteris torquata (Dobson) 

Cynonycteris torquata Dobson, Cat. Chiroptera Brit. Mus., pp. 71, 76, pi. 5, f. 1, 

June 1878. Angola. 
Cynopterus collaris Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. 

Mus., p. 123, 1870 (not Kolenati; not Geoffroy). "W. Africa" = Lower 

Congo; Angola. 

Myonycteris wroughtoni K. Andersen 

Myonycteris wroughtoni K. Andersen, Ann. Mag. Nat. Hist. (8) 2: 450, Nov. 
1908. River Likand, Uele district, northeastern Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 59 

Subgenus Phygetis K. Andersen 

Phygetis K. Andersen, Cat. Chiroptera Brit. Mas., 1: 579, 1912. As a subgenus; 
type Cynonyderis brachycephala Bocage. 

Myonycteris brachycephala (Bocage) 

Cynonyderis brachycephala Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 
1 : 198, Dec. 1889. Island of San Thome, Gulf of Guinea. 

Nanonycteris Matschie 

Nanonyderis Matschie, Megachiroptera Berlin. Mus., pp. 36, 58, 1899. As a 

subgenus of Epomophorus; type Epomophorus veldkampii Jentink. 
Epomophorus Jentink, Notes Leyden Mus., 10: 51, Apl. 1888 (in part). 

Nanonycteris veldkampii (Jentink) 

Epomophorus veldkampii Jentink, Notes Leyden Mus., 10: 51, Apl. 1888. 
Buluma, Fisherman Lake, Liberia. 
Range: Liberia and Gold Coast to southern Nigeria. 

Plerotes K, Andersen 

Plerotes K. Andersen, Ann. Mag. Nat. Hist. (8) 5: 97, 3 Jan. 1910. Genotype 
Epomophorus anchietse Seabra. 

Plerotes anchiet^ (Seabra) 

Epomophorus anchietse Seabra, Jorn. Sci. Math., Phya. e Nat., Lisboa, (2) 6: 

116, Aug. 1900; (2) 5: 166, pi. 1, fig. 3 (palate ridges) no specific name 

given, June 1898. Galanga, Benguela, Angola. 



Pteropus Brisson. Flying Foxes 

Pteropus Brisson, Regne Animal, ed. 2, pp. 13, 153, 1762. Genotype Pteropus 
niger (Kerr). 

Spectrum Lacepede, Tabl. Mamm., p. 15, 1799 (not Spectrum Scopoli, 1777, 
Lepidoptera). Genotype Pteropu,s niger (Kerr). 

Eunycteris Gray, Proc. Zool. Soc. London, p. 64, May 1866. Genotype Pteropus 
phaiops Temminck = Pteropus melanopogon Peters. 

Pselaphon Gray, Cat. Monkeys, Lemurs, and Fruit-eating Bats Brit. Mus., 
p. 110, 1870. Genotype Pteropus pselaphon Layard. 

Sericonyderis Matschie, Megachiroptera Berlin. Mus., pp. 6, 30, 1899. As a 
subgenus of Pteropus; type Pteropus subniger (Kerr). 

Desmalopex Miller, Bull. U. S. Nat. Mus., no. 57, p. 60, 29 June 1907. Geno- 
type Pteropus leucopterus Temminck. 



60 bulletin: museum of comparative zoology 

Pteropus aldabrensis True 

Pteropiis aldabrensis True, Proc. U. S. Nat. Mus., 16: 533, 14 July 1893. Alda- 
bra Island. 
Range: Aldabra Island, north of Comoro group. 

Pteropus comorensis Nicoll 

Pteropus comorensis Wallace, Island Life, p. 400, 1880 (nomen nudum). Nicoll, 
Three Voyages of a Naturalist, pp. 87, 88, 90, 1908. Comoro Islands. 

Pteropus livingstonii Gray 

Pteropus livingstonii Gray, Proc. Zool. Soc. London, p. 66, May 1866. Johanna 
Island, Comoro group. 

Pteropus nicer (Kerr) 

Vespertilio vampirus niger Kerr, Linnajus's Animal Kingdom, 1: xvii, 90, 1792. 

No locality. 
Vespertilio caninus Blumenbach, Handbuch Naturg., ed. 5, p. 73, 1797 (in 

part; not of Wied). 
Spectrum vampirus Lacepede, Tabl. Mammiferes, p. 15, 1799; Daudin, in 

Buffon's Hist. Nat., Quadrupedes, Didot ed., 14: 188, 1802. "L'ile de 

Bourbon" = Mauritius. 
Pteropus fuscus E. Geoffroy, Cat. Mamm. Mus. National d'Hist. Nat., Paris, 

p. 46, 1803. Reunion and Madagascar. 
Vespertilio mauritianus Hermann, Observationes Zool., p. 19, 1804. Mauritius. 
Pteropus vulgaris E. Geoffroy, Ann. Mus. d'Hist. Nat., Paris, 15: 92, 1810. 

Reunion and Madagascar. 
Pteropus pteropus Merriam, ex Brisson, Science, n. ser., 1 : 376, 5 Apl. 1895. 
Range: The Mascarene Islands, Reunion and Mauritius. 

Pteropus rodricensis Dobson 

Pteropus rodricensis Dobson, Cat. Chiroptera Brit. Mus., p. 36, 1878. Island 

of Rodriguez. 
Pteropus mascarinus Mason, Ann. Mag. Nat. Hist. (7) 20: 221, Sept. 1907. 

Round Island (La Ronde), 15 miles northeast of Mauritius. 

Pteropus rufus rufus E. Geoffroy 

Pteropus rufus E. Geoffroy, Cat. Mamm. Mus. National d'Hist. Nat., Paris, 

p. 47, 1803. Madagascar. 
Pteropus edwardsii E. Geoffroy, Ann. Mus. d'Hist. Nat., Paris, 15: 92, 1810. 

Madagascar (renaming of P. rufus). 
Pteropus madagascariensis Oken, Lehrbuch d. Naturgesch.,pt.3: sect. 2, p. 936, 

1816. Madagascar (renaming of P. edwardsii). 
Pteropus phaiops Temminck, Monogr. Mammal., 1: 178, 1825. Madagascar. 
Range: Northern and central Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 61 

Pteropus rtjfus princeps K. Andersen 

Pteropus rufus princeps K. Andersen, Ann. Mag. Nat. Hist. (8) 2: 367, Oct. 
1908. Ft. Dauphin, Madagascar. 
Range: Southern Madagascar. 

Pteropus seychellensis Milne-Edwards 

Pteropus seychellensis Milne-Edwards, Bull. Soc. Philom. Paris, (7) 2: 221, 

1877. Seychelles Ids. 
Range: Seychelles Islands. 

Pteropus subniger (Kerr) 

Vespertilio vampyrus subniger Kerr, Linnseus's Animal Kingdom, 1 : xvii, 91, 1792. 

No locality. 
Pteropus collaris Illiger, Abhandl. K. Preuss. Akad. Wiss., Berlin, 1804r-ll, 

p. 78, 84, 1815. 
Pteropus fuscus Desmarest, Nouv. Diet. d^Hist. Nat., 19: 544, 1803 (not E. 

Geoff roy). Reunion. 
Pteropus torquatus G. Fischer, Zoognosia, 3: 553, 1814. 
Spectrum rubidum Daudin, in Buffon, Hist. Nat., Quadrupedes, Didot ed., 14: 

188, 1802. (?) 
Pteropus rubricollis E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 15: 93, 1810. 

Reunion Id. 
Range: The Mascarene Islands, Reunion and Mauritius. 

Pteropus voeltzkowi Matschie 

Pteropus {Spectrum) voeltzkowi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 486, Oct. 1909. Fufuni, Pemba Island, off coast of Tanganyika Terri- 
tory. 
Range: Pemba Island. 

RousETTUS Gray. Rousette Bats 
Subgenus Rousettus Gray 

Rousettus Gray, London Medical Repository, 15: 299, 1 Apl. 1821. Genotype 

Pteropus segypliacus E. Geoffroy. 
Cercopteropus Burnett, Quart. Journ. Sci., Lit. and Art, pt. 1, p. 269, Apl-June, 

1829. Genotype Pteropus segyptiacus E. Geoffroy. 
Xantharpyia Gray, List Spec. Mamm. Brit. Mus., pp. xix, 37, 13 May 1843; 

Genotype Pteropus amplexicaudatus E. Geoffroy. 
Eleutherura Gray, List. Spec. Mamm. Brit. Mus., p. xix, 13 May 1843 (nomen 

nudum); Voyage "Sulphur," 1: 29, 1844. Genotype Pteropus leachii A. 

Smith. 
Senonycteris Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., 

p. 115, 1870. As a subgenus of Xantharpyia; type Pteropus seminudus 

Kelaart. 



62 bulletin: museum of comparative zoology 

Cynonyderis Peters, Reise nach Mossambique, Saugeth., p. 25, 1852. Geno- 
type Pteropus collaris Illiger = Pteropus leachii A. Smith. 

RousETTTJS ^GYPTiAcus (E. Geoffroy) 

Pteropus egyptiacus E. Geoffroy, Ann. Mus. d' Hist. Nat., Paris, 15: 96, 1810 

(misprint); corrected to segyptiacus in Descrip. de I'Egypte, Hist. Nat., 

2: 134, pi. 3, f. 2, 1818. From the Great Pyramid, Gizeh, Egypt. 
Pteropus geoffroyi Temminck, Monogr. Mamm., 1: 197, pi. 15, f. 14, 15, 1825. 

Senegal and probably north coast of Africa. 
Eleuiherura unicolor Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. 

Mus., p. 117, 1870. Gaboon. 
Range: Loando and Gaboon to Palestine and Cyprus. 

RousETTUs LEACHII (A. Smith) 

Pteropus collaris Lichtenstein, Verzeichn. Doubl. Mus. Berlin, pp. 3, 5, 

1823 (in part), not Illiger. Terra Caffrorum. 
Pteropus leachii A. Smith, Zool. Journ., 4: 433, May 1829. "Gardens about 

Cape Town," South Africa. 
Pteropus holtentottus Temminck, in Smuts, Enum. Mamm. Cap., p. 3, 

1832. Capetown, Cape Province. 
Rousettus sjostedti Lonnberg, Wiss. Ergebn. Swedischen Zool. Exp. u. Leitung 

Sjostedt, Kilimandjaro-Meru, 1: no. 2, p. 7, 1910. Mkulumusi caves, near 

Tanga, Tanganyika Territory. 
Range: Cape Colony northeastward to Kenya Colony. 



Subgenus Stenonycteris Andersen 

Stenonycteris Andersen, Cat. Chiroptera Brit. Mus., 1: 23, 1912. Type Rou- 
settus lanosus Thomas. 

Rousettus kempi Thomas 

Rousettus kempi Thomas, Ann. Mag. Nat. Hist. (8) 4: 543, Dec. 1909. Kirui, 
Mt. Elgon, Kenya Colony, 6000 feet. 
Range: Mt. Elgon to Shoa. 

Rousettus lanosus Thomas 

Rousettus lanosus Thomas, Ann. Mag. Nat. Hist. (7) 18: 137, Aug. 1906; 

Trans. Zool. Soc. London, 19: 486, pi. 20 (col.), Mar. 1910. Mubuku 

Valley, Ruwenzori East, Uganda, 13,000 feet. 
Range: Ruwenzori. 

Rousettus madagascariensis G. Grandidier 

Rousettus madagascariensis G. Grandidier, Bull. Acad. Malgache, 11: 91, pi., 4 

text-figs, (skull), 1928. Beforona, between Tananarive and Andevoranto, 

eastern Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 63 



Subgenus Lissonycteris Andersen 

Lissonyderis Andersen, Cat. Chiroptera Brit. Mus., 1: 23, 1912. Subgenus of 
Rousettus; type Rousettus angolensis (Bocage). 

RousETTUs ANGOLENSIS (Bocage) 

Cynonycteris angolensis Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 5: 
133, 138, text-f., June 1898. Pungo Andongo, Cahata, Quibula, Angola. 
Range: Angola and Canaeroons east to Kenya Colony and Tanganyika 
Territory. > 

Rousettus crypticola Cabrera 

Rousettus crypticola Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 20: 
106, Mar. 1920. Cave of San Fernando, Basile, Fernando Po. 

Rousettus smithii Thomas 

Rousettus smithii Thomas, Ann. Mag. Nat. Hist. (8) 2: 375, Oct. 1908. Sierra 
Leone. 
Range: Sierra Leone to southern Nigeria. 

ScoTONYCTERis Matschic 

Scotonycteris Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 200, 1894. 
Genotype Scotonycteris zenkeri Matschie. 

Scotonycteris zenkeri Matschie 

Scotonycteris zenkeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 202, 

1894. Yaunde, southern Cameroons. 
Scotonycteris bedfordi Thomas, Abstract Proc. Zool. Soc. London, no. 4, 

p. 14, 8 Mar. 1904; Proc. Zool. Soc. London, 1: 372, 2 Aug. 1904. Fish 

Town, Fernando Po. 



Subfamily MACROGLOSSIN^. Long-tongued Fruit Bats 
Megaloglossus Pagenstecher 

Megaloglossus Pagenstecher, Zool. Anzeiger, 8: 245, 27 Apl. 1885. Genotype 

Megaloglossus woermanni Pagenstecher. 
Trygenycteris Lydekker, in Flower and Lydekker, Mammals Living and 

Extinct, p. 655, 1891. Genotype Megaloglossus woermanni: proposed to 

replace Megaloglossus, thought to be preoccupied by Megaglossa Rondani, 

1865 (Diptera). 

Megaloglossus woermanni Pagenstecher 

Megaloglossus woermanm Pagenstecher, Zool. Anzeiger, 8: 245, 27 Apl. 1885; 
Jahrb. Wiss. Anst. Hamburg, 2: 123, pi., 1885. Sibange Farm, Gaboon. 
Range: Liberia to Cameroons and lower Congo. 



64 bulletin: museum of comparative zoology 

Suborder MICROCHIROPTERA 

Family RHINOPOMIDiE. Mouse-tailed Bats 

Rhinopoma Oken 

Rhinopoma Oken, Lehrbuch d. Naturgesch., pt. 3, zool., sect. 2, p. 926, 1816; 

E. Geoffrey, Descrip. de I'Egypte, 2: 113, 1818 (see Lyon, M. W., Jr., 

Proc. Biol. Soc. Washington, 27: 217, 1914). Genotype Vespertilio 

microphyllus Briinnich. 
Rhinopomus Gervais, Hist. Nat. des Mammiferes, p. 202, 1854 (lapsus). 

Genotype Vespertilio microphyllus Briinnich. 
Rhynopoma Bowditch, Anal. Nat. Class. Mamm., p. 30, 1821. Genotype 

Vespertilio microphyllus Briinnich. 

Review: Thomas, O. On the species of the genus Rhinopoma. Ann. Mag. 
Nat. Hist. (7) 11: 496-499, May 1903. 

Rhinopoma cystops cystops Thomas 

Rhinopoma cystops Thomas, Ann. Mag. Nat. Hist. (7) 11: 496, May 1903. 
Luxor, Egypt. 
Range: Egypt and Sudan west to Air, Sahara. 

Rhinopoma cystops macinnesi Hayman 

Rhinopoma cystops macinnesi Hayman, Ann. Mag. Nat. Hist. (10) 19: 530, 
May 1937. Bat Island, near Central Island, Lake Rudolf. 

Rhinopoma microphyllum (Briinnich) 

Vespertilio microphyllus Briinnich, Dyrenes Historie og Dyre-Samlingen udi 

Univ. Natur-Theater, 1 : 50, pi. 6, f. 1-4, 1782. Arabia and Egypt. 
Rhinopoma sennaariense Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien, 

54: sect. 1, 547, 1866 (nomen nudum). Sennaar and Fazogli, near Ro- 

seires, Anglo-Egyptian Sudan. 
Rhinopoma longicaudatum Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien. 

54: sect. 1, 547, 1866 (nomen nudum). Sennaar. 
Rhinopoma lepsianum Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 222, 

1859. Blue Nile. 
For Herrn Lepsius. 
Rhinopoma cordofanicum Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien, 

54: sect. 1, 547, 1866 (nomen nudum); Heuglin, Reise Nordost-Afrika, 

2: 24, 1877 (nomen validum). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 65 

Family EMBALLONURID.E. Sheath-tailed Bats 
Emballonura Temminck 

Emballonura Temminck, van der Hoeven's Tijdschr. Nat. Gesch. en Physiol., 
5: 22, 1838. Genotype, by elimination, Emballonura monticola Temminck. 

Emballonura atrata Peters 

Emballonura atrata Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 693, 
1874. Interior of Madagascar. 

Range: Madagascar and adjacent island of Nossi-Be. 

CoLEURA Peters 

Coleura Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 479, 1867. Geno- 
type Emballonura afra Peters. 
Review: Thomas, O. Notes on bats of the genus Coleura. Ann. Mag. Nat. 
Hist. (8) 15: 576-579, 1915. 

Coleura afra (Peters) 

Emballonura afra Peters, Reise nach Mossambique, Saugethiere, p. 51, pi. 12; 
pi. 13, f. 18, 19, 1852. Tette, Portuguese East Africa. 

Coleura gallarum gallarum Thomas 

Coleura gallarum Thomas, Ann. Mag. Nat. Hist. (8) 15: 576, June 1915. 
Zeyla, British Somaliland. 
Range: Somaliland; Aden, Arabia. 

Coleura gallarum nilosa Thomas 

Coleura gallarum nilosa Thomas, Ann. Mag. Nat. Hist. (8) 15: 576, June 1915. 
Upper Nile, near mouth of Bahr-el-Zeraf, Sudan. 
Range: Upper Nile to northeastern Congo Beige. 

Coleura seychellensis Peters 

Coleura seychellensis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 
1868, p. 367, 1869. Seychelles Ids. 
Range: Seychelles group (? and Zanzibar). 

Coleura silhouette Thomas 

Coleura silhouettse Thomas, Ann. Mag. Nat. Hist. (8) 15: 576, June 19154 
Silhouette Island, Seychelles group. 



66 bulletin: museum of comparative zoology 

Taphozous E. Geoffroy. Tomb Bats 

Taphozous E. Geoffroy, Description de I'Egypte, 2: 113, 1818. Genotype 
Taphozous perjoratxis Geoffroy. Oken, Lehrbuch d. Naturg., pt. 3, zool., 
sect. 2, p. 926, 1816, is by some considered an authority. 

Taphozous hamiltoni Thomas 

Taphozous hamiltoni Thomas, Ann. Mag. Nat. Hist. (9) 5: 142, Jan. 1920. 
Mongalla, Sudan. 

Taphozous hildegarde.e Thomas 

Taphozous hildegardese Thomas, Ann. Mag. Nat. Hist. (8) 4: 98, Aug. 1909. 
Rabai, Mombasa district, Kenya Colony. 

Taphozous mauritianus mauritianus E. Geoffroy 

Taphozous mauritianus E. Geoffroy, Description de I'Egypte, 2: 127, 1818. 

Island of Mauritius. 
Taphozous leucopterus Temminck, Monogr. de. Mammalogie, 2: 284, pi. 60, 

f. 7, 1835-41. Interior of South Africa. 
Taphozous dobsoni Jentink, Notes Leyden Mus., 1: 123, Apl. 1879. Mahambo, 

Madagascar. 

Taphozous mauritianus cinerascens Seabra 

Taphozous maritianus (sic) cinerascens Seabra, Jorn. de Sci. Math., Phys. e 
Nat., Lisboa, (2) 6: 77, Aug. 1900. Benguella, Angola (? valid race). 

Taphozous perforatus perforatus E. Geoffroy 

Taphozous perforatus E. Geoffroy, Description de I'Egypte, 2: 126, 1818. 
Egypt. 

Taphozous perforatus h^dinus Thomas 

Taphozous perforatus h^dinus Thomas, Journ. Bombay Nat. Hist. Soc, 24: 
62, 30 Sept. 1915. Chanler Falls, northern Guaso Nyiro, Kenya Colony. 
Range: Kenya Colony to Aden. 

Taphozous senegalensis Desmarest 

Taphozous senegalensis Desmarest, Encyclop. Methodique, Mammalogie, 1 : 

130, 1820. Senegal (based on "le lerot-volant" of Daubenton, M6m. 

Acad. Paris, p. 386, 1759). 

Taphozous sudani Thomas 

Taphozous sudani Thomas, Ann. Mag. Nat. Hist. (8) 15: 561, June 1915. 
Mongalla, north of Lado, Upper Nile, Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 67 



LiPONYCTERis Thomas 

Liponycieris Thomas, Ann. Mag. Nat. Hist. (9) 9: 266, Mar. 1922. Genotype 
Taphozous nudiventris Cretzschmar. 

LiPONYCTERis NUDIVENTRIS (Cretzschmar) 

Taphozous nudiventris Cretzschmar in Rlippell's Atlas, Reise Nord-ost Afrika, 

Saugethiere, p. 70, fig. 27b, 1826. 
Taphozous nudiventer Temminck, Monogr. de Mammalogie, 2: 280, 1835-41 

(lapsus cal.). 
Nydicejus serratus Heuglin, Reise in Nordost- Afrika, 2: 35, 1877. "Scherq el 

Aqabah," on the Araschkol and nearby mountains, ?Abyssinia. 
Taphozous perforatus var. assabensis Monticelli, Ann. Accad. Aspir. Nat., 

Napoli, 1 : 78, pi. 7, 1885. Assab, Eritrea. A synonym of perforatus fide 

MonticeUi, but Senna, A., 1905, who examined the type says it is T. 

nudiventris. 

Saccolaimus Lesson 

Saccolaimus Lesson, Nouv. Tabl. Regne Anim., Mamm., p. 19, 1842 (pub- 
lished as a syn. of Taphozous), a nomen nudum; Gray, Ann. Mag. Nat. 
Hist., (3) 17: 92, Feb. 1866 (no type designated); Fitzinger, Sitzb. K. 
Akad. Wiss. Wien, math.-nat. CI., 61: sect. 1, 483, Apl. 1870. Genotype 
Taphozous saccolaimus Temminck. 

Taphonycteris Dobson, Proc. Zool. Soc. London, p. 548, 1875. As a subgenus 
of Taphozous for Taphozous saccolaimus, T. affinis and T. peli. 

Saccolaimus peli (Temminck) 

Taphozous peli Temminck, Esquisses zool. sur la cote de Guine, p. 82, 1853. 
Boutry River, Gold Coast. 



Family NYCTERIDyE. HoUow-faced Bats 
Nycteris G. Cuvier and E. Geoff roy 

Nycteris Cuvier and Geoffroy, Methode Mammalogique, in Mag. Encyclo- 
pcdique, 1 annee, 2: 186, 1795; Geoffroy, Cat. Mamm. Mus. National 
d'Hist. Nat., Paris, p. 64, 1803. Genotype Vespertilio hispidus Schreber. 
Not Nycteris Borkhausen, Der Zoologe (Compendiose Bibliothek gemein- 
niitzigsten Kentnisse fiir alle Stande, pt. 21), no. 4-7, p. 66, 1797; geno- 
type Vespertilio borealis S. L. Miiller = Lasiurus borealis (Miiller). 
Adopted by Opinion 111 of International Comm. of Zool. Nomenclature, 

see Smithsonian Misc. Coll., 73: no. 6, p. 18, 8 June 1929. 

Nicteris Desmarest, Nouv. Dictionnaire d'Hist. Nat., 15: 501, 1803. 



68 bulletin: museum of comparative zoology 

Petalia Gray, Mag. Zool. and Bot., 2: 494, 1838. Genotype Nyderis javanica 

Geoffrey. 
Nycterops Gray, Proc. Zool. Soc. London, p. 83, May 1866. Genotype Nycierops 

pilosa Gray = Nyderis hispida (Schreber). 

Review (as Petalia) : Andersen, K. Brief diagnoses of eight new Petalia, with 
a list of the known forms of the genus. Ann. Mag. Nat. Hist. (8) 10: 546-550, 
Nov. 1912. 

Nyctehis ^thiopica ^thiopica Dobson 

Plecotus sethiopicus Heughn and Fitzinger, Sitzb. K. Akad. Wiss. Wien, 54: 

pt. 1, p. 546, 1866; Heughn, Reise in Nordost-Afrika, 2: 31, 1877. Bahr- 

el-Abiad, Anglo-Egyptian Sudan; a nomen nudum. 
Nyderis sethiopica Dobson, Cat. Chiroptera Brit. Mus., p. 165, 1878. Kordofan. 

Nycteris arge Thomas 

Nyderis arge Thomas, Ann. Mag. Nat. Hist. (7) 12: 633, Dec. 1903. Efulen, 
Cameroons. 
Range: Northeastern Congo Beige to Cameroons and Liberia. 

Nycteris aurita (K. Andersen) 

Petalia aurita K. Andersen, Ann. Mag. Nat. Hist. (8) 10: 547, Nov. 1912. Kilifi, 
Kenya Colony. 
Range: Somaliland and Kenya Colony. 

Nycteris avaktjbia J. A. Allen 

Nycteris avakubia J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 426, fig. 3, 29 
Sep. 1917. Avakubi, Congo Beige. 

Nycteris capensis A. Smith 

Nyderis capensis A. Smith, Zool. Journ.,4: 434, 1829. Interior of South Africa. 

Nycteris affinis A. Smith, Zool. Journ., 4: 434, 1829. South Africa. 

Nycteris discolor Wagner, Schreber's Saugthiere, Suppl., 1: 440, 1840; 5: 646, 

1855. "Siidspitze von Afrika." 
Nycteris fuliginosa Peters, Reise nach Mossambique, Saugethiere, p. 46, pi. 10, 

1852. Boror, 12 miles northwest of Quellimane, Mozambique. 

Nycteris damarensis damarensis Peters 

Nycteris damarensis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 1870, 
p. 905, 1871. Otjimbingue, DamaralaDd, Southwest Africa. "^ 

Nycteris damarensis brockmani (K. Andersen) 

Petalia damarensis brockmani Andersen, Ann. Mag. Nat. Hist. (8) 10: 548, Nov. 
1912. Upper Sheikh, British Somaliland. 
Range : Somaliland to Eritrea. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 69 

Nycteris damarensis media (K. Andersen) 

Petalia damarensis media K. Andersen, Ann. Mag. Nat. Hist. (8) 10: 548, 
Nov. 1912. Harar, Abyssinia. 
Range: Abyssinia. 

Nycteris gambiensis (K. Andersen) 

Petalia gambiensis K. Andersen, Ann. Mag. Nat. Hist. (8) 10: 548, Nov. 1912. 
Dialocote, French Gambia. 

Nycteris grandis Peters 

Nycteris grandis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 358, 1865; 

for 1870, p. 906, 1871. "Guinea." 
Nycteris baikii Gray, MS., in Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 

for 1866, p. 672, 1867. West Africa. 

Nycteris hispida (Schreber) 

Vespertilio hispidus Schreber, Saugthiere, 1: 169, 188, pi. 56, 1775. Senegal. 

Nycteris villosa Peters, Reise nach Mossambique, Saugethiere, p. 48, pi. 11, 
1852. Inhambane, Mozambique. 

Rhinolophus martini Fraser, Proc. Zool. Soc. London, p. 25, July 1843. Fer- 
nando Po. 

Nycteris daubentonii E. Geoffroy, Ann. Mus. d'Hist. Nat., Paris, 20: 19, Aug. 
1813. Senegal. 

Nycteris poensis Gray, Cat. Spec. Mamm. Brit. Mus., p. 24, 13 May 1843 
(nomen nudum). Fernando Po. 

Nycterops pilosa Gray, Proc. Zool. Soc. London, p. 83, May 1866. "Africa." 

Nycteris luteola Thomas 

Nycteris sethiopica luteola Thomas, Ann. Mag. Nat. Hist. (7) 8: 30, July 1901. 
Kitui, Kenya Colony. 
Range: Zanzibar and LTganda, west to Loanda. 

Nycteris macrotis Dobson 

Nycteris macrotis Dobson, Monogr. Asiatic Chiroptera, p. 80, 1876. Sierra 

Leone. 
Nycteris geoffroyi Desmarest, Encyclop. Methodique, Mammalogie, 1: 127, 

1820. Based on an individual from 40 leagues east of Podoo, Senegal, but 

the name also includes the Egyptian thebaica. 
"May be an earlier name" for N. macrotis but "unless the type should have 
been preserved, the identification appears rather doubtful" (Andersen, K., 
Ann. Mag. Nat. Hist. (8) 10: 549, footnote, Nov. 1912). 

Nycteris madagascariensis G. Grandidier 

Nycteris madagascariensis G. Grandidier, Bull. Mus. d'Hist. Nat., Paris, (2) 9: 
353, Nov. 1937. Valley of the Rodo, Madagascar. 



70 bulletin: museum of comparative zoology 

Nycteris major (K. Andersen) 

Petalia major K. Andersen, Ann. Mag. Nat. Hist. (8) 10: 547, Nov. 1912. Ja 
River, Cameroons. 
Range: Northeastern Congo Beige to Cameroons. 

Nycteris marica Kershaw 

Nycteris marica Kershaw, Ann. Mag. Nat. Hist. (9) 12: 534, Oct. 1923. Tindiga, 
Kilosa, Tanganyika Territory. 

Nycteris nana nana (K. Andersen) 

Petalia nana K. Andersen, Ann. Mag. Nat. Hist. (8) 10: 547, Nov. 1912. 
Benito River, French Congo. 

Nycteris nana tristis G. M. Allen and Lawrence 

Nycteris nana tristis G. M. Allen and Lawrence, Bull. Mus. Comp. Zool., 79: 
47, 30 Jan. 1936. Kaimosi, Kakamega district, Kenya Colony. 

Nycteris oriana Kershaw 

Nycteris oriana Kershaw, Ann. Mag. Nat. Hist. (9) 10: 179, Aug. 1922. Chi- 
romo, Shir6 Valley, Nyasaland. 

Nycteris pallida J. A. Allen 

Nycteris pallida J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 425, 29 Sep. 1917. 
Farad] e, Congo Beige. 
Range: Northeastern Congo Beige to Liberia. 

Nycteris parish (de Beaux) 

Petalia parisii de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. Nat. Milano, 
62: 254, Feb. 1924. Balli, southern Italian Somaliland. 

Nycteris proxima Lonnberg and Gyldenstolpe 

Nycteris proxima Lonnberg and Gyldenstolpe, Arkivf. Zool., 17B: no. 9, p. 1, 
14 May 1925. Kartoushi, Semliki Valley, Congo Beige. 

Nycteris revoilii Robin 

Nycteris revoilii Robin, Bull. Soc. Philom. Paris, (7) 5: 90, 1881. Somaliland. 

Nycteris labiata Heuglin, Nova Acta Acad. Caes. Leop. -Carol., Halle, 29: no. 8, 
p. 5, 1861. Keren, Bogosland, Abyssinia. Perhaps antedates AT. revoilii 
Robin (Andersen, 1912: 550, footnote). Dobson makes it a synonym of 
N. thebaica. 

Nycteris thebaica thebaica E. Geoffroy 

Nycteris thehaicus E. Geoffroy, Description de I'Egypte, 2: 119, pi. 1, no. 2, 

1818. Egypt. 
Nycteris albiventer Wagner, Schreber's Saugthiere, Suppl., 1 : 439, 1840. Nubia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 71 

Nyderis angolensis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 1870, 
p. 903, 1871. Caconda, Biballa, Rio Coroca, Angola. Probably distinct. 
Range: Egypt and northern Arabia; Island of Corfu. ? Angola. 

Nycteris thebaica aurantiaca (de Beaux) 

Petalia (Nyderis) thebaica aurantiaca de Beaux, Atti Soc. Ital. Sci. Nat. Milano, 
62: 91, July 1923. Archer's Post, Northern Guaso Nyiro, Kenya Colony. 

Nycteris woodi Andersen 

Nycteris woodi Andersen, Ann. Mag. Nat. Hist. (8) 13, 563, June 1914. Chi- 
langa, northwestern Rhodesia, 4100 feet. 



Family MEGADERMID.^. Big-eared Bats 

Review: Andersen, K., and Wroughton, R. C. On the bats of the family 
Megadermatidas. Ann. Mag. Nat. Hist. (7) 19: 129-145, 1907. 



Cardioderma Peters 

Cardioderma Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 488, 1873' 
As a subgenus of Megaderma; type Megaderma cor Peters. 

Cardioderma cor (Peters) 

Megaderma cor Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 194, 1872. 
Abyssinia. 

Lavia Gray. Yellow-winged Bat 

Lavia Gray, Mag. Zool. and Bot., 2: 490, 1838. Genotype Megaderma frons 

E. Geoff roy. 
Livia L. Agassiz, Nomenclator Zool., Mamm., addenda, p. 6, 1846 (misspelling 

of Lavia). 
Megaderma E. Geoflfroy, Ann. Mus. d'Hist. Nat., Paris, 15: 192, 1810 (in part) 

Lavia frons frons (E. Geoffroy) 

Megaderma frons E. Geoffroy, Ann. Mus. d'Hist. Nat., Paris, 15: 192, 1810. 
Senegal. 

Lavia frons affinis Andersen and Wroughton 

Lavia frons affinis Andersen and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 
140, Feb. 1907. Kaka, White Nile, Anglo-Egyptian Sudan. 

Lavia frons rex Miller 

Lavia rex Miller, Proc. Biol. Soc. Washington, 18: 227, 9 Dec. 1905. Taveta, 
Tanganyika Territory. 



72 bulletin: museum of comparative zoology 



Family RHINOLOPHID^. Horseshoe Bats 

Reviews: Andersen, Knud. On some bats of the genus Rhinolophus, with 
remarks on their mutual aflfinities, and descriptions of twenty-six new forms. 
Proc. Zool. Soc. London, 2: 75-145, pi. 3, 4, Oct. 1905. 

Andersen, Knud. A list of the species and subspecies of the genus Rhino- 
lophus, with some notes on their geographical distribution. Ann. Mag. Nat. 
Hist. (7) 16: 648-662, Dec. 1905. 



Rhinolophus Lacepede 

Rhinolophus Lacepede, Tabl. des Mamm., p. 15, 1799. Genotype Vespertilio 

ferrum-equinum Schreber. 
Rhinocrepis Gervais, Dictionnaire Pittoresque d'Hist. Nat., 4: pt. 318, p. 617, 

1836. Genotype Vespertilio ferrum-equinum Schreber, by implication. 
Aquias Gray, Proc. Zool. Soc. London, p. 15, 13 Apl. 1847. Based on Rhino- 
lophus luctus Temminck and R. trifoliatus Temminck. 
Phyllotis Gray, Proc. Zool. Soc. London, p. 81, May 1866 (not of Waterhouse, 

1837). Genotype Rhinolophus philippinensis Waterhouse. 
Coelophyllus Peters, Proc. Zool. Soc. London, for 1866, p. 427, pi. 35, Apl. 1867. 

Genotype Rhinolophus ccelophyllus Peters. 
Euryalus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 225, 1901. As 

a subgenus of Rhinolophus; type Rhinolophus m&idyi Matschie = i2. 

euryale Meh61y. 

The simplex group (of 1905) or R. viegaphyllus group of 1918 

Rhinolophus ab^ J. A. Allen 

Rhinolophus abse J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 428, 29 Sept. 
1917. Aba, Uele district, Congo Beige. 

Rhinolophus acrotis acrotis Heuglin 

Rhinolophus acrotis Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 29: 
no. 8, p. 4, 10, 1861. Keren, Eritrea, about 15°45' N., 38°30' E. 

Rhinolophus acrotis andersoni Thomas 

Rhinolophus andersoni Thomas, Ann. Mag. Nat. Hist. (7) 14: 156, Aug. 1904. 

Eastern desert of Egypt, about 22°N., 35°E. 
Rhinolophus acrotis andersoni K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 454, 

Dec. 1904. 

Rhinolophus acrotis brachyqnathus K. Andersen 

Rhinolophus acrotis brachygnathus K. Andersen, Ann. Mag. Nat. Hist. (7) 
15: 73, Jan. 1905. Giza, Egypt. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 73 

Rhinolophus acrotis schwarzi Heim de Balsac 

Rhinolophus acrotis schwarzi Heim de Balsac, Bull. Mus. d'Hist. Nat., Paris, 
(2) 7: 483, Nov. 1934. Djanet, Tassali des Azdjers, Sahara. 

Rhinolophtjs ALCYONE ALCYONE Temminck 

Rhinolophus alcyone Temminck, Esquisses Zool. sur la Cote de Guin6, p. 80 
1852. Boutry River, Gold Coast. 
Type in Leiden Mus., but description has not "one single word of any value 
for identifying the species or determining its aflfinities" — K. Andersen. 

Rhinolophus alcyone alticoltjs Sanborn 

Rhinolophus alcyone alticolus Sanborn, Zool. Ser. Field Mus. Nat. Hist., 20: 
108, 15 Aug. 1936. Mt. Cameroon, Cameroon Mandate, 5800 feet. 

Rhinolophus andreinii Senna 

Rhinolophus andreinii Senna, Archivio Zool. Ital., 2: 256, pi. 16, f. 1; pi. 18, 
f. 7-16, 30 Sept. 1905. Adi Ugri, Eritrea. 

Rhinolophus axillaris J. A. Allen 

Rhinolophus axillaris J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 429, 29 Sept. 

1917. Aba, Uele district, Congo Beige; "apparently a member of the 

landeri group." 

Rhinolophus bembanicus Senna 

Rhinolophus bembanicus Senna, Annuario Mus. Zool. R. Univ. Napoli, (2) 4: 
no. 9, p. 1, 2, text-fig. 1, 2, 27 Mar. 1914. Lake Bengueolo, Rhodesia. 

Rhinolophus blasii Peters 

Rhinolophus blasii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 1866, 
p. 17, 1867. Based on the R. clivosus of Blasius. 

Rhinolophus brockmani Thomas 

Rhinolophus brockmani Thomas, Ann. Mag. Nat. Hist. (8) 5: 192, Feb. 1910. 
Upper Sheikh, British SomalUand, 4500 feet. 

Rhinolophus capensis Lichtenstein 

Rhinolophus capensis Lichtenstein, Verzeichniss d. Doubletten Mus. Berlin, 

p. 4, 1823. Cape of Good Hope. 
Rhinolophus auritus Sundevall, in Grill, Kongl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 2: no. 10, p. 13, footnote, 1860. Belvedere, near Knysna, 

Cape of Good Hope. 
Range: Western part of Cape Colony north to mountains from which the 

coastal rivers drain, east to Winter-Bergen. 



74 bulletin: museum of comparative zoology 

Rhinolophus clivosus Cretzschmar 

Rhinolophus clivosus Cretzschmar, in Riippell's Atlas zu d. Reise im Nordlichen 
Afrika, Saugeth., p. 47, 1826. Mohila, Arabia. 
Range: Red Sea coasts of Arabia and Gulf of Aden, African side (Berbera). 

Rhinolophus darlingi darlinqi K. Andersen 

Rhinolophus darlingi K. Andersen, Ann. Mag. Nat. Hist. (7) 15: 70, Jan. 1905. 
Mazoe, Mashonaland, Rhodesia. 
Range: Across South Africa from Rhodesia to Angola. 

Rhinolophus darlingi barbertonensis Roberts 

Rhinolophus darlingi barbertonensis Roberts, Ann. Transvaal Mus., 10: 59, 31 
Jan. 1924. Louw's Creek, Barberton, South Africa. 

Rhinolophus deckenii Peters 

Rhinolophus deckenii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 
1867, p. 705, 1868. Zanzibar coast. 

Rhinolophus denti Thomas 

Rhinolophus denti Thomas, Ann. Mag. Nat. Hist. (7) 13: 386, May 1904. 
Kuruman, Bechuanaland, South Africa. 

Rhinolophus dobsoni Thomas 

Rhinolophus dobsoni Thomas, Ann. Mag. Nat. Hist. (7) 14: 156, footnote, Aug. 
1904. Kordofan, Anglo-Egyptian Sudan. 

Rhinolophus euryale barbarus (K. Andersen and Matschie) 

Euryalus barbarus K. Andersen and Matschie, Sitzb. Ges. Naturf. Freunde 

Berlin, p. 79, 1904. Tangiers, Morocco. 
Rhinolophus algirus Loche, Exploration Sci. de I'Algerie, Zool., Mammif^res, 
p. 83, 1867. "Jardin Marengo," Algeria. 

This name is not certainly identifiable but may possibly prove to supersede 
barbarus if the types can be found. This, too, is probably the bat intended by 
Heim de Balsac in recording Rhinolophus meyeli { = mehelyi) from Laghouat, 
Algeria, Suppl. .in Bull. Biol, de France et de Belgique, Paris, 21: 320, 1936). 

Range: Coastwise from Tangiers to Tunis. 

Rhinolophus euryale judaicus (K. Andersen and Matschie) 
Euryalus judaicus K. Andersen and Matschie, Sitzb. Ges. Naturf. Freunde, 
Berlin, p. 80, 1904. Jerusalem, Palestine, Cave Adullam. 
Range: Palestine to Lower Egypt. "Specimens from Lower Egypt are in- 
distinguishable from those of Palestine." — K. Andersen. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS (D 

Rhinolophus etjryale meridionalis (K. Andersen and Matschie) 
Euryalus meridionalis K. Andersen and Matschie, Sitzb. Ges. Naturf. Freunde, 
p. 79, 1904. Algeria ("probably a mountain form"). 

Rhinolophus ferrum-eqtjinum obscurus Cabrera 

Rhinolophus ferrum-equinum obscurus Cabrera, Mem. Soc. Espanola Hist. 
Nat., Madrid, 2: 257, 1904. Valencia, Spain. 
Range: Spanish peninsula, Balearic Islands, and Algeria. 

Rhinolophus foxi Thomas 

Rhinolophus foxi Thomas, Ann. Mag. Nat. Hist. (8) 11: 314, Mar. 1913. 
Kabwir, Bauchi plateau, northern Nigeria. 

Rhinolophus geoffroyii geoffroyii A. Smith 

Rhinolophus geoffroyii A. Smith, Zool. Journ., 4: 433, May 1829. South Africa. 
Replaces R. augur according to Roberts, Ann. Transvaal Mus., 6: 112, Apl. 
1919. 
Rhinolophus augur K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 380, Nov. 1904. 
Kuruman, Bechuanaland, South Africa. 
Range: Orange River area, Bechuanaland to Transvaal. 

Rhinolophus geoffroyii keniensis HoUister 

Rhinolophus keniensis Hollister, Smithsonian Misc. Coll., 66: no. 1, p. 2, 10 
Feb. 1916. Mount Kenya, Kenya Colony, 7000 feet. 

Rhinolophus geoffroyii zambesiensis K. Andersen 

Rhinolophus augur zambesiensis K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 383, 
Nov. 1904. Fort Hill, northern Nyasaland. 

Rhinolophus geoffroyii zuluensis K. Andersen 

Rhinolophus augur zuluensis K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 383, 
Nov. 1904. Jususic Valley, 20 miles northwest of Eshowe, Zululand, 

Natal. 

Rhinolophus landeri Martin 

Rhinolophus landeri Martin, Proc. Zool. Soc. London, for 1837, p. 101, 25 May 
1838. Fernando Po. 

Rhinolophus lobatus Peters 

Rhinolophus lobatus Peters, Reise nach Mossambique, Saugethiere, p. 41, pi. 9; 
pi. 13, f. 16, 17, 1852. Sena, Tette, Mozambique. 

Rhinolophus simulator K. Andersen 

Rhinolophus simulator K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 384, 
Nov. 1904. Mazoe, Mashonaland, Rhodesia. 



76 bulletin: museum of comparative zoology 

The Rhinolophus lepidus group (of 1905) or 
R. pusillus group of 1918 

Rhinolophus empusa K. Andersen 

Rhinolophus empusa K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 378, Nov. 1904. 
Zomba, Nyasaland. 



The Rhinolophus ?nidas group (of 1905) or 
R. hipposideros group of 1918 

Rhinolophus hipposideros escaler^ K. Andersen 

Rhinolophus hipposideros escalerse K. Andersen, Ann. Mag. Nat. Hist. (9) 2: 
378, Oct. 1918. Ha-ha, Mogador, Morocco. 

Rhinolophus hipposideros minimus Heuglin 

Rhinolophus minimus Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 29: 

no. 8, p. 4, 6, 1861. Keren, Eritrea. 
Rhinolophus bihastatus E. Geoffroy, Description de I'Egypte, 2: 132, 1818; 

Ann. Mus. d'Hist. Nat., Paris, 20:265, 1813. Europe. 
Rhinolophus hipposiderus minimus K. Andersen, Ann. Mag. Nat. Hist. (7) 14: 

455, Dec. 1904. 
Range: Mediterranean subregion, south to Sennaar, east to Eritrea; not 

known from Egypt. 

Rhinolophus hipposideros vespa Laurent 

Rhinolophus hipposideros vespa Laurent, Bull. Soc. d'Hist. Nat. de I'Afrique 
du Nord, 28: 157, text f. 1-2, Feb. 1937. Korifla, Morocco. 



The R. philippinensis group (of 1905) or 
the R. luctus group of 1918 

Rhinolophus maclaudi Pousargues 

Rhinolophus maclaudi Pousargues, Bull. Mus. d'Hist. Nat., Paris, 3: 358, f. 1,2, 
1897. Konakri Island, French Guinea. 



The Rhinolophus macrotis group 

Rhinolophus ^thiops Peters 

Rhinolophus selhiops Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 637, 
1868 ( = 1869?). Otjimbingue, Southwest Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 77 

Rhinolophtjs eloquens K. Andersen 

Rhinolophus hildebrandti eloquens K. Andersen, Ann. Mag. Nat. Hist. (7) 15: 

74, Jan. 1905. Entebbe, Uganda. 
Rhinolophus eloquens Andersen, Ann. Mag. Nat. Hist. (7) 16: 651, Dec. 1905. 

Rhinolophus fumigatus fumigatus Riippell 

Rhinolophus fumigatus Riippell, Museum Senckenburgianum, 3 : 132, 155, 1842. 

Shoa. 
Rhinolophus macrocephalus Heuglin, Reise in Nordost-Afrika, 2: 22, 1877. 

Adowa, Abyssinia. 
Rhinolophus antinorii Dobson, Ann. Mus. Civ. Stor. Nat. Geneva, (2) 2: 16, 

22 May 1885. Daimbi, Shoa. 
Range: Somaliland, Abyssinia, Eritrea. 

Rhinolophus fumigatus exsul K. Andersen 

Rhinolophus fumigatus exsul K. Andersen, Ann. Mag. Nat. Hist. (7) 15: 74, 
Jan. 1905. Kitui, Kenya Colony. 

Rhinolophus hildebrandtii hildebrandtii Peters 

Rhinolophus hildebrandtii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 
p. 195, pi. 1, f. 1, la, 1878. Ndi, Taita, Kenya Colony. 

Rhinolophus hildebrandtii perauritus de Beaux 

Rhinolophus hildebrandti perauritus de Beaux, Atti See. Ital. Sci. Nat. e Mus. 

Civ. Stor. Nat. Milano, 61 : 22, Feb. 1922. Territory of the Rahanuin, 

southern Somaliland. 



, Group uncertain 

Rhinolophus angolensis Seabra 

Rhinolophus angolensis Seabra, Jorn. de Sci. Math., Phys. e Nat., Lisboa (2) 
5: 250, 1898; (2) 6: 117, Aug. 1900. Hanha, Angola. 

Rhinolophus swinnyi swinnyi Gough 

Rhinolophus swinnyi Gough, Ann. Transvaal Mus., 1: 71, 2 text-f., Apl. 1908. 
Ngqeleni district, western Pondoland, South Africa. 

Rhinolophus swinnyi piriensis Hewitt 

Rhinolophus swinnyi piriensis Hewitt, Albany Mus. Records, 2: 402, 6 Feb. 
1913. Pirie, eastern Cape Colony, South Africa. 



78 bulletin: museum of comparative zoology 

Family HIPPOSIDERID.E. Horseshoe Bats 

Asellia Gray. Trident Bats 

Asellia Gray, Mag. Zool. and Bot., 2: 493, 1838 (as a subgenus of Hipposideros) ; 
Gray, List Spec. Mamm. Brit. Mus., p. 24, 13 May 1843 (as a genus). 
Genotype Rhinolophus tridens E. Geoffroy. 

Asellia patrizii de Beaux 

Asellia patrizii de Beaux, Ann. Mus. Civ. Stor. Nat. Genova, 55: 186, 6 June 
1931. Gaarre, Dancalia, Ethiopia. 

Asellia tridens tridens (E. Geoffroy) 

Rhinolophus tridens E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 20: 265, 1813. 
Description de I'Egypte, 2: 130, 1818. Egypt. 
Range: Northeastern Africa, Egypt to Zanzibar. 

Asellia tridens diltjta K. Andersen 

Asellia tridens diluia K. Andersen, Ann. Mag. Nat. Hist. (9) 2: 379, Oct. 1918. 
El Golea, Algerian Sahara. 

Asellia tridens italosomalica de Beaux 

Asellia tridens italosomalica de Beaux, Ann. Mus. Civ. Stor. Nat. Genova, 55: 
190, 6 June 1931. Oddur, Italian Somaliland. 

Clceotis Thomas 

Cteofis Thomas, Ann. Mag. Nat. Hist. (7) 8: 28, July 1901. Genotype Clceotis 
percivali Thomas. 

Clceotis percivali percivali Thomas 

Clceotis percivali Thomas, Ann. Mag. Nat. Hist. (7) 8: 28, July 1901. Takaungu, 
north of Mombasa, Kenya Colony. 

Clceotis percivali australis Roberts 

Clceotis percivali australis Roberts, Ann. Transvaal Mus., 5: 264, 16 May 1917. 
Mooimeisjesfontein, Rustenburg, South Africa. 



Hipposideros Gray 

Hipposideros Gray, Zool. Miscellany, p. 37, 1831. Genotype Vespertilio speoris 

Schreber. 
Gloionycteris Gray, Proc. Zool. Soc. London, p. 82, May 1866. Type Phyllo- 

rhina armiger Hodgson. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 79 

Phyllorrhina Bonaparte, Iconogr. della Fauna Italica, pt. 21, 3d page, 1837. 

As a subgenus of Rhinolophus ; type Rhinolophus diadema E. Geoffrey. 
Speorifera Gray, Proc. Zool. Soc. London, p. 82, May 1866. Genotype Hippo- 

sideros vulgaris Blyth. 
Chrysonyderis Gray, Proc. Zool. Soc. London, p. 82, May 1866. Genotype 

Hipposideros fulvus Gray. 
Rhinophylla Gray, Proc. Zool. Soc. London, p. 82, May 1866. Genotype 

Phyllorhina labuanensis Tomes. 
Macronycteris Gray, Proc. Zool. Soc. London, p. 82, May 1866. Genotype 

Rhinolophus gigas Wagner. 
Doryrhina Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 314, 1871. Asa 

subgenus; type Phyllorhina cy clops Temminck. 
Sideroderma Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 324, 1871. As 

a subgenus; type Phyllorhina fuliginosa Temminck. 
Syndesmotis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 329, 1871. As 

a subgenus; type Phyllorhina megalotis Heuglin. 
Ptychorhina Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 325, 1871. 

As a subgenus; type Rhinolophus caffer Sundevall. 
Cyclorhina Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 326, 1871. 

As a subgenus; types Phyllorhina obscura and doripe. 
Thyreorhina Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 327, 1871. 

As a subgenus; type Phyllorhina coronata Peters. 
Hipposiderus Blanford, Proc. Zool. Soc. London, for 1887, p. 637, Apl. 1888 

(variant). 

Hipposideros ab.e J. A. Allen 

Hipposideros abae J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 432, 29 Sep. 
1917. Aba, Uele district, Congo Beige. 

Hipposideros beatus K. Andersen 

Hipposiderus beatus K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 279, Mar, 1906. 
Fifteen miles from Benito River, West Africa. 
Range: Cameroons and Liberia. 

Hipposideros caffer caffer (Sundevall) 

Rhinolophus caffer Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. For- 

handl., Stockholm, 3: no. 4, p. 118, 1846. Near Port Natal. 
Phyllorrhina gracilis Peters, Reise nach Mossambique, Saugeth., p. 36, pi. 7, 

f. 1-4; pi. 13, f. 14-15, 1852. Tette, lower Zambesi, Mozambique. 
Phyllorrhina bicornis Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Halle, 29: 

no. 8, p. 4, 7, 1861. Keren, Eritrea. 

Hipposideros caffer angolensis (Seabra) 

Phyllorhina angolensis Seabra, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 5: 
256, Dec. 1898. Rio Coroca, Angola. 



80 bulletin: museum of comparative zoology 

Hipposiderus caffer angolensis K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 279, 
Mar. 1906 (valid race). 

HipPOSiDEROS CAFFER CENTRALIS K. Andersen 

Hipposiderus caffer centralis K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 275, 

277, Mar. 1906. Entebbe, Uganda. 

HipposiDEROS CAFFER GUiNEENSis K. Andersen 

Hipposiderus caffer guineensis K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 275, 

278, Mar. 1906. Como River, seventy miles from Gaboon. 

HipposiDEROS CAFFER NiAPU J. A. Allen 

Hipposideros caffer niapu J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 431, 
29 Sep. 1917. Niapu, northeastern Congo Beige. 

Hipposideros caffer tephrus Cabrera 

Hipposiderus tephrus Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 6: 

358, July 1906. Mogador, Morocco. 
Hipposiderus caffer tephrus K. Andersen, Ann. Mus. Civ. Stor. Nat. Genova, 

43: 12, 10 Apl. 1907. 

Hipposideros commersoni commersoni (E. Geoffroy) 

Rhinolophus commersoni E. Geoffroy, Ann. Mus. d'Hist. Nat., Paris, 20: 263, 
pi. 5, 1813. Fort Dauphin, Madagascar. 

Hipposideros commersoni marungensis (Noack) 

Phyllorhina commersonii var. marungensis Noack, Zool. Jahrb., Syst., 2: 272, 

pi. 10, f. 31-33, 7 May 1887. Qua Mpala, Marungu, western Tanganyika 

Territory. 
Hipposideros commersoni mostellum Thomas, Ann. Mag. Nat. Hist. (7) 13: 385, 

May 1904. Tana River, Kenya Colony. 

Hipposideros commersoni thomensis (Bocage) 

Phyllorhina commersoni, var. thomensis Bocage, Jorn. Sci. Math., Phys. e Nat., 

Lisboa, (2) 2: 88, Sep. 1891; (2) 7: 67, 1904. Island of S. Thome. 
Hipposiderus thomensis K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 41, Jan. 

1906. 

Hipposideros cyclops (Temminck) 

Phyllorrhina cyclops Temminck, Esquisses Zool. sur la Cote de Guine, p. 75, 
1853. Boutry River, Gold Coast. 

Rhinolophus micaceus De Winton, Ann. Mag. Nat. Hist. (6) 20: 524, Dec. 1897. 
Como River, 75 miles from Gaboon. Synonym of Hipposideros cyclops, see 
De Winton, Ann. Mag. Nat. Hist. (7) 4: 454, footnote, 1899. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 81 

Hipposideros langi J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37 : 434, text-fig. 
4-6, 29 Sep. 1917. Avakubi, northeastern Congo Beige. 
See Hayman, R.W., Ann. Mag. Nat. Hist. (10) 15: 47, Jan. 1935, on status. 

Hipposideros fuliginosus (Temminck) 

Phyllorrhina fuliginosa Temminck, Esquisses Zool. sur la Cote de Guine, p. 77, 

1853. "La cote de Guine." 
Hipposiderus fuliginosus K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 271, 

Mar. 1906 (valid species). | 

Hipposideros gigas gigas (Wagner) 

Rhinolophus gigas Wagner, Arch. f. Naturgesch., 11: sect. 1, p. 148, 1845. 

Benguela, Angola. 
Phyllorrhina vittata Peters, Reise nach Mossambique, Saugeth., p. 32, pi. 6; 

pi. 13, f. 7-13, 1852. Ibo Island, Cap Delgado group, 12° 20' S. (in part, 

as to the male co-type). 

Hipposideros gigas gambiensis K. Andersen 

Hipposiderus gigas gambiensis K. Andersen, Ann. Mag. Nat. Hist. (7) 17: 42, 
Jan. 1906. Gambia. 

Hipposideros gigas niangar^ J. A. Allen 

Hipposideros gigas niangarse J . A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 438, 
pi. 51, f. 1, 29 Sep. 1917. Niangara, Uele district, Congo Beige. 

Hipposideros megalotis (Heuglin) 

Phyllorrhina megalotis Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 29: 
no. 8, p. 4, 8, 1861. Bogos Land, Eritrea. 

Hipposideros nanus J. A. Allen 

Hipposideros nanus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 434, 29 Sep. 
1917. Faradje, Uele district, eastern Congo Beige. 

Hipposideros ruber (Noack) 

Phyllorhina rubra Noack, Zool. Jahrb., Syst., 7: 586, pi. 18, f. 14, 15, 23 Dec. 

1893. "Lugerrunjere Fluss," Tanganyika Territory. 
Hipposideros ruber Hollister, Bull. U. S. Nat. Mus., no. 99, part 1, p. 85, 1918 

(distinct species from H. caffer) . 



Unidentifiable 

Hipposideros sandersoni Sanderson, Animal Treasure, New York (Viking 
Press), p. 290 (fig.), p. 296 ("bluish-grey and small"), Sep. 1937. Near 
Mamfe, Nigeria. 



82 bulletin: museum of comparative zoology 



Tri^nops Dobson 

Trisenops Dobson, Journ. Asiatic Soc. Bengal, 40; pt. 2, 455, pi. 28, 29 Dec. 
1871. Genotype Trisenops persicus Dobson. 

Tri^nops afer Peters 

Trisenops afer Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 1876, p. 913, 
f. 2, 1877. Mombasa, Kenya Colony. 

Tri^nops furctjla Trouessart 

Trisenops furcula Trouessart, Bull. Mus. d'Hist. Nat. Paris, for 1906, no. 7, 

p. 446, 1906. Grotto of Sarondrana near Tulear (bay Saint- Augustin), 

west coast of Madagascar. 
Trisenops aurila G. Grandidier, Bull. Mus. d'Hist. Nat. Paris, 18: 8, text-f., 

1912. Near Diego-Suarez, extreme northern Madagascar. 

Trisenops humbloti Milne-Edwards 

Tn'ceriops /ium6Zo<i Milne- Edwards, Comptes Rend. Acad. Sci. Paris, 91: 1035, 
1881. Madagascar (east coast). 

Trlsinops persicus Dobson 

Trisenops persicus Dobson, Journ. Asiatic Soc. Bengal, 40: pt. 2, 455, pi. 28, 
29 Dec. 1871. Near Shiraz, Persia. 
Range: Persia to Egypt. 

Tri^nops rtjfus Milne-Edwards 

Trisenops rufus Milne-Edwards, Comptes Rend. Acad. Sci. Paris, 91: 1035, 
1881. Madagascar (east coast). 



Family MYZOPODID.^. Sucker-footed Bat 

Myzopoda Milne-Edwards and A. Grandidier 

Myzopoda Milne-Edwards and A. Grandidier, Bull. Soc. Philom., Paris, (7) 2: 
220, 1878. Genotype Myzopoda aurita Milne-Edwards and A. Grandidier. 

Myxopoda Dobson, Proc. Zool. Soc. London, for 1878, p. 871, Apl. 1879 (lapsus 
calami). 

Myzopoda aurita Milne-Edwards and A. Grandidier 

Myzopoda aurita Milne-Edwards and A. Grandidier, Bull. Soc. Philom., Paris, 
(7) 2: 220, 1878. Madagascar. 
Range: Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 83 

Family VESPERTILIONIDAE. Simple-nosed Bats 

Barbastella Gray. Barbastelle 

Barbastella Gray, London Medical Repository, 15: 300, 1 Apl. 1821. Genotype 

Vespertilio barbastellus Schreber. 
Synotus Keyserling and Blasius, Arch. f. Naturgesch., 5: sect. 1, 305, 1839. 

Genotype Vespertilio barbastellus Schreber. 

Barbastella barbastellus (Schreber) 

Vespertilio barbastellus Schreber, Saugthiere, 1: 168, pi. 55, 1775. Burgundy 

and Germany. 
Vespertilio leucomelas Cretzschmar, Riippell's Atlas, Reise im Nordl. Afrika, 

Saugeth., p. 73, pi. 28b, 1826. Arabia Petraea. 

CiSTUGO Thomas 

Cistugo Thomas, Ann. Mag. Nat. Hist. (8) 10: 205, Aug. 1912. Genotype 
Cistugo seabrse Thomas. 

Cistugo lesueuri Roberts 

Cistugo lesueuri Roberts, Ann. Transvaal Mus., 6: 112, Apl. 1919. Lormarins, 
Paarl district, Cape Colony. 

Cistugo seabrjE Thomas 

Cistugo seabrx Thomas, Ann. Mag. Nat. Hist. (8) 10: 205, Aug. 1912. Mos^ 
samedes, Angola. 

Eptesicops Roberts 

Eptesicops Roberts, Ann. Transvaal Mus., 11: 245, 14 Sep. 1926. Genotype 
Scotophilus rusticus Tomes. 

Eptesicops rusticus (Tomes). 

Scotophilus rusticus Tomes, Proc. Zool. Soc. London, pp. 31, 35, May 1861. 
Damaraland, Southwest Africa; lectotype from Olifants Vlei. 

Eptesicus Rafinesque. Serotine Bats 

Eptesicus Rafinesque, Annals of Nature, p. 2, 1820. Genotype Eptesicus 

melanops Rafinesque = Vespertilio fuscus Beauvois. 
Cnephasus Kaup, Skizzirte Entwick.-Gesch. u. Natiirlich. Syst. d. Europ. 

Thierwelt, 1: 103, 1829. Genotype Vespertilio serotinus Schreber. 
Meteorus Kolenati, Allgemein. Deutsch. Naturh. Zeitung, Dresden, (2) 2: 

131, 1856 (in part). 



84 bulletin: museum of comparative zoology 

Cateorus Kolenati, Allgemein. Deutsch. Naturh. Zeitung, Dresden, (2) 2: 

131, 1856. Genotype Vespertilio serotinus Schreber. 
Amblyotus Kolenati, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 29: 252, 

1858. Genotype Amblyotus atratus }^o\enati = Eptesicus nilssoni (Key- 

serling and Blasius). 
Scotophilus H. Allen, Monogr. Bats No. Amer., Smithsonian Misc. Coll., 

no. 165, p. 27, June 1864 (in part). 
Pachyomus Gray, Ann. Mag. Nat. Hist. (3) 17: 90, Feb. 1866. Genotype 

Scotophilus pachyomus Tomes. 
Nyctiptenus Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 62: sect. 1, 

424, 1870. Genotype Vespertilio smiihii Wagner. 
Vesperus Dobson, Cat. Chiroptera Brit. Mus., p. 184, 1878 (as a subgenus of 

Vesperugo; in part). 
Adelonycteris H. Allen, Proc. Acad. Nat. Sci. Philadelphia for 1891, p. 466, 

19 Jan. 1892 (in part; substitute for Vesperus, preoccupied). 
Neoromicia Roberts, Ann. Transvaal Mus., 11: 245, 14 Sep. 1926. Genotype 

Eptesicus zuluensis Roberts. 
Vespertilio and Vesperugo of various older authors, in part. 

Eptesicus ater J. A. Allen 

Eptesicus ater J. A. Allen, Bull. Amer. Mus. Nat. Hist. 37: 443, 29 Sep. 1917. 
Faradje, northeastern Congo Beige. 

Eptesicus bicolor (Bocage) 

Vesperus bicolor Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 1: 5, Mar. 
1889. Caconda, Angola. 

Eptesicus brunneus (Thomas) 

Vesperugo (Vesperus) brunneus Thomas, Ann. Mag. Nat. Hist. (5) 6: 165, 
1 text-f., Aug. 1880. Old Calabar, West Africa. 

Eptesicus capensis capensis (A. Smith) 

Vespertilio capensis A. Smith, Zool. Journ., 4: 435, Jan.-May 1829. Cape 
Colony, South Africa. 

Eptesicus capensis angolensis Hill 

Eptesicus capensis angolensis Hill, Amer. Mus. Novitates, no. 916, p. 1, 17 
Apl. 1937. Chitau, Angola. 

Eptesicus capensis damarensis (Noack) 

Vesperus damarensis Noack, Zool. Jahrb., Syst., 4: 213, pi. 5, f. 59, 28 Feb. 
1889. Omburo and Golabie, Damaraland, Southwest Africa. 

Eptesicus capensis gracilior (Thomas and Schwann) 

Vespertilio capensis gracilior Thomas and Schwann, Proc. Zool. Soc. London, 
1: 257, 10 Aug. 1905. Eshowe, Zululand, Natal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 85 

Eptesicus capensis nkatiensis Roberts 

Eptesicus capensis nkatiensis Roberts, Ann. Transvaal Mus., 15: 16, 1 Oct. 
1932. Nkate, northern Bechuanaland, South Africa. 

Eptesicus capensis somalicus (Thomas) 

Vespertilio minutus somalicus Thomas, Ann. Mag. Nat. Hist. (7) 8: 32, July 
1901. Hargaisa, British Somaliland. 

Eptesicus capensis somalicus Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 1, p. 92, 
1918. 

Vespertilio hesperida Temminck, Monographies de Mammalogie, 2: 211, 
1835-41. "Lesbordsde la Mer rouge vers les cotes d' Abyssinia." Inde- 
terminate; Heuglin refers this to Miniopterus. 

Vesperugo senarensis Fitzinger and Heuglin, Sitzb. K. Akad. Wiss., Wien, 
math.-nat. CI., 54: sect. 1, p. 546, 1866; Heuglin, Reise in Nordost- 
Afrika, 2: 32, 1877 (no description). Khartoum, Anglo- Egyptian Sudan; 
a nomen nudum. 

Eptesicus faradjius J. A. Allen 

Eptesicus faradjius J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 444, 29 Sep. 
1917. Faradje, northeastern Congo Beige. 

Eptesicus flavescens (Seabra) 

Vesperugo (Vesperus) flavescens Seabra, Jorn. Sci. Math., Phys. e Nat., Lisboa, 
(2) 6: 23, 1900; (2) 6: 119, Aug. 1900. Galanga, Angola". 

Eptesicus garamb^e J. A. Allen 

Eptesicus garamhse J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 445, 29 Sep. 
1917. Garamba, Congo Beige. 

Eptesicus grandidieri (Dobson) 

Vesperugo (Vesperus) grandidieri Dobson, Ann. Mag. Nat. Hist. (4) 18: 500, 
Dec. 1876. Zanzibar. 

Eptesicus guineensis (Bocage) 

Vesperus guineensis Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 1: 
6, Mar. 1889. Bissau, Portuguese Guinea. 

Eptesicus humbloti (Milne-Edwards)" 

Vesperus humbloti MQne-Edwards, Compt. Rendus Acad. Sci. Paris, 91 : 1035, 
1881. Madagascar. 

Eptesicus innesi (Lataste) 

Vesperugo (Vesperus) innesi Lataste, Ann. Mus. Civ. Stor. Nat. Genova, 

(2) 4 (=24): 625, 2 text-figs., 25 July 1887. Cairo, Egypt. 
Eptesicus innesi Thomas, Ann. Mag. Nat. Hist. (9) 4: 350, Nov. 1919. 



86 bulletin: museum of comparative zoology 

Eptesicus loveni Granvik 

Epetiscus (sic) loveni Granvik, Acta Univ. Lund., (2) 21: no. 3, p. 12, 1924. 
Eastern slopes of Mt. Elgon, Kenya Colony, 8000 feet. 

Eptesicus matroka (Thomas and Schwann) 

Vespertilio matroka Thomas and Schwann, Proc. Zool. Soc. London, 1: 258, 
10 Aug. 1905. Ambositra, Betsileo, Madagascar, 1100 meters. 

Eptesicus megalurus (Temminck) 

Vespertilio megalurus Temminck, Monogr. de Mammalogie, 2: 206, 1835-41. 
"L' interieur de 1' Afrique meridionale." 
Range: South Africa to Nyasaland (see Thomas, 1917: 280). 

Eptesicus melckorum Roberts 

Eptesicus melckorum Roberts, Ann. Transvaal Mus., 6: 113, Apl. 1919. Kers- 
fontein. Berg River, Cape Province. 

Eptesicus phasma G. M. Allen 

Eptesicus phasma G. M. Allen, Bull. Mus. Comp. Zool., 54: 327, Dec. 1911. 
Meru River, Northern Guaso Nyiro, Kenya Colony. 

Eptesicus platyops (Thomas) 

Vespertilio platyops Thomas, Ann. Mag. Nat. Hist. (7) 8: 31, July 1901. Lagos; 
Gold Coast. 

Eptesicus pusillus (Leconte) 

Vespertilio pusillus Leconte, Proc. Acad. Nat. Sci., Philadelphia, p. 10, 1857. 

West Africa, doubtless Gaboon. This is probably the same as Vesperugo 

pusillus Noack, in which case the latter becomes a synonym, and the name 

should be credited to Leconte; the description, however, is insufficient for 

identification. 
Vesperugo pusillus Noack, Zool. Jahrb., Syst., 4: 216, pi. 2, f. 2; pi. 5, f. 60-61, 

28 Feb. 1887. Boma, mouth of the Congo, Congo Beige. (?Not Vespertilio 

pusillus Leconte, 1857). 
Vespertilio minuta Temminck, Monogr. de Mammalogie, 2: 209, 1835-41. 

Cape of Good Hope, South Africa. (Not Vespertilio minutus Montagu, 

1808). 

Eptesicus rectitragus Wettstein 

Eptesicus rectitragus Wettstein, Anzeiger K. Akad. Wiss. Wien, 53: 191, 1916. 
Billing, southern Kordofan, Sudan. 

Eptesicus rendalli (Thomas) 

Vesperugo {Vesperus) rendalli Thomas, Ann. Mag. Nat. Hist. (6) 3: 362, fig., 
Apl. 1889; Thomas, Proc. Zool. Soc. London, for 1896, p. 791 and foot- 
note, Apl. 1897. Bathurst, Gambia. 
Range: Mongalla to Nyasaland west to Gambia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 87 

Eptesicus serotinus gabonensis (Trouessart) 

Vesperugo serotinus var. gabonensis Trouessart, Cat. Mamm. Viv. Foss., ed. 2, 

pt. 1, p. 108, 1897. Based on Dobson's Vesperugo serotinus, var. a, in Cat. 

Chiroptera Brit. Mus., p. 192, 1878. Gaboon. 

Eptesicus serotinus isabellinus (Temminck) 

Vespertilio isabellinus Temminck, Monogr. de Mammalogie, 2: 205, pi. 52, f. 1, 
2, 1835-41. "Environs de Tripoli." 

Eptesicus smithii (Wagner) 

Vespertilio smithii Wagner, Schreber's Saugthiere, Suppl., 5: 747, footnote, 
1855. South Africa (based on A. Smith's lUustr. Zool. South Africa, 
1848; larger than E. capensis, see Thomas and Schwann, Proc. Zool. Soc. 
London, 1: 258, 1905). 

Eptesicus tenuipinnis (Peters) 

Vesperiis tenuipinnis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 263, 
1872. Guinea. 

Eptesicus ugand^ HoUister 

Eptesicus ugandse Hollister, Smithsonian Misc. CoU., 66: no. 1, p. 3, 10 Feb. 
1916. Ledgus, Uganda. 

Eptesicus vansoni (Roberts) 

Neoromida vansoni Roberts, Ann. Transvaal Mus., 15: 15, 1 Oct. 1932. 
Zweizwe Waterhole, north of Tsotsoroga Pan, northern Bechuanaland. 

Eptesicus zuluensis Roberts 

Eptesicus zuluensis Roberts, Ann. Transvaal Mus., 10: 60, text-f. 1, 31 Jan. 
1924. White Umfolosi Game Reserve, Zululand, Natal. 



Glauconycteris Dobson 

Glauconycteris Dobson, Proc. Zool. Soc. London, p. 383, Oct. 1875. As a sub- 
genus of Chalinolobus, type Kerivoula poensis Gray; originally included also 
G. variegatus and G. argentatus. 

Glauconycteris alboguttatus J. A. Allen 

Glauconycteris alboguttatus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 449, 
text-fig. 7, B, 29 Sep. 1917. Medje, Congo Beige. 

Glauconycteris argentatus (Dobson) 

Chalinolobus argentatus Dobson, Proc. Zool. Soc. London, p. 385, Oct. 1875. 
Cameroon Mts., French Equatorial Africa. 



88 bulletin: museum of comparative zoology 

Chalinolohus congicus Noack, Zool. Jahrb., Syst., 4: 223, pi. 2, f. 1 ; pi. 5, f. 66-68, 
28 Feb. 1889. Netonna, near mouth of Congo River, French Equatorial 
Africa. (See Thomas, Ann. Mag. Nat. Hist. (8) 11: 144, Jan. 1913, as syn. 
of G. argentatus). 

Glauconycteris BEATRIX Thomas 

Glauconycteris beatrix Thomas, Ann. Mag. Nat. Hist. (7) 8: 256, Sep. 1901. 
Benito River 15 miles from mouth, French Equatorial Africa. 
Range: Cameroons to Entebbe, Uganda. 

Glauconycteris egeria Thomas 

Glauconycteris egeria Thomas, Ann. Mag. Nat. Hist. (8) 11: 144, Jan. 1913. 
Bibundi, Cameroons. 

Glauconycteris humeralis J. A. Allen 

Glauconycteris humeralis J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 448, text- 
fig. 7, A, 29 Sep. 1917. Medje, Congo Beige. 

Glauconycteris poensis (Gray) 

Kerivoula poensis Gray, Ann. Mag. Nat. Hist. (1) 10: 258, Dec. 1842. "Fer- 
nando Po," but in reality Abo, Lower Niger (see Allen and Thomson, 
Exp. Niger, 2: 479, 1848). 

Vesperus (Hesperoptenus) kraussii Peters, Monatsb. K. Preuss. Akad. Wiss., 
Berlin, p. 638, 1868. Yoruba, southern Nigeria (see Thomas, Ann. Mag. 
Nat. Hist. (8) 11: 145, Jan. 1913). 

Glauconycteris variegata variegata (Tomes) 

Scotophilus variegatus Tomes, Proc. Zool. Soc. London, p. 36, May 1861. 
Otjoro, Southwest Africa. 

Glauconycteris variegata papilio Thomas 

Glauconycteris papilio Thomas, Ann. Mag. Nat. Hist. (7) 15: 77, Jan. 1905. 

Entebbe, Uganda. 
Glauconycteris variegata papilio Thomas, Ann. Mag. Nat. Hist. (8) 15: 560, 

footnote, June 1915. 
Range: Zambesi north to Uganda, and thence west to the Gold Coast. 

Glauconycteris variegata phal^en'a Thomas 

Glauconycteris phalsena Thomas, Ann. Mag. Nat. Hist. (8) 15: 560, June 1915. 
Near Fashoda, White Nile, Anglo-Egyptian Sudan. 

L.EPHOTis Thomas 

Lxphotis Thomas, Ann. Mag. Nat. Hist. (7) 7: 460, May 1901. Genotype 
Lsephotis wintoni Thomas. 



I 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 89 

L^PHOTis wiNTONi Thomas 

L^photis wintoni Thomas, Ann. Mag. Nat. Hist. (7) 7: 460, May 1901. Kitui, 
Kenya Colony, 1150 meters. 

MiMETiLLUS Thomas 

Mimetillus Thomas, Abstr. Proc. Zool. Soc. London, no. 10, p. 12, 22 Nov. 

1904; Proc. Zool. Soc. London, for 1904, 2: 188, 18 Apl. 1905. Genotype 

Vesperugo moloneyi Thomas. 
The African representative of Tylonyderis. 
Vesperugo Thomas, Ann. Mag. Nat. Hist. (6) 7: 528, June 1891 (in part). 

Mimetillus berneri Monard 

Mimetillus berneri Monard, Bull. Soc. Neuchateloise des Sci. Nat., Neuchatel, 
57: 49, 1932. Vila da Ponte, Angola. 

Mimetillus moloneyi (Thomas) 

Vesperugo (Vesperus) moloneyi Thomas, Ann. Mag. Nat. Hist. (6) 7: 528, text- 
f., June 1891. Lagos, southern Nigeria. 
Range: Sierra Leone and Fernando Po to Nigeria. 

Mimetillus thomasi Hinton 

Mimetillus thomasi Hinton, Ann. Mag. Nat. Hist. (9) 6: 240, Aug. 1920. 
N'dola, Northern Rhodesia. 



Myotis Kaup. Mouse-eared Bats 

Myotis Kaup, Skizzirte Entw.-Gesch. u. Natiirl. Syst. d. Europ. Thierwelt, 1 : 

106, 1829. Genotype Vespertilio myotis Bechstein. 
Nystades Kaup, Skizzirte Entw.-Gesch. u. Natiirl. Syst. d. Europ. Thierwelt, 

1: 108, 1829 (not of Gloger, 1827). Genotype Vespertilio leisleri Kuhl. 
Leuconoe Boie, Oken's Isis, p. 256, 1830. Genotype "Die Wasserfledermause" 

= Vespertilio daubentonii Kuhl. 
Selysius Bonaparte, Iconogr. d. Fauna Italica, 1: Mamm., Introd., p. 3, 1841. 

Genotype Vespertilio mystacinus Leisler. 
Capaccinius Bonaparte, Iconogr. d. Fauna ItaHca, 1: Indice distributivo [6], 

1841. Genotype Vespertilio capaccinii Bonaparte. 
Trilatitus Gray, Ann. Mag. Nat. Hist. (1) 10: 258, Dec. 1842. Included three 

species. 
Brachyotus Kolenati, Allgemein. Deutsch. Naturhist. Zeitung, Dresden, (2) 

2: 131, 1856 (not of Gould, 1837). Included V. mystacinus, daubentonii, 

and dasycneme. 
Isotus Kolenati, Allgemein. Deutsch. Naturhist. Zeitung, Dresden, (2) 2: 131, 

1856. Included Vespertilio nattereri and emarginatus. 



90 bulletin: museum of comparative zoology 

Tralatitius Gray, Ann. Mag. Nat. Hist. (3) 17: 90, Feb. 1866. Genotype not 

specified. 
Pternopterus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 706, 1867. 

As a subgenus of Vespertilio; type Vespertilio (Pternopterus) lobipes Peters 

= Myotis muricola (Hodgson). 
Exochurus Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 62: sect. 1, 

75, 1870. Included Vespertilio macrodactylus, horsfieldii = adversus, and 

macrotarsus. 
Aeorestes Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 62: sect. 1, 

427, 1870. Included Vespertilio villosissimus, albescens, Isevis and nigricans. 
Comastes Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 62: sect. 1, 

565, 1870. Included Vespertilio capaccinii, megapodius, dasycneme, and 

limnophilus. 
Vespertilio Dobson, Cat. Chiroptera Brit. Mus., p. 284, 1878, and other 

authors, but not of Linnseus. 

Myotis africanus (Dobson) 

Vespertilio africanus Dobson, Ann. Mag. Nat. Hist. (4) 16: 260, Oct. 1875. 
Gaboon. 

Myotis bocagii bocagii (Peters) 

Vespertilio bocagii Peters, Jorn. Sci. Math., Phys. e Nat., Lisboa, (1) 3: 125, 
Dec. 1870. Duque de Bragan^a, Angola. 

Myotis bocagii cupreolus Thomas 

Myotis bocagei cupreolus Thomas, Ann. Mag. Nat. Hist. (7) 13: 407, June 1904. 
Efulen, Cameroons. 

Myotis bocagii hildegarde^ Thomas 

Myotis hildegardese Thomas, Ann. Mag. Nat. Hist. (7) 13: 209, March 1904. 
Fort Hall, Kenya Colony. 

Myotis capaccinii (Bonaparte) 

Vespertilio capaccinii Bonaparte, Iconogr. Fauna Ital., 1 : Mamm., pt. 20 and 
pi., fig. 2, 1837. Sicily. 

Vespertilio megapodius Temminck, Monogr. de Mammalogie, 2: 189, 1839. 
Sardinia. 

Brachyotus blasii Kolenati, Monogr. Europ. Chiroptera, p. 102, 1859. 

Vespertilio majori Ninni, Atti R. 1st. Veneto Sci., Lett, ed Arti, Venezia, (5) 
4: 688, 721, 1878. Venice, Italy. 
Range: Northern Morocco (Tetuan) eastward to Algeria and the Mediter- 
ranean region of southern Europe. 

Note. — Myotis emarginatus (Geoffroy) is credited to Algeria by Lataste 
(Actes Soc. Linn.de Bordeaux, 39: 75, 1885), but the record requires confir- 
mation. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 91 

Myotis goudotii (A. Smith) 

Vespertilio goudotii A. Smith, South African Quart. Journ., 2: 244, June 1834. 

Madagascar. 
Vespertilio madagascariensis Tomes, Proc. Zool. Soc. London, p. 89, 13 Apl. 

1858. Madagascar. 
Vespertilio sylvicola A. Grandidier, Rev. et Mag. Zool., (2) 22: 49, Feb. 1870. 

Forests of Ambaravaranvatou, Madagascar. 

Myotis megalopus (Dobson) 

Vespertilio megalopus Dobson, Ann. Mag. Nat. Hist. (4) 16: 261, Oct. 1875. 
Gaboon. 
The African representative of M. daubentonii,? = Myotis longipes fide Thomas 
1915. 

Myotis oxygnathus (MonticelH) 

Vespertilio oxygnathus MonticeUi, Ann. Accad. O. Costa de Aspir. Nat., Napoli, 
(1) 1: 82, 1885. Matera, Basihcata, Italy. 

Vespertilio murinus Dobson, Cat. Chiroptera Brit. Mus., p. 309, 1878 (in part; 
not of Linnaeus). "It is probable that all specimens from N. Africa hitherto 
referred to M. myotis {Vespertilio murinus of Dobson and earlier authors) 
are really referable to" Myotis oxygnathus (MonticeUi) (O. Thomas, 
Novitates Zool., 20: 29, Feb. 1913). 
Range: Mediterranean region, from Greece to Spain and northern Africa, 
Tunis and Algeria. 

Myotis scotti Thomas 

Myotis scotti Thomas, Ann. Mag. Nat. Hist. (9) 19: 554, May 1927. Djem- 
djem Forest, about 40 miles west of Addis Ababa, Ethiopia, 8000 feet. 

Myotis tricolor (Temminck) 

Vespertilio tricolor Temminck, in Smuts, Enum. Mamm. Capensium, p. 106, 

1832; Monogr. de Mammalogie, 2: 207, 1835-41. Capetown, Cape of 

Good Hope. 

Myotis venustus (Matschie) 

Vespertilio venustus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 74, 1899. 
Kinole, Ukami Mts., Tanganyika Territory. 

Myotis welwitschii (Gray) 

Scotophilus welwitschii Gray, Proc. Zool. Soc. London, p. 211, pi. 24, Sept. 1866. 
Angola. This is very like M. venustus (Matschie). 

Nyctalus Bowdich. Noctule Bats 

Nyctalus Bowdich, Excursions in Madeira and Porto Santo, p. 36, and footnote, 
1825. Genotype Nyctalus verrucosus Bowdich. 



92 bulletin: museum of comparative zoology 

Pterygistes Kaup, Skizzirte Entw.-Gesch. u. Natiirl. Syst. d. Europ. Thierwelt, 
1 : 90, 1829. Genotype Vespertilio noctula Schreber. 

Vesperugo Keyserling and Blasius, Arch. f. Naturgesch., 5: sect. 1, p. 317, 1839 
(in part). As a subgenus of Vespertilio; type not specified; seven species 
included. 

Noctula Bonaparte, Iconogr. Fauna Ital., 1: p. [6], Indice distributivo, 1841. 
Genotype Vespertilio noctula Schreber. 

Noctulinia Gray, Ann. Mag. Nat. Hist. (1) 10: 258, Dec. 1842 (in part). In- 
cluded Noctulinia proterus=Nyctalus noctula, and fulvus = Scotophilus 
kuhlii. 

Panugo Kolenati, Allgemein. Deutsch. Naturhist. Zeitung, Dresden, (2) 2: 131, 
1856. Included Vespertilio noctula and leisleri. 

Noctula Gerbe, Le Naturaliste, 2: 187, 15 March 1880. As a subgenus of Ves- 
perugo = Nyctalus; type Vesperugo plancyi Gerbe. 

Nyctaltjs verrucosus Bowdich 

Nyctalus verrucosus Bowdich, Excursions in Madeira and Porto Santo, p. 36, 

and footnote, 1825. Madeira. 
Vesperugo leisleri Dobson, Cat. Chiroptera Brit. Mus., p. 215, 1878 (in part; 

specimens from Madeira). 
Pterygistes madeine Barrett-Hamilton, Ann. Mag. Nat. Hist. (7) 17: 98, Jan. 

1906. Madeira. 
Range: Madeira Islands. 

Nyctalus macuanus (Peters) 

Vespertilio macuanus Peters, Reisfe nach Mossambique, Saugethiere, p. 61, pi. 
16, f. 1, 1852. Cabaceira peninsula, Mozambique. Reference uncertain. 

Otonycteris Peters. Desert Big-eared Bats 

Otonycteris Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 223, 1859i 
Genotype Otonycteris hemprichii Peters. 

Otonycteris hemprichii Peters 

Otonycteris hemprichii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 223, 
1860. No locality; Hemprich and Ehrenberg's collecting. 

Plecotus ustus Fitzinger and Heuglin, Sitzb. K. K. Akad. Wiss., Wien, math.- 
nat. CI., 54: sect. 1, p. 546, 1866. Heuglin, Reise in Nordost-Afrika, 2: 30, 
1877. Wadi Haifa, in Baten-el-Hadjar, Egypt. A nomen nudum; the type 
is a specimen of Otonycteris (see Anderson, J., and De Winton, W. E., Zool. 
of Egypt, Mamm., p. 118, 1902). 

Plecotus auritus saharse Laurent, Bull. Soc. d'Hist. Nat. de I'Afrique du Nord, 
27: 408, fig., Dec. 1936. El Golea, Algeria. (See Heim de Balsac, H., 
Bull. Soc. d'Hist. Nat. de I'Afrique du Nord, 28: 321, Apl. 1937, as to 
status of this name.) 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 93 

PiPiSTRELLUS Kaup. Pipistrelles 

Pipistrellus Kaup, Skizzirte Entwickl.-Gesch. u. Natiirl. Syst. d. Europ. Thier- 

welt, 1: 98, 1829. Genotype Vesperiilio pipistrellus Schreber. 
Romicia Gray, Mag. Zool. and Bot., 2: 495, 1838. Genotype Romicia calcarata 

Gray = Pipistrellus kuhlii (Kuhl). 
Vesperugo Keyserling and Blasius, Arch. f. Naturgesch., 5: sect. 1, p. 312, 1839 

(in part). Included 13 species now referred to the genera Eptesicus and 

Pipistrellus. 
Romicius Blyth, Cuvier's Animal Kingdom, p. 75, 1840. Variant of Romicia 

Gray. 
Nannugo Kolenati, Allgemein. Deutsch. Naturhist. Zeitung, Dresden, (2) 2: 

131, 1856. Included Vespertilio nathusii, kuhlii, and pipistrellus. 
Alobus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 707, 1867 (not of 

Leconte, 1856). Genotype Vespertilio temminckii = Pipistrellus ruppellii 

(Fischer). 
Hypsugo Kolenati, Allgemein. Deutsch. Naturhist. Zeitung, Dresden, (2) 2: 

131, 1856. Included Vespertilio niaurus and krascheninikoivii. 
Scotophilus H. Allen, Monogr. Bats No. Amer., Smithsonian Misc. Coll., no. 165, 

p. 27, June 1864 (in part). 
Scotozous Dobson, Proc. Zool. Soc. London, p. 372, Oct. 1875 (in part). Geno- 
type Scotozous dormeri Dobson. 

Pipistrellus abaensis J. A. Allen 

Pipistrellus abaensis J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 442, 29 Sep. 
1917. Aba, northeastern Congo Beige. 

Pipistrellus aero Heller 

Pipistrellus aero Heller, Smithsonian Misc. Coll., 60: no. 12, p. 3, 4 Nov. 1912. 
Mt. Gargues, Mathews Range, Kenya Colony. 

Pipistrellus anchiet^ (Seabra) 

Vesperugo anchietae Seabra, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 6: 26, 
120, Aug. 1900. Cahata, Angola. 

Pipistrellus ariel Thomas 

Pipistrellus ariel Thomas, Ann. Mag. Nat. Hist. (7) 14: 157, Aug. 1904. Eastern 
Egyptian desert, lat. 22° N., long. 35° E., 2000 feet. 

Pipistrellus crassulus Thomas 

Pipistrellus crassulus Thomas, Ann. Mag. Nat. Hist. (7) 13: 206, March 1904. 
Efulen, Cameroons. 

Pipistrellus culex Thomas 

Pipistrellus culex Thomas, Ann. Mag. Nat. Hist. (8) 7: 458, May 1911. Kabir, 
northern Nigeria. 



94 bulletin: museum of comparative zoology 

PiPISTRELLTJS DESERTI ThomaS 

Pipistrellus deserti Thomas, Proc. Zool. Soc. London for 1902, 2: 4, Oct. 1902. 
Tripoli, Mursuk. 

Pipistrellus fouriei Thomas 

Pipistrellus fouriei Thomas, Proc. Zool. Soc. London, p. 288, 29 Apl. 1926. 
Ukualukasi, northwest Ovamboland, Southwest Africa. 

Pipistrellus helios Heller 

Pipistrellus helios Heller, Smithsonian Misc. Coll., 60: no. 12, p. 3, 4 Nov. 1912. 
Merelle Water, 30 miles south of Mt. Marsabit, Kenya Colony. 

Pipistrellus kuhlii kuhlii (Natterer) 

Vespertilio kuhlii Natterer in Kuhl, Deutschen Fledermause, in Wetterau. 

Ann., Hanau, p. 55, 1817. Triest, Austria. 
Vespertilio niarginatus Cretzschmar in Riippell's Atlas zu d. Reise Nordl. 

Afrika, Zool., Saugeth., p. 74, pi. 29a, 1826. Arabia Petraea and Nubia; 

according to Anderson, J., and De Winton, W. E. (Zool. of Egypt, Mamm., 

p. 127, 1902), however, on the type specimen in the Frankfort Museum, 

"Egypt is given as the locality." 
Vespertilio pipistrellus, var. /? segyptius J. B. Fischer, Synopsis Mamm., 

p. 105, 1829. "In cryptis Thebarum ^Egypti." 
V esp\ertilio] albo-limbatus Ktister, Oken's Isis, pt. 1, p. 75, 1835. Sardinia. 
Pipistrella tninuta Loche, Exploration Sci. de I'Algerie, Zool., Mammiferes, p. 

78, 1867. Oasis of Messad, southern Algeria. 

Pipistrellus kuhlii fuscatus Thomas 

Pipistrellus kuhlii fuscatus Thomas, Ann. Mag. Nat. Hist. (7) 8: 34, July 1901. 
Naivasha, Kenya Colony. 

Pipistrellus kuhlii pallidus Heim de Balsac 

P[ipistrellus] k\uhlii] pallidus Heim de Balsac, Suppl. au Bull. Biol, de France 

et de Belgique, Paris, 21: 43, 50 (nomen nudum), 180, 1936. Northern 

Sahara, to the A'haggar, Algeria. 

Pipistrellus kuhlii subtilis (Sundevall) 

Vesperugo subtilis Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. Forhandl., 
Stockholm, 3: no. 4, p. 119, 1846. Interior of Caffraria. 

Pipistrellus leucomelas Monard 

Pipistrellus leucomelas Monard, Bull. Soc. Neuchateloise des Sci. Nat., Neuch- 
atel, 57: 47, 1932. Vila da Ponte, Angola. 

Pipistrellus maderensis (Dobson) 

Vesperugo maderensis Dobson, Cat. Chiroptera Brit. Mus., p. 231, pi. 12, f. 5, 
1878. Madeira. 
Range: Madeira and Canary Islands. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 95 

PiPiSTRELLUS MARRENSis Thomas and Hinton 

Pipistrellus marrensis Thomas and Hinton, Proc. Zool. Soc. London, p. 249, 
6 July 1923. Foothills, southern Jebel Marra, Dafur. 

Pipistrellus maurus (Blasius) 

Vesperugo maurus Blasius, Arch. f. Naturgesch., 19: sect. 1, p. 35, 38, 1853. 

Central Alps, Germany. 
Scotophilus darwini Tomes, Proc. Zool. Soc. London, p. 70, (Feb. -June) 1859. 

La Palma, Canary Islands. May be a valid race. 

Pipistrellus musciculus Thomas 

Pipistrellus musciculus Thomas, Ann. Mag. Nat. Hist. (8) 11: 316, March 1913. 
Bitye, Ja River, Cameroons, 2000 feet. 

Pipistrellus nanulus Thomas 

Pipistrellus nanulus Thomas, Ann. Mag. Nat. Hist. (7) 14: 198, Sept. 1904. 
Efulen, Cameroons. 

Pipistrellus nanus nanus (Peters) 

Vespertilio nanus Peters, Reise nach Mossambique, Saugethiere, p. 63, pi. 16, 

f. 2, 1852. Inhambane, Mozambique. 
Vesperugo pusillulus Peters, Jorn. de Sci. Math., Phys. e Nat., Lisboa, (1) 3: 

124, Dec. 1870. Loango, Angola. 

Pipistrellus nanus australis Roberts 

Pipistrellus nanus australis Roberts, Ann. Transvaal Mus., 4: 67, 20 Oct. 1913. 
Port St. Johns, South Africa. 

Pipistrellus pagenstecheri (Noack) 

Vesperugo pagenstecheri Noack, Zool. Jahrb., Syst., 4: 220, pi. 2, f. 3; pi. 5, f. 
64-65, 28 Feb. 1889. Netonna, Banana Creek, Congo mouth, 6° S. 

Pipistrellus pulcher (Dobson) 

Vesperugo pulcher Dobson, Proc. Zool. Soc. London, p. 471, Oct. 1875. Zanzi- 
bar. 

Pipistrellus RfjppELLii ruppellii (J. B. Fischer) 

Vespertilio temminckii Cretzschmar in Rlippell's Atlas zu d. Reise Nordl. 

Afrika, Zool., Saugeth., p. 17, pi. 6, 1826. (Not Horsfield.) Dongola. 
Vespertilio riippelii J. B. Fischer, Synopsis Mamm., p. 109, 1829. Dongola 

(renaming of V. temminckii Cretzschmar). 
Vesperugo sennaariensis Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien, 

math.-nat. CI., 54: pt. 1, p. 546, 1866; nomen nudum. Khartoum. 
Vesperugo senarensis Heuglin, Reise in Nordost-Africa, 2: 32, 1877; nomen 

nudum. 



96 bulletin: museum of comparative zoology 

Vesperugo hypoleucus Heuglin, in Fitzinger and Heuglin, Sitzb. K. Akad. 
Wiss., Wien, math. nat. CI., 54: sect. 1, p. 546, 1866. Sennaar, between 
Kereri, Halfaye and Surerdt on the Nile near Khartoum, Anglo-Egyptian 
Sudan. 

PiPISTRELLUS RXJPPELLII FUSCIPES ThomaS 

Pipistrellus fuscipes Thomas, Ann. Mag. Nat. Hist. (8) 11: 315, March 1913. 
Sixty miles west of Entebbe, Uganda. 

Pipistrellus RiJppELLii vernayi Roberts 

Pipistrellus vernayi Roberts, Ann. Transvaal Mus., 15: 16, 1 Oct. 1932. Maun, 
Ngamiland, South Africa. 

Pipistrellus stampflii (Jentink) 

Vesperugo stampflii Jentink, Notes Leyden Mus., 10: 54, Apl. 1888. Farming- 
ton River, Liberia. 

Pipistrellus minusculus Miller, Proc. Washington Acad. Sci., 2: 647, f. 43, 
28 Dec. 1900. Mount Coffee, Liberia. 



Plecotus E. Geoffroy. Long-eared Bats 

Plecotus E. Geoffroy, Description de I'Egypte, 2: 112, pi. 2, no. 3, 1818. Geno- 
type Vespertilio auritus Linnaeus. 

Plecotus auritus ^gyptiaous I. Geoffroy. Egyptian Big-eared Bat 

Vespertilio auritus /9. segyptius J. B. Fischer, Synopsis Mamm., p. 117, 1829; 
based on description by E. Geoffroy, Description de I'Egypte, 2: 119, 
1818. Egypt. Not Vespertilio pipistrellus var. ^, xgyptius J. B. Fischer, 
ibid., p. 105, 1829. 

Plecotus segyptiacus I. Geoffroy, Mag. de Zool., 2: cl. 1, p. 2 of description of 
pi. 2-3, 1832, who repeats characters given by E. Geoffroy, in Descrip- 
tion de I'Egypte, 2: 119, 1818. Egypt. 

Plecotus christii Gray, Mag. Zool. and Bot., 2: 495, 1838. "North Africa." 
For Dr. Turnbull Christie. 

Plecotus aethiopicus Heuglin, in Fitzinger and Heuglin, Sitzb. K. K. Akad. 
Wiss. Wien, math.-nat. Cl., 54: sect. 1, p. 546, 1866; Reise in Nordost- 
Afrika, 2: 31, 1877. Bahr-el-Abiad, Anglo- Egyptian Sudan. Nomen 
nudum. 

Range: Egypt, to Aswan. 

Plecotus auritus teneriff^ Barrett-Hamilton 

Plecotus teneriffse Barrett- Hamilton, Ann. Mag. Nat. Hist. (7) 20: 520, Dec. 
1907. Teneriffe, Orotava. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 97 



Rhinopterus Miller. Shagreened Bats 

Rhinopterus Miller, Proc. Biol. Soc. Washington, 19: 85, 4 June 1906. Geno- 
type Glauconycteris floweri De Winton. 

Scabrifer G. M. Allen, Bull. Mus. Comp. Zool., 52: 46, July 1908. Substitute 
for Rhinopterus, not Rhinoptera Kuhl. 

Rhinopterus floweri (De Winton) 

Glauconycteris floweri De Winton, Ann. Mag. Nat. Hist. (7) 7: 46, Jan. 1901. 
Wad Marium near Khartoum, Anglo-Egyptian Sudan. 

Rhinopterus lowei Thomas 

Rhinopterus lowei Thomas, Ann. Mag. Nat. Hist. (8) 15: 559, June 1915. 
Blue Nile, 20 miles above Sennaar, Anglo-Egyptian Sudan. 

Rhinopterus notius (G. M. Allen) 

Scabrifer notius G. M. Allen, Bull. Mus. Comp. Zool., 52: 46, July 1908. Cape 
Town, South Africa. 



ScoTEiNUS Dobson 

Scoteinus Dobson, Proc. Zool. Soc. London, p. 371, Oct. 1875. As a subgenus; 
type Scotophilus emarginatus Dobson; but Scotophilus rueppellii Peters 
and S. greyii Gray (Dobson) were also included. 

Scoteinus schlieffenii schlieffenii (Peters) 

Nycticejus schlieffenii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 223, 

1860. Cairo, Egypt. 
N[ycticejus] adovanus Heuglin, Reise in Nordost-Afrika, 2: 34, 1877. Asam 

River, at Adowa, Tigre, Abyssinia. 
According to Peters, this was Nycticejus schlieffenii. 

Scoteinus schlieffenii albiventer Thomas and Wroughton 

Scoteinus schlieffeni albiventer Thomas and Wroughton, Proc. Zool. Soc. 

London, p. 540, 30 Oct. 1908. Naikhala, Upper Egypt. 
Scotophilus minimus Noack, Zool. Jahrb., Syst., 2: 280, pi. 10, f. 36, 37, 7 May 

1887. Qua Mpala, Marungu, Tanganyika Territory. (See Thomas, Ann. 

Mus. Civ. Stor. Nat., Genova, 29: 87, 3 Feb. 1890, as to status.) 
Nycticeius africanus G. M. Allen, Bull. Mus. Comp. Zool., 54: 328, Dec. 1911. 

Meru River, Northern Guaso Nyiro, Kenya Colony. 

Scoteinus schlieffenii australis Thomas and Wroughton 
Scoteinus schlieffeni australis Thomas and Wroughton, Proc. Zool. Soc. London, 
p. 539, 30 Oct. 1908. Inhambane, Mozambique. 



98 bulletin: museum of comparative zoology 

SCOTEINUS SCHLIEFFENII FITZSIMONSI RobcrtS 

Scoteinus schlieffeni fitzsimonsi Roberts, Ann. Transvaal Mus., 15: 17, 1 Oct. 
1932. Tsotsoroga Pan, northern Bechuanaland, South Africa. 

Note. — The record of Scoteinus pallidus by Thomas, O., and Doria, G. 
(Ann. Mus. Civ. Stor. Nat. Genova, (2) 4: 206, 1887) based on a specimen 
supposed to have been taken at Massawa, Eritrea, proves to have been a 
mistake, as the specimen was wrongly labelled. 



ScoTCECUS Thomas 

Scotoecus Thomas, Ann. Mag. Nat. Hist. (7) 7: 263, March 1901. Genotype 
Scotophilus albofuscus Thomas. 

ScoTCECUS ALBiGULA Thomas 

Scotoecus albigula Thomas, Ann. Mag. Nat. Hist. (8) 4: 544, Dec. 1909. Kirui, 
Mt. Elgon, Kenya Colony, 6000 feet. 

ScoTCECUS ALBOFUSCUS (Thomas) 

Scotophilus albofuscus Thomas, Ann. Mus. Civ. Stor. Nat. Genova, 29: 84, 
3 Feb. 1890. Bathurst, Gambia. 

ScoTCECus ARTiNii de Beaux 

Scotoecus artinii de Beaux, Atti Soc. Ital. Sci. Nat. Milano, 62: 98, July 1923. 
Archer's Post, Northern Guaso Nyiro, Kenya Colony. 

ScoTCECus ciNNAMOMEUs Wettstcin 

Scotoecus cinnamomeus Wettstein, Anzeiger K. Akad. Wiss. Wien, 53: 191, 1916. 
Nubbaka, Kordofan, Anglo-Egyptian Sudan. 

ScoTCECUS falabjE Thomas 

Scotoecus falabse Thomas, Ann. Mag. Nat. Hist. (8) 16: 447, Nov. 1915. Kabwir, 
northern Nigeria. 

ScoTCECUS HiNDEi Thomas 

Scotoecus hindei Thomas, Ann. Mag. Nat. Hist. (7) 7: 264, Mar. 1901. Kitui, 
Kenya Colony, 1150 meters. 
For S. L. Hinde. 

ScoTCECUS HiRUNDO (De Winton) 

Scotophilus hirundo De Winton, Ann. Mag. Nat. Hist. (7) 4: 355, Nov. 1899. 
Gambaga, Gold Coast. 

ScoTCECus wooDi Thomas 

Scotoecus woodi Thomas, Ann. Mag. Nat. Hist. (8) 19: 280, Mar. 1917. Chi- 
romo, southern Nyasaland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 99 



ScoTOPHiLUS Leach. Brown Bats 

Scotophilus Leach, Trans. Linn. Soc. London, 13: 69, 182L Genotype Scoto- 

philus kuhlii Leach. (Name not invahdated by Scotophila Hiibner, 1816, 

in entomology.) 
Pachjotus Gray, Zool. Miscellany, no. 1, p. 38, 1831 (in part, to include Nyctice- 

jus and Scotophilus); Miller, Bull. U. S. Nat. Mus., no. 57, p. 219, 1907. 

Genotype Scotophilus kuhlii Leach. 

Review: Thomas, O., and Wroughton, R. C, Proc. Zool. Soc. London, 2: 
539, 30 Oct. 1908 (South African forms). 

Scotophilus borbonicus (E. Geoffroy) 

Vespertilio borbonicus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 8: 201, pi. 46, 
1806. "Ile-Bourbon," i.e. Mauritius. 

Scotophilus gigas Dobson 

Scotophilus gigas Dobson, Ann. Mag. Nat. Hist. (4) 16: 122, Aug. 1875. Lagos, 
southern Nigeria. 
Range: Gold Coast to Nyasaland. 

Scotophilus nigrita nigrita (Schreber) 

Vespertilio nigrita Schreber, Saugthiere, 1: 171, pi. 58, 1775. Senegal. 

Scotophilus nigrita colias Thomas 

Scotophilus nigrita colias Thomas, Ann. Mag. Nat. Hist. (7) 13: 207, Mar, 
1904. Fort Hall, Kenya Colony. 

Scotophilus nigrita dinganii (Wahlberg) * 

Vesperugo dinganii Wahlberg, in Sundevall, Ofversigt af Kongl. Svenska 
Vet.-Akad. Forhandl., Stockholm, 3: no. 4, 119, 1846. The mountainous 
interior of Caffraria, near the Tropic ("nomine Ducis Caffrorum ap- 
pellatus")- 

Scotophilus nigrita herero Thomas 

Scotophilus nigrita herero Thomas, Ann. Mag. Nat. Hist. (7) 17: 174, Feb. 1906. 

Olifant Vlei, about 18° S., 17°30' W., northern Damaraland, Southwest 

Africa. 

Scotophilus nigrita leucogaster (Cretzschmar) 

Nycticejus leucogaster Cretzschmar, in Riippell, Atlas zu d. Reise im Nordl. 
Afrika, Saugeth., p. 71, pi. 28a, 1826. Kordofan. 

Nycticejus serratus Heuglin, Reise in Nordost-Afrika, 2: 35, 1877. "Scherq el 
Aqabah, am Araschkol und den benachbarten Granitgebirgen," Abys- 
sinia. 



100 bulletin: museum of comparative zoology 

ScoTOPHiLUS NiGRiTA Nux Thomas 

Scotophilus nigrita nux Thomas, Ann. Mag. Nat. Hist. (7) 13: 208, Mar. 1904. 
Efulen, Cameroons. 

Scotophilus nigrita planirostris (Peters) 

Nycticejus planirostris Peters, Reise nach Mossambique, Saugethiere, p. 65, 
pi. 17, f. 1, a-e, 1852. Tette, Mozambique. 

Scotophilus nigritellus De Winton 

Scotophilus nigritellus De Winton, Ann. Mag. Nat. Hist. (7) 4: 355, Nov. 1899. 
Gambaga, Gold Coast. 

Scotophilus robustus Milne-Edwards 

Scotophilus robustus Milne-Edwards, Compt. Rend. Acad. Sci. Paris, 91 : 
1035, 1881. Madagascar. 

Scotophilus viridis viridis (Peters) 

Nycticejus viridis Peters, Reise nach Mossambique, Saugeth., p. 67, pi. 17, 
f. 2, a-e, 1852. Island of Mozambique. 

Scotophilus viridis damarensis Thomas 

Scotophilus damarensis Thomas, Ann. Mag. Nat. Hist. (7) 17: 175, Feb. 1906. 

Olifant Vlei, northern Damaraland, Southwest Africa. 
Scotophilus viridis damarensis Thomas and Wroughton, Proc. Zool. Soc. 

London, 2: 539, 30 Oct. 1908. 

Scotophilus murino-flavus (Heuglin). Smaller Brown Bat. 

Nycticejus murino-flavus Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 

29: no. 8, pp. 5, 15, 1861. M'kullu, coast of Abyssinia (Ethiopia). 
Nycticejus flavigaster Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Halle, 29: 

no. 8, p. 5, 14, 1861. Keren, Eritrea. 
Scotophilus altilis G. M. Allen, Bull. Mus. Comp. Zool., 58: 350, July 1914. 

Aradeiba, above Roseires, Blue Nile, Anglo-Egyptian Sudan. 
? syn. of murino-flavus Heuglin. 



Subfamily KERIVOULIN.E. Forest Bats 
Kerivoula Gray 

Kerivoula Gray, Ann. Mag. Nat. Hist. (1) 10: 258, Dec. 1842. Genotype 
Vespertilio hardunckii Horsfield; five other species also included. 

Kirivoula Gervais, Diet. Univ. d'Hist. Nat., 13: 213, 1849. 

Nyctophilax Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. Kl., 42: 390, 
1860. To replace Kerivoula. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 101 

Nyctophylax Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. Kl., 62: sect. 

1, p. 544, Nov. 1870. 
Cerivoula Blanford, Fauna British India, Mamm., p. 338, 1891 (emendatio). 

Kerivoitla aerosa Tomes 

Kerivoula aerosa Tomes, Proc. Zool. Soc. London, p. 333, 9 Nov. 1858. South 
Africa, eastern coast. 

Kerivoula africana Dobson 

Kerivoula africana Dobson, Cat. Chiroptera Brit. Mus., p. 335, 1878. Zanzibar. 

Kerivoula argentata Tomes 

Kerivoula argentata Tomes, Proc. Zool. Soc. London, p. 32, May 1861. Otjoro, 
Southwest Africa. 

Kerivoula brunnea Dobson 

Kerivoula brunnea Dobson, Cat. Chiroptera Brit. Mus., p. 334, 1878. South 
Africa. 

Kerivoula cuprosa Thomas 

Kerivoula cuprosa Thomas, Ann. Mag. Nat. Hist. (8) 10: 41, July 1912. 
Bitye, Ja River, Cameroons. 

?Kerivoula eriophora (Heuglin) 

Nycticejus eriophorus Heuglin, Reise in Nordost-Afrika, 2: 34, 1877. Belegaz 
valley, between Semien and Wogara, Abyssinia. 

Kerivoula harrisoni Thomas 

Kerivoula harrisoni Thomas, Proc. Zool. Soc. London, for 1900, p. 802, footnote, 
1 Apl. 1901. Walamo, between Lakes Zwai and Margherita, north- 
eastern Africa. For J. .1. Harrison, coll. 

Kerivoula lanosa (A. Smith) 

Vespertilio lanosus A. Smith, Illustr. Zool. South Africa, Mamm., pi. 50 and 
text, Dec. 1847. Coast, 200 miles east of Cape Town. 

Kerivoula lucia Hinton 

Kerivoula lucia Hinton, Ann. Mag. Nat. Hist. (9) 6: 240, Aug. 1920. N'dola, 
Northern Rhodesia. 

Kerivoula muscilla Thomas 

Kerivoula muscilla Thomas, Ann. Mag. Nat. Hist. (7) 18: 294, Oct. 1906. 
Ja River, Cameroons. 

Kerivoula nidicola nidicola (Kirk) 

Nycticejus nidicola Kirk, Proc. Zool. Soc. London, for 1864, p. 650, May 1865. 
Shupanga, Zambesia. 



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Kerivoula nidicola zuluensis Roberts 

Kerivoula nidicola zuluensis Roberts, Ann. Transvaal Mus., 10: 61, 31 Jan. 

1924. White Umfolosi River, Zululand, Natal; from nest of a weaver bird, 

with two others. 

Kerivoula phal^na Thomas 

Kerivoula -phalaena Thomas, Ann. Mag. Nat. Hist. (8) 10: 281, Sep. 1912. 
Bibianaha, inland from Denkwa, Gold Coast. 

Kerivoula smithii Thomas 

Kerivoula smithii Thomas, Ann. Mag. Nat. Hist. (5) 6: 166, text-f., Aug. 1880. 
Old Calabar, southern Nigeria. 

Subfamily MINIOPTERIN.^. Long-fingered Bats 

South African forms reviewed: Jameson, H. L., Ann. Mag. Nat. Hist. (8) 4: 
471-472, Nov. 1909. 

MiNiOPTERUS Bonaparte 

Miniopterus Bonaparte, Iconogr. d. Fauna Italica, 1: pt. 21, 1837. As a sub- 
genus of Vespertilio; type Vespertilio ursinii Bonaparte = V. schreibersii 
Kuhl. 

Miniopteris Gray, Ann. Mag. Nat. Hist. (3) 17: 91, Feb. 1866 (variant). 

Minneopterus Lampe, Jahrb. Nassau Ver. Naturkunde, 53: Cat. Saugeth.- 
Sammlung, p. 12, 1900 (lapsus cal.). 

MiNiOPTERUS AFRiCANUS Saubom 

Miniopterus africanus Sanborn, Zool. Ser. Field Mus. Nat. Hist., 20: 111, 15 

Aug. 1936. Sanford's Ranch, Mulo, Shoa district, northwest of Addis 

Ababa, Ethiopia. 

MiNiOPTERUS BREYERI BREYERI JameSOU 

Miniopterus breyeri Jameson, Ann. Mag. Nat. Hist. (8) 4: 471, Nov. 1909. 

Gatkoppies, Waterberg district, Transvaal. 
= Miniopterus natalensis, Roberts, Ann. Transvaal Mus., 4: 68, 20 Oct. 1913. 

MiNiOPTERUS BREYERI VICINIOR J. A. Allen 

Miniopterus breyeri vicinior J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 450, 
29 Sep. 1917. Aba, northeastern Congo Beige. 

Miniopterus fraterculus Thomas and Schwann 

Miniopterus fraterculus Thomas and Schwann, Proc. Zool. Soc. London, 1: 162, 
7 June 1906. Knysna, Cape Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 103 

MiNIOPTERTTS INFLATUS ThomaS 

Miniopterus inflatus Thomas, Ann. Mag. Nat. Hist. (7) 12: 634, Dec. 1903. 
Efulen, Cameroons. 

Miniopterus majori Thomas 

Miniopterus majori Thomas, Ann. Mag. Nat. Hist. (7) 17: 175, Feb. 1906. 
Imasindrary, northeastern Betsileo, Madagascar. 

Miniopterus manavi Thomas 

Miniopterus manavi Thomas, Ann. Mag. Nat. Hist. (7) 17: 176, Feb. 1906. 
Imasindrary, northeastern Betsileo, Madagascar. 

Miniopterus minor Peters 

Miniopterus minor Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, for 1866, 
p. 885, 1867. Coast of Zanzibar. 

Miniopterus natalensis natalensis (A. Smith) 

Vespertilio natalensis A. Smith, South African Quart. Journ., 2: 59, 1934. 

Natal. 

Miniopterus natalensis arenarius Heller 

Miniopterus natalensis arenarius Heller, Smithsonian Misc. Coll., 60: no. 12, 
p. 2, 4 Nov. 1912. Guaso Nyiiki, Northern Guaso Nyiro, Kenya Colony. 

Miniopterus natalensis dasythrix (Temminck) 

Vespertilio dasythrix Temminck, Monogr. de Mammalogie, 2: 268, 1840. 

Interior of Caffraria = South Africa. 
Miniopterus dasythrix A. Smith, Illustr. Zool. So. Africa, Mamm., pi. 52 and 

text, 1848. 

Miniopterus natalensis smitianus Thomas 

Miniopterus smitianus Thomas, Proc. Zool. Soc. London, p. 373, 12 July 1927. 

Witvlei, 40 miles west of Gobabis, eastern Damaraland, Southwest Africa. 
Miniopterus natalensis smitianus Shortridge, Mammals South West Africa, 1 : 

67, 1934. 

Miniopterus newtoni Bocage 

Miniopterus newtoni Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 1: 
198, Dec. 1889. Island of St. Thomas. 

Miniopterus rufus Sanborn 

Miniopterus rufus Sanborn, Zool. Ser. Field Mus. Nat. Hist., 20: 112, 15 Aug. 
1936. Katobwe, 40 miles below Bukama, Lualaba River, Congo Beige. 



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MiNIOPTERUS SCHREIBERSII (Kuhl) 

Vespertilio schreibersii Kuhl, Wetterau. Anstalt, 4: 41, 1819. Germany. 
Range: Southern Europe and Algeria (not known from Egypt). 

MiNioPTERtrs scoTiNUS (Sundevall) 

Vesperugo scotinus Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. For- 
handl., Stockholm, 3: no. 4, p. 119, 1846. Various places in Caffraria. 

Indeterminable 

Vespertilio epichrysus Temminck, in Smuts, Enum. Mamm. Cap., p. 106, 

1832. Capetown, South Africa. 
Vespertilio platycephalus Temminck, in Smuts, Enum. Mamm. Cap., p. 106, 

1832. Capetown, South Africa. 



Family MOLOSSID^. Free-tailed Bats 

Ch^rephon Dobson 

Chserephon Dobson, Journ. Asiatic Soc. Bengal, 43: pt. 2, 144, 17 Oct. 1874. 
Subgenus of Nyctinomus. Thomas, Journ. Bombay Nat. Hist. Soc, 22: 
91, 21 Apl. 1913 (generic characters keyed, and species listed). Genotype 
Nyctinomus johorensis Dobson. 

Ch^rephon bivittatus (Heuglin) 

Nyctinomus bivittatus Heuglin, Nova Acta Acad. Caes. Leop. -Carol., Halle, 29: 

no. 8, p. 4, 13, 1861. Keren, Eritrea. 
Dysopes talpinus Heuglin, Reise Nordost-Afrika, 2: 28, 1877. Upper Abiad 

River, Anglo-Egyptian Sudan. 
Dysopes hepaticus Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Halle, 31: 

no. 7, p. 14, 1864. Country of the Req natives to the Djur River, i.e., 

Bahr-el-Ghazal, Anglo-Egyptian Sudan. 

Ch^rephon frater J. A. Allen 

Chserephon f rater J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 456, 29 Sep. 
1917. Malela (near Boma), southwestern Congo Beige. 

Ch^rephon gambianus (De Winton) 

Nyctinomus gambianus De Winton, Ann. Mag. Nat. Hist. (7) 7: 39, Jan. 1901. 
Gambia. 

Cbmerephon hindei (Thomas) 

Nyctinomus hindei Thomas, Ann. Mag. Nat. Hist. (7) 13: 210, Mar. 1904. 
Fort Hall, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 105 

Ch^rephon leucogaster (A. Grandidier) 

Nydinomus leucogaster A. Grandidier, Revue et Mag. de Zool., (2) 21: 337, 
Sep. 1869. Madagascar, Mahab, near Menabe. 

Ch^rephon lexjcostigma G. M. Allen 

Chxrephon leucostigma G. M. Allen, Bull. Mus. Comp. Zool., 61: 513, 27 
Feb. 1918. Tananarive, Madagascar. 

Ch^rephon limbatus (Peters) 

Dysopes limbatus Peters, Reise nach Mossambique, Saugeth., p. 56, pi. 14, 

1852. Mozambique Island, Sena, 15-17° S. 
Dysopes dubius Peters, Reise nach Mossambique, Saugeth., p. 60, pi. 15, f. 2, 

1852. Sena (a young individual). Not Nydinomus dubius A. Smith, 1833. 

Ch^rephon major (Trouessart) 

Nydinomus pumilus var. major Trouessart, Cat. Mamm. Viv. Foss., ed. 1, 
pt. 1, p. 146, 1897. "Cataract of the Nile," Anglo-Egyptian Sudan; based 
on Dobson's description of a skin and skull (Cat. Chiroptera Brit. Mus., 
p. 428, 1878), apparently similar to a specimen described by HoUister 
(Bull. U. S. Nat. Mus., no. 99, pt. 1. p. 98, 1918) from Gondokoro, and re- 
garded as identical with Chserephon emini. 

Nydinomus major De Winton, Ann. Mag. Nat. Hist. (7) 7: 40, Jan. 1901. 

Nydinomus emini De Winton, Ann. Mag. Nat. Hist. (7) 7: 40, Jan. 1901. 
"Mosambiro," but actually Usambiro, Tanganyika Territory (see Thomas, 
ibid. (7) 13: 210, footnote. Mar. 1904). 

Ch^rephon NIGERIA Thomas 

Chserephon nigerise Thomas, Ann. Mag. Nat. Hist. (8) 11: 319, Mar. 1913. 
Zaria Province, northern Nigeria. 

Ch^rephon pumilus pumilus (Cretzschmar) 

Dysopes pumilus Cretzschmar, in Rlippell's Atlas, Reise im Nordlichen Afrika, 

Saugeth., p. 69, pi. 27, 1826. Massawa, Eritrea. 
Nydinomus pusilus Lesson, Complements de Buffon, ed. 2, Paris, 1: 328, 

1838. Massawa, Eritrea. Error for Dysopes pumilus Cretzschmar. 

Ch^rephon pumilus elphicki Roberts 

Chasrophon (sic) pumilus elphicki Roberts, Ann. Transvaal Mus., 11: 245, 14 
Sep. 1926. Malelane estate, eastern Transvaal. 

Ch^rephon pumilus naivash^ Hollister 

Chserephon pumilus naivashse Hollister, Smithsonian Misc. Coll., 66: no. 1, 
p. 4, 10 Feb. 1916. Naivasha Station, Kenya Colony. 



106 bulletin: museum of comparative zoology 

Ch^rephon pusillus (Miller) 

Nyctinomus pusillus Miller, Proc. Biol. Soc. Washington, 15: 245, 16 Dec. 

1902. Aldabra Island. Not Nyctinomus pusilus Lesson, 1838, which is a 

misprint for putnilus. 

ChjErephon russatus J. A. Allen 

Chserephon russatus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 458, text- 
fig. 25, 29 Sep. 1917. Medje, Congo Beige. 

Ch^rephon websteri Dollman 

ChcPrephon websteri Dollman, Ann. Mag. Nat. Hist. (8) 2: 546, Dec. 1908. 
Yola, northern Nigeria. 



Subgenus Lophomops J. A. Allen 

Lophotnops J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 460, text-fig. 9-11, 
29 Sep. 1917. Subgenus of Chsprephon; type Chserephon (Lophomops) 
chapini J. A. Allen. 

Ch^rephon ABM J. A. Allen 

Chsprephon (Lophomops) abse J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 464, 
text-fig. 11, 29 Sep. 1917. Aba, northeastern Congo Beige. 

Ch^rephon chapini J. A. Allen 

Chserephon (Lophomops) chapini J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 
461, text-fig. 9, 29 Sep. 1917. Faradje, northeastern Congo Beige. 

Ch^rephon cristatus J. A. Allen 

Chserephon (Lophomops) cristatus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 

463, text-fig. 10 A, B, 29 Sep. 1917. Boma, near mouth of Congo River, 

Congo Beige. 

Ch^rephon lancasteri Hayman 

Chserephon lancasteri HsLyman, Ann. Mag. Nat. Hist. (11) 1: 383, Apl. 1938. 
Lunzi River, Lundazi district, Loangwa Valley, northeastern Rhodesia. 

Ch^rephon langi Roberts 

Chcerephon (Lophomops) langi Roberts, Ann. Transvaal Mus., 15: 17, 1 Oct. 
1932. Tsotsoroga Pan, northern Bechuanaland, South Africa. 

Ch^rephon nigri Hatt 

Chserophon (sic) (Lophomops) nigri Hatt, Bull. Soc. Zool. de France, 53: 374, 

text-f. B, 10 Dec. 1928. Bourem, on the Niger River, Timbuctou district,- 

French Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 107 

Ch^rephon shortridgei Thomas 

Chserephon (Lophomops) shortridgei Thomas, Proc. Zool. Soc. London, p. 289, 

29 Apl. 1926. Ukualukasi, northwestern Ovamboland, Southwest Africa, 

3400 feet. 

Mops Lesson 

Mops Lesson, Nouv. Tabl. Regne Animal, Mamm., p. 18, 1842. Genotype 
Mops indicus hesson = Dysopes mops F. Cuvier. See Thomas, Journ. Bom- 
bay Nat. Hist. Soc, 22: 91, 21 Apl. 1913, for generic characters and list of 
species. 

Mops angolensis angolensis (Peters) 

Nydinomus angolensis Peters, Jorn. Sci. Math., Phys. e Nat., Lisboa, (1) 3: 
124, Dec. 1870. (Quenza River), Angola. 

Mops angolensis aloysii-sabaudi^ (Festa) 

Nydinomus aloysii-sabaudise Festa, Boll. Mus. Zool. ed Anat. Comp. Univ. di 

Torino, 22: no. 546, p. 1, 21 Jan. 1907. Ruwenzori. 
Chserephon angolensis sahaudise de Beaux, Ann. Mus. Civ. Stor. Nat. Genova, 

(3) 9=49:371, 28 Aug. 1922. 

Mops angolensis niveiventer Cabrera and Ruxton 

Mops angolensis niveiventer Cabrera and Ruxton, Ann. Mag. Nat. Hist. (9) 17: 
594, May 1926. St. Joseph de Luluabourg, southern Congo Beige. 

Mops angolensis wonderi Sanborn 

Mops angolensis wonderi Sanborn, Zool. Ser. Field Mus. Nat. Hist., 20: 114, 
15 Aug. 1936. Sotuba, 7 km. east of Bamako, French Sudan. 

Mops brachypterus (Peters) 

Dysopes brachypterus Peters, Reise nach Mossambique, Saugeth., p. 59, pi. 15 
f. 1, 1852. Island of Mozambique, lat. 15° S. 

Mops chitauensis Hill 

Mops chitauensis Hill, Amer. Mus. Novitates, no. 916, pp. 2, 3, text-f. 1, 17 
Apl. 1937. Chitau, Angola. 

Mops congicus J. A. Allen 

Mops congicus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 467, pi. 55, 29 Sep. 
1917. Medje, Congo Beige. 

Mops demonstrator (Thomas) 

Nydinomus demonstrator Thomas, Ann. Mag. Nat. Hist. (7) 12: 504, Nov. 1903. 

Mangala, 25 miles north of Gondokoro, Kordofan, Anglo-Egyptian Sudan 

( = Mongalla). 



108 bulletin: museum of comparative zoology 

Mops leonis (Thomas) 

Nyctinomus leonis Thomas, Ann. Mag. Nat. Hist. (8) 2: 373, Oct. 1908. Sierra 
Leone. 
Range: Sierra Leone to Cameroons, Fernando Po. 

Mops miarensis (A. Grandidier) 

Nyctinomus miarensis A. Grandidier, Revue et Mag. de Zool., (2) 21: 337, Sep. 

1869. Madagascar, Miari, between Soua-haze and Soua-hane. 
Nyctinomus unicolor A. Grandidier, Revue et Mag. de Zool., (2) 22: 49, Feb. 

1870. Madagascar. See Dobson, Cat. Chiroptera Brit. Mus., p. 430, foot- 
note, 1 878, as to this name being a synonym of A'^. miarensis. 

Mops niangar^e J. A. Allen 

Mops niangarx J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 468, 29 Sep. 1917. 
Niangara, northeastern Congo Beige. 

Mops ruppellii (Temminck) 

Dysopes rupelii Temminck (sic), Monographies de Mammalogie, 1 : 224, pi. 18, 

1827. Egypt. 
There seems no doubt that this name must replace midas Sundevall. 
Dysopes midas Sundevall, Kongl. Svenska Vet.-Acad. Handl., Stockholm, for 

1842, p. 207, pi. 2, f. 7, 1843. Bahr-el-Abiad, White Nile, Anglo-Egyptian 

Sudan. 
Nyctinomus (Dysopes) ventralis Heuglin, Nova Acta Acad. Cses. Leop.-Carol., 

Halle, 29: no. 8, p. 4, 11, 1861. Keren, Abyssinia. 
Nyctinomus ruppellii Fitzinger and Heuglin, Sitzb. K. Akad. Wiss. Wien, 

math.-nat. Kl., 54: sect. 1, p. 545, 1866. 

Mops thersites (Thomas) 

Nyctinomus thersites Thomas, Ann. Mag. Nat. Hist. (7) 12: 634, Dec. 1903. 
Efulen, Cameroons. 

Mops trevori (J. A. Allen) 

Mops trevori J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 469, pi. 48, f. 2, 29 
Sep. 1917. Faradje, northeastern Congo Beige. 

Subgenus Allomops J. A. Allen 

Allomops J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 470, text-fig. 12-15, 
29 Sep. 1917. Subgenus of Mops, type Chxrephon {Allomops) osborni J. A. 
Allen = Mops osborni (J. A. Allen). 

Mops faradjius J. A. Allen 

Mops {Allomops) faradjius J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 476, 
29 Sep. 1917. Faradje, northeastern Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 109 

Mops nanulus J. A. Allen 

Mops (Allomops) nanulus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 477, 
29 Sep. 1917. Niangara, Congo Beige. 

Mops occipitalis J. A. Allen 

Mops {Allomops) occipitalis J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 474, 
text-f. 15, 29 Sep. 1917. Avakubi, Congo Beige. 

Mops osborni (J. A. Allen) 

Chserephon (Allomops) osborni J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 

470, text-f. 12-15, 29 Sep. 1917. 
Mops (Allomops) osborni J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 473, 

text-f. 12-14, 26, 29 Sep. 1917. Kinshasa, near Leopoldville, Congo Beige. 

MoRMOPTERUs Petcrs 

Mormopterus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 258, 1865. 
As a subgenus of Dysopes; type Nyctinomus jugularis Peters. 

Mormopterus acetabulosus acetabulosus (Hermann) 

Vespertilio acetabulosus Hermann, Observationes Zoologicse, p. 19, 1804. Port 

Louis, Island of Mauritius (ex Commerson MS.). 
Nyctinomus mauritianus Horsfield, Zool. Researches in Java, pt. 5, 9th page 

of text in acct. of A^". tenuis. Port Louis, Island of Mauritius (renaming of 

Vespertilio acetabulosus Hermann, ex Commerson). 
Nyctinomus (Mormopterus) jugularis Peters, in Sclater, Proc. Zool. Soc. London, 

p. 468, Oct. 1865. Antananarivo, Madagascar. 

Mormopterus acetabulosus natalensis (A. Smith) 

Dysopes natalensis A. Smith, Illustr. Zool. South Africa, Mamm., pi. 49, Dec. 
1847. Near Port Natal, South Africa. 

Mormopterus albiventer (Dobson) 

Nyctinomus albiventer Dobson, Proc. Zool. Soc. London, for 1876, p. 733, Apl. 
1877. Madagascar. 

Mormopterus setiger (Peters) 

M[ormopterus] setiger Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 483, 
pi., f. 3, 1881. Taita, Kenya Colony. 

Myopterus Oken 

Myopterus Oken, Lehrbuch d. Naturgesch., pt. 3: zool., sect. 2, p. 932, 1816. 

Genotype Myopterus senegalensis Oken the "rat-volant" of Daubenton. 

See Miller, Bull. U. S. Nat. Mus., no. 57, p. 246, 1907. 
Myopteris Desmarest, Encyclop. Methodique, Mammalogie, 1: 131, 1820; 

emendation. 



110 bulletin: museum of comparative zoology 

Myoptera Blainville, Compt. Rend. Acad. Sci. Paris, 5: 815, 1837, variant. 
Eomops Thomas, Ann. Mag. Nat. Hist. (7) 16: 572, Nov. 1905. Genotype 
Mormopterus whitleyi Scharff . 

Myopterus albatus Thomas 

Myopterus albatus Thomas, Ann. Mag. Nat. Hist. (8) 16: 469, Dec. 1915. 
River Uele, Congo Beige. 

Myopterus senegalensis Oken 

Myopterus senegalensis Oken, Lehrbuch d. Naturgesch., pt. 3: zool., sect. 2, 

p. 932, 1816. Senegal. Based on Daubenton's account. 
Myopterus dauhentonii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 574, 

1866. Senegal. Peters credits the name to E. Geoffroy, Descrip. de 

I'Egypte, 2: 113, 1818, in which Trouessart follows, but only the generic 

name appears there. 
Myopteris (sic) dauhentonii Desmarest, Encyclop. Methodique, Mammalogie, 

1: 132, 1820. Senegal; based on the "rat-volant" of Daubenton (Mem. 

Acad. Roy. Sci. Paris, p. 386, 1759). 

Myopterus whitleyi (Scharff) 

Mormopterus whitleyi Scharff, Ann. Mag. Nat. Hist. (7) 6: 569, text-f., Dec. 
1900. Benin City, West Africa. 

Nyctinomus E. Geoffroy. Wrinkle-lipped Bats 

Nydinomus E. Geoffroy, Description de I'Egypte, 2: 114, pi. 2, 1818. Geno- 
type Nyctinomus segyptiacus E. Geoffroy. 

Nyctinomus ^qyptiacus E. Geoffroy 

Nyctinomus segyptiacus E. Geoffroy, Description de I'Egypte, 2: 128, pi. 2, 

no. 2, 1818. Egypt. 
Dysopes geoffroyi Temminck, Monogr. de Mammalogie, 1 : 226, pi. 19, 1827. 

Egypt. Substitute for Nyctinomus segyptiacus E. Geoffroy. 

Nyctinomus africanus Dobson 

Nyctinomus africanus Dobson, Ann. Mag. Nat. Hist. (4) 17: 348, May 1876. 
Transvaal. 

Nyctinomus anchiet^ Seabra 

Nyctinomus anchietse Seabra, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 6: 82, 
Aug. 1900. Quibula, Angola. 

Nyctinomus ansorgei Thomas 

Nyctinomus ansorgei Thomas, Ann. Mag. Nat. Hist. (8) 11: 318, Mar. 1913. 
Malange, northern Angola. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 111 

Nyctinomus bemmeleni Jentink 

Nyctinomus bemmeleni Jentink, Notes Leyden Mus., 1 : 125, Apl. 1879. Liberia. 
Nyctinomus vemmeleni Trouessart, Cat. Mamm. Viv. Foss., pt. 1, p. 146, 1897 
(mistake for bemmeleni). 

Nyctinomus bocagei Seabra 

Nyctinomus bocagei Seabra, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 6: 84, 
Aug. 1900. Galanga, Angola. 

Nyctinomus brunneus Seabra 

Nyctinomus brunneus Seabra, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 6: 83, 
Aug. 1900. Quissange, Angola. 

Nyctinomus cisturus Thomas 

Nyctinomus cisturus Thomas, Ann. Mag. Nat. Hist. (7) 12: 502, Nov. 1903. 
Mangala, 25 miles north of Gondokoro, Kordofan, Anglo-Egyptian Sudan. 

Nyctinomus condylurus A. Smith 

Nyctinomus condylurus A. Smith, South African Quart. Journ., 1: 54, 1833. 
Port Natal, South Africa. 
Probably valid for the larger Nyctinomus of South Africa. 

Nyctinomus dubius A. Smith 

Nyctinomus dubius A. Smith, South African Quart. Journ., 1 : 54, 1833. South 
Africa, between Cape Colony and Natal. 
Status uncertain, but probably applicable to some one of the species of 
Chasrephon or Nyctinomus of South Africa. 

Nyctinomus fulminans Thomas 

Nyctinomus fulminans Thomas, Ann. Mag. Nat. Hist. (7) 12: 501, Nov. 1903. 
Fianarantsoa, eastern Betsileo, Madagascar. 

Nyctinomus lobatus Thomas 

Nyctinomus lobatus Thomas, Ann. Mag. Nat. Hist. (6) 7: 303, Mar. 1891. 
Turquel, Siik, Kenya Colony. 

Nyctinomus ochraceus J. A. Allen 

Nyctinomus ochraceus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 37: 455, 29 Sep. 
1917. Medje, northeastern Congo Beige. 

Nyctinomus spillmanni Monard 

Nyctinomus spillmanni Monard, Bull. Soc. Neuchateloise des Sci. Nat., Neuch- 
atel, 57: 51, 1932. Vila da Ponte, Angola. 



112 bulletin: museum of comparative zoology 

Nyctinomus tongaensis Wettstein 

Nydinomus (Nyctinomus) tongaensis Wettstein, Anzeiger K. Akad. Wiss. Wien, 

math.-nat. Kl., 53: 192, 1916. Tonga, Upper White Nile, Anglo-Egyptian 

Sudan. 

Otomops Thomas 

Otomops Thomas, Journ. Bombay Nat. Hist. Soc, 22: 90, 21 Apl. 1913. Geno- 
type Nyctinomus wroughtoni Thomas. 

Otomops icarus Chubb 

Otomops icarus Chubb, Ann. Durban Mus., 1: 433, pi. 21, 21 May 1917. Dur- 
ban, Natal. 

Otomops martiensseni (Matschie) 

Nyctinomus martiensseni Matschie, Arch. f. Naturg., 63: sect. 1, p. 84, Oct. 

1897. Magrotto plantation near Tanga, Tanganyika Territory. The 

African representative of 0. wroughtoni of India. 

Platymops Thomas 
Subgenus Platymops Thomas 

Platymops Thomas, Ann. Mag. Nat. Hist. (7) 17: 499, May 1906. Genotype 
Platymops macmillani Thomas. 

Platymops macmillani Thomas 

Platymops macmillani Thomas, Ann. Mag. Nat. Hist. (7) 17: 500, May 1906. 
Between Addis Ababa and Lake Rudolf, Abyssinia. 

Subgenus Sauromys Roberts 

Sauromys Roberts, Ann. Transvaal Mus., 6: 5, 28 June 1917. As a subgenus 
of Platymops Thomas, with p ^ present; type Platymops haagneri Roberts. 

Platymops haagneri Roberts 

Platymops haagneri Roberts, Ann. Transvaal Mus., 6: 5, 28 June 1917. Keet- 
manshoop, Damaraland, Southwest Africa. 

Platymops petrophiltjs Roberts 

Platymops petrophilus Roberts, Ann. Transvaal Mus., 6: 4, 28 June 1917; 
Bleskop, near Rustenburg, Transvaal. 

Xiphonycteris Dollman 

Xiphonycteris Dollman, Ann. Mag. Nat. Hist. (8) 7: 210, Feb. 1911. Genotype 
Xiphonycteris spxirrelli Dollman. 

Xiphonycteris spurrelli Dollman 

Xiphonycteris spurrelli Dollman, Ann. Mag. Nat. Hist. (8) 7: 211, Feb. 1911. 
Bibianaha, 60 miles west of Kumasi, Gold Coast, Nigeria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 113 

Order PRIMATES. Lemurs and Monkeys, Man 
Suborder LEMUROIDEA. Lemurs 

Lemuriformes Schwarz, Proc. Zool. Soc. London, p. 400, 29 June 1931. 

Family LORISIDiE. Galagos and Slow Lemurs 

Subfamily LORISINiE. Slow Lemurs 

Review: Schwarz, E. On the African short-tailed lemurs or pottos. Ann. 
Mag. Nat. Hist. (10) 8: 249-256, Sep. 1931. 

Arctocebus Gray. Golden Potto 

Arctocebus Gray, Proc. Zool. Soc. London, p. 150, Oct. 1863. Genotype 

Perodicticus calabarensis J. A. Smith. 
Perodicticus, Nycticehus, in part, of older authors. 

Arctocebus calabarensis calabarensis (J. A. Smith) 

Perodicticus calabarensis J. A. Smith, Proc. Roy. Phys. Soc. Edinburgh, 2: 

177, text-f. 1-3, pi. 8, 1860, 1863 on title page. Old Calabar, southern 

Nigeria. 
Range: Cameroon Mt. region, north to the limit of forest, west to the Niger 

and Sanaga Rivers. 

Arctocebus calabarensis aureus De Winton 

Arctocebus aureus De Winton, Ann. Mag. Nat. Hist. (7) 9: 48, Jan. 1902. 

Spanish Guinea, 50 miles from the mouth of the Benito River. 
Arctocebus ruficeps Thomas, Ann. Mag. Nat. Hist. (8) 12: 387, Oct. 1913. 

Metet, near the Nyong River, southern Cameroons. 
Range: Spanish Guinea and the Cameroons. 

Perodicticus Bennett. Pottos 

Perodicticus Bennett, Proc. Zool. Soc. London, p. 109, 1 Sep. 1831; Philos. 

Mag. (2) 10: 389, 1831. Genotype Perodicticus geoffroyi Bennett = Lemur 

potto P.L.S. MuUer. 
Potto Lesson, Species Mamm. Bimanes et Quadrumanes, pp. 207, 237, 1840. 

Genotype Potto bosmanii Lesson = Lemur potto P.L.S. Miiller. 

Perodicticus potto potto (P.L.S. Muller) 

Lemur potto P.L.S. Muller, Linne's Natursyst., Suppl., p. 12, 1766. Elmina, 

Gold Coast (based on the Potto of Bosman). 
Galago guineensis Desmarest, Encyclop. Methodique, Mammalogie, 1: 104, 

1820. Guinea. New name for Lemur potto P.L.S. Muller. 



114 bulletin: museum of comparative zoology 

Perodidicus geoffroyi Bennett, Proc. Zool. Soc. London, p. 109, 1 Sep. 1831. 

Sierra Leone. 
Potto bosmanii Lesson, Species Mamm. Bimanes et Quadrumanes, p. 238, 

1840. Elmina, Gold Coast (renaming oi Lemur potto P. L. S. Miiller). 
Range: Upper Guinea, from the Niger River west to the Gold Coast. 

Perodicticus potto edwardsi a. Bouvier 

Perodidicus edwardsi A. Bouvier, "Guide du Naturaliste", Revue Bibl. Sci. 
Nat., 1 : 10, 1879. North bank of Congo River. 

Perodidicus batesi De Winton, Ann. Mag. Nat. Hist. (7) 9: 48, Jan. 1902. 
"Benito River, 15 miles from the mouth"? = Spanish Guinea. 

Perodicticus ju-ju Thomas, Ann. Mag. Nat. Hist. (8) 5: 351, Apl. 1910. South- 
ern Nigeria. 
Range: Forest region from lower Guinea coast (Cameroons, Rio Muni) east 
to between the Niger, Ubangi, and Congo Rivers. 

Perodicticus potto faustus Thomas 

Perodicticus faustus Thomas, Ann. Mag. Nat. Hist. (8) 6: 426, Oct. 1910. 
Irneti, Bompona, Maringa River, northwestern Congo Beige. 
Range: Forests of left bank of the Congo. 

Perodicticus potto ibeanus Thomas 

Perodidicus ibeanus Thomas, Abstract Proc. Zool. Soc. London, no. 81, p. 17, 

22 Mar. 1910; Proc. Zool. Soc. London, p. 536, 11 Oct. 1910. Kakamega 

Forest, near Mt. Elgon, Kenya Colony. 
Perodictius (sic) arrhenii Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 58: no. 2, p. 45, 1 Sep. 1917. Near Masisi, three days west 

of Lake Ivivu, Congo Beige. 
Periodidicus (sic) nebulosus Lorenz, Ann. K. K. Naturhist. Hofmus. Wien, 31 : 

239, 1917. Ukaika, Ituri forest, Congo Beige. 
Range: Forest region and isolated areas north and east of the Congo, from 

the Ubangi east to Mau forest, Kenya Colony. 



Subfamily GALAGINiE. Galagos or Bush Babies 
Galago E. Geoffroy 

Galago E. Geoffroy, Mag. Encyclopedique, 2e ann., 1 : 49, pi., 1796. Genotype 

Galago senegalensis E. Geoffroy. 
Otolicnus Illiger, Prodromus Syst. Mamm. et Avium, p. 74, 1811. Genotype 

Lemur galago Schreber = Galago senegalensis E. Geoffroy. 
Macropus G. Fischer, Mem. Soc. Imp. des Nat., Moscou, 1: 12, 1811. New 

name for Galago E. Geoffroy, hence genotjrpe Galago senegalensis E. 

Geoffroy. Preoccupied by Macropus Shaw, 1790, for kangaroos. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 115 

Galagoides A. Smith, South African Quart. Journ., (2) 2: 32, 1833. Genotype 
Galago demidovii G. Fischer (see Elliot, Review of the Primates, Mono- 
graphs Amer. Mus. Nat. Hist., 1: xxix, 1912 = 15 June 1913). 

Chirosciurus Gervais, Dictionnaire Pittoresque d' Hist. Nat., 4: pt. 318, p. 617, 
1836. Based on the Khoyak of G. Cuvier and E. Geoffroy, Mag. Encyclop., 
(1) 2: 182, 1795, which is Galago senegalensis E. Geoffroy. 

Hemigalago Dahlbom, Zool. Studier, 1 : pt. 3, p. 224, 225, pi. 10, 1857. Geno- 
type Galago demidovii G. Fischer. 

Otolemur Coquerel, Revue et Mag. de Zool., (2) 11: 458, pi. 17-18, 1859. 
Genotype Otolemur agisymbanus Coqxierel = Galago crassicaudatus agisym,- 
banus (Coquerel). 

Callotus Gray, Proc. Zool. Soc. London, p. 132, 145, 1863. Genotype Calloius 
monteiri Gra,y = Galago crassicaudatus tnonteiri Gray. 
Gray misspells the name Callotis in Proc. Zool. Soc. London, p. 458, 1864. 

Otogale Gray, Proc. Zool. Soc. London, p. 132, 139, Oct. 1863. Type by sub- 
sequent designation Otolicnus garnettii Ogilby = Galago crassicaudatus 
garnettii (Ogilby). 

Sciurocheirus Gray, Proc. Zool. Soc. London, for 1872, p. 857, text-f. 5, Apl. 
1873. Genotype by monotypy Galago alleni Waterhouse. 

Review: Schwarz, E. On the African long-tailed lemurs or galagos. Ann. 
Mag. Nat. Hist. (10) 7: 41-66, Jan. 1931. 

Galago alleni Waterhouse 

Galago alleni Waterhouse, Proc. Zool. Soc. London, for 1837, p. 87, 13 Feb. 

1838. Island of Fernando Po. 
Galago allenii, var. gabonensis Gray, Proc. Zool. Soc. London, p. 146, Oct. 

1863. Gaboon. 
Otolicnus alleni var. cameronensis Peters, Monatsb. K. Preuss. Akad. Wiss., 

Beriin, p. 472, 1876. Aqua Town (Duala), Cameroons. 
Galago gabonensis balesi Elliot, Ann. Mag. Nat. Hist. (7) 20: 187, Sep. 1907. 

Como ("Kemo") River, 75 miles from mouth, Gaboon. 
Range: Forest region from the lower Niger and the Ubangi Rivers west to 

the Gulf of Guinea and Island of Fernando Po. 

Galago crassicaudatus crassicaudatus E. Geoffroy 

Galago crassicaudatus E. Geoffroy, Ann. Mus. d' Hist. Nat. Paris, 19: 166, 

1812. No locality given, but -fixed by Thomas (Ann. Mag. Nat. Hist. 

(8) 20: 48, July 1917), as Quelimane, Mozambique. 
Otogale crassicaudata var. kirkii Gray, Proc. Zool. Soc. London, for 1864, p. 456, 

May 1865. Quelimane, Mozambique. 
Otolemur badius Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 277, 1905. 

Ugalla (or Ualaba) River, Tanganyika Territory. 
Range: South slope of Uluguru Mts., Tanganyika Territory, south to the 

Zambesi River and west to the central Rift Valley. 



116 bulletin: museum of comparative zoology 

Galago crassicaudatus agisymbanus (Coquerel) 

Otolemur agisymbanus Coquerel, Revue et Mag. de Zool., (2) 11: 459, pi. 17-18, 
f. 1-1 b, 1859. Agisymbana Island, northern Zanzibar, East Africa. 
Range: Zanzibar and Pemba Islands. 

Galago crassicaudatus argentatus Lonnberg 

Galago argentatus Lonnberg, Ann. Mag. Nat. Hist. (8) 11: 167, Jan. 1913. 
Ukina, east of Shirati, east of Lake Victoria, Tanganyika Territory. 
Range: East shores of Lake Victoria, Ukerewe Island. 

Galago crassicaudatus garnettii (Ogilby) 

Otolicnus garnettii Ogilby, Proc. Zool. Soc. London, p. 6, July 1838. No 
locality, but regarded usually as "Natal", although "accepted" as Zanzi- 
bar by Thomas (Ann. Mag. Nat. Hist. (8) 20: 48, July 1917); but see 
Schwarz (ibid., (10) 5: 49-50, Jan. 1930). 

Galago zuluensis Elliot, Ann. Mag. Nat. Hist. (7) 20: 186, Sep. 1907. Zululand, 
Natal. 

Galago crassicaudatus kikuyuensis Lonnberg 

Galago (Otolemur) kikuyuensis Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 64, Jan. 
1912. Escarpment Station, Kikuyu, Naivasha Province, Kenya Colony. 

Galago crassicaudatus lasiotis Peters 

Galago lasiotis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 912, text-f. 

1, 1876. Mombasa, Kenya Colony. 
Galago hindei ElHot, Ann. Mag. Nat. Hist. (7) 20: 186, Sep. 1907. Kitui, Athi 

River, Kenya Colony. 
Galago hindsi Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 

Hist., 1: 62, 1912 =15 June 1913 (lapsus cal.). 
Range : Italian Somaliland and Kenya Colony, from lower Juba River, south 

to Tanga, Tanganyika Territory. 

Galago crassicaudatus lonnbergi Schwarz 

Galago crassicaudatus lonnbergi Schwarz, Ann. Mag. Nat. Hist. (10) 5: 48, 
Jan. 1930. Tambarara, Gorongoza Mts., Mozambique. 
Range: South of the Zambezi River to the Limpopo, exact limits unknown. 

Galago crassicaudatus monteiri (Gray) 

Callotus monteiri Gray (ex Bartlett MS.), Proc. Zool. Soc. London, p. 145, 

Oct. 1863. Cuvo Bay, south of Loanda, Angola. 
Range: South-central and southwestern Africa from the central African 

Rift Valley in the east, between Baraka and the Zambezi River. 

Galago crassicaudatus panganiensis (Matschie) 

Otolemur panganiensis Matschie, Sitzb. Ges. Xaturf. Freunde, Berlin, p. 278, 

Dec. 1905. Arusha, near Mt. Kilimanjaro, Tanganyika Territory. 
Range: Tanganyika Territory from Mt. Kilimanjaro and Mt. Meru south 

to the Uluguru Mts., and on Mafia Island. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 117 

Galago crassicaudatus umbrosus Thomas 

G[alago] c[rassicaudatus] umbrosus Thomas, Ann. Mag. Nat. Hist. (8) 20: 49, 

July 1917. Woodbush Mountain, Tzaneen Estate, Zoutpansberg district, 

northern Transvaal. 
Range : Only recorded from type locality. 

Galago demidovii demidovii G. Fischer 

Lemur minutus G. Cuvier, Tabl. Element. Hist. Nat. des Anim., p. 101, 1798. 

Senegal. "Unidentifiable. . . . Some authors have synonymized the 

species with Galago demidoffi, G. Fischer, described eight years later" 

(Allen, J. A., BuU. Amer. Mus. Nat. Hist., 47: 302, 6 Feb. 1925). 
Galago cuvieri G. Fischer, Mem. Soc. Imp. des Naturalistes, Moscou, 1: 23, 

1806. New name for Lemur minutus G. Cuvier, which is unidentifiable. 
Galago demidovii G. Fischer, Comment. Soc. Phys. Med. Moscou, 1 : 57, 1808, 

apud Sherborn. Senegal. 
Galagoides demidoffi, A. Smith, South African Quart. Journ., 2: no. 1, pt. 2, 

p. 32, 1833 (fide Sherborn). 
Mioxicebus rufus Lesson, Species Mamm. Bimanes et Quadrumanes, p. 219, 

1840. "Madagascar," but said to be a synonym of Galago demidovii G. 

Fischer (see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 303, 6 Feb. 

1925). 
Octolicnus (sic) peli Temminck, Esquisses Zool. sur la Cote de Guine, p. 42, 

1853. Dabacrom, Gold Coast. 
Range: Forest region from Senegal to Liberia, Gold Coast and Niger River. 

Galago demidovii anomurus Pousargues 

Galago (Hemigalago) anomurus Pousargues, Bull. Soc. Zool. de France, 18: 51, 

1893; Nouv. Arch. Mus. d'Hist. Nat. Paris, (3) 6: 157, pi. 2 (col.), 1894. 

Upper Kemo River, right affluent of the Ubangi, French Congo. 
Hemigalago demidoffi, medius Thomas, Ann. Mag. Nat. Hist. (8) 16: 466, 

Dec. 1915. Poko, upper Uele district, Congo Beige. 
Range : Forests of right bank of the Congo, from the upper Shari and east of 

the Ubangi Rivers, south to Panga on the middle Aruwimi. 

Galago demidovii orinus Lawrence and Washburn 

Galago demidovii orinus Lawrence and Washburn, Occasional Papers Boston 

Soc. Nat. Hist., 8: 259, 8 Jan. 1936. Bagilo, Uluguru Mts., Tanganyika 

Territory, 5000 feet. 

Galago demidovii phasma Cabrera and Ruxton 

Galagoides demidoffi phasma Cabrera and Ruxton, Ann. Mag. Nat. Hist. (9) 

17: 596, May 1926. St. Joseph de Luluabourg, southern Congo Beige. 
Range: Forest region on left bank of the Congo, from near its mouth east to 

Tanganyika-Morro district. 



118 bulletin: museum of comparative zoology 

Galago demidovii poensis Thomas 

Galago demidoffi poensis Thomas, Abstract Proc. Zool. Soc. London, no. 10, 

p. 12, 22 Nov. 1904; Proc. Zool. Soc. London, for 1904, 2: 186, 18 Apl. 

1905. Bantaberi, Island of Fernando Po, 1800 m. 
Range: Island of Fernando Po. 

Galago demidovii pusillus (Peters) 

Otolicnus pusillus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 473, 
1876. Donguila, Gaboon, French Equatorial Africa. 
According to Schwarz, E., (Ann. Mag. Nat. Hist. (10) 7: 62, Jan. 1931), the 
type is from Old Gaboon Station (Libreville), Gaboon. 

Galago demidovii thomasi Elliot 

Galago (Hemigalago) ihomasi Elliot, Ann. Mag. Nat. Hist. (7) 20: 189, Sep. 

1907. Fort Benin, Semliki River, border of Uganda and Congo Beige. 
Galago matschiei Lorenz, Ann. K. K. Naturhist. Hofmus., Wien, 31: 237, 1917. 

Moera, Ituri forest, Congo Beige. 
Range: Central Ituri forest region east to Lake Victoria and the Victoria 

Nile. 

Galago senegalensis senegalensis E. Geoff roy 

Galago senegalensis E. Geoffroy, Mag. Encycloped., 1: 38-41, pi. 1, 1796. 
Senegal. 

Lemur galago G. Cuvier, Tabl. Element. Hist. Nat. des Anim., p. 101, 1798. 
Senegal. 

Lemur calago Shaw, Gen. Zool., Mamm., 1 : 108, 1800. Senegal. 

Galago geoffroyi G. Fischer, Mem. Soc. Imp. des Nat., Moscou, 1 : 25, 1806. 
Substitute name for G. senegalensis E. Geoffroy. 

Galago acaciarum Lesson, Species Mamm. Bimanes et Quadrumanes, p. 246, 
1840. New name for G. senegalensis E. Geoffroy. 

Otolicnus teng Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 
1842, p. 201, 1843. Bahr-el-Abiad, Anglo-Egyptian Sudan. 

Galago murinus Murray, Edinburgh Philosophical Journ., 10: 243, 1859. 

Galago sennariensis Gray, Proc. Zool. Soc. London, p. 147, text-f., Oct. 1863. 
White Nile, Sennaar, Anglo-Egyptian Sudan. According to Schwarz (Ann. 
Mag. Nat. Hist. (10) 7: 53, footnote, Jan. 1931) this is the first publication 
of the name in a technical sense; Otolicnus galago, a. sennariensis Wagner 
(Schreber's Saugthiere, Suppl., 5: 158, 1855) is only a geographical desig- 
nation. 

Galago pupulus EUiot, Ann. Mag. Nat. Hist. (8) 5: 77, Jan. 1910. Yola, 
Nigeria. 
Range: French and Anglo-Egyptian Sudan in the savanna country, from 
Senegal and Gambia east to Abyssinia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 119 

Galago senegalensis albipes Dollman 

Galago braccatus albipes Dollman, Ann. Mag. Nat. Hist. (8) 4: 549, Dec. 1909. 
Kirui, Mt. Elgon, Kenya Colony. 
Range: Uplands of Kenya Colony west of the Rift Valley. 

Galago senegalensis braccatus Elliot 

Galago braccatus Elliot, Ann. Mag. Nat. Hist. (7) 20: 187, Sep. 1907. (Tsavo 

River north of) Mount Kilimanjaro, Kenya Colony. 
Range: Uplands of Kenya and Tanganyika, west to the Great Rift Valley, 

south to Pangani River. 



Galago senegalensis bradfieldi Roberts 

Galago moholi bradfieldi Roberts, Ann. Transvaal Mus., 14: 224, 28 July 1931. 
No locality given, but according to Shortridge (Mamm. South West 
Africa, 1:11, 1934) the type was from Waterberg, Southwest Africa. 



Galago senegalensis dunni Dollman 

Galago dunni Dollman, Ann. Mag. Nat. Hist. (8) 5: 92, Jan. 1910. Headwaters 
of Fafan, 35 miles east of Harar, SomaUland. 
Range : Abyssinian plateau. 



Galago senegalensis gallarum Thomas 

Galago gallarum Thomas, Ann. Mag. Nat. Hist. (7) 8: 27, July 1901. Webi Dau, 

Boran Galla country, southern Abyssinia. 
Range: Somaliland and Galla country south of the Abyssinian plateau and 

Chercher Range, south to Northern Guaso Nyiro and Tana River, 

Kenya Colony. 



Galago senegalensis granti Thomas and Wroughton 

Galago granti Thomas and Wroughton, Proc. Zool. Soc. London, p. 286, 1 Aug. 

1907. Coguno, Inhambane, Mozambique. 
Galago mertensi Frade, Bull. Soc. Portuguaise Sci. Nat., Lisbonne, 9: 128, 1924. 

Chibuto =Shibuto, Inhambane, Prov. Louren^o Marques, Mozambique. 
Range: Southern part of Mozambique. 



Galago senegalensis intontoi Monard 

Galago moholi var. intontoi Monard, Bull. Soc. Neuchateloise Sci. Nat., 
Neuchatel, 55: 67, 1931. Type locahty not designated, specimens re- 
corded from Mbale River, Caquindo, Chimporo, and Vila da Ponte, 
Angola. 



120 bulletin: museum of comparative zoology 

Galago senegalensis inustus Schwarz 

Galago senegalensis inustus Schwarz, Revue Zool. et Bot. Africaines, 19: 391, 
1 Oct. 1930. Djugu, west of Lake Albert, south of Mahagi, Ituri district, 
Congo Beige. According to Hayman (Ann. Mag. Nat. Hist. (10) 20: 150, 
July 1937) this should stand as a full species. 
Range: Semliki-Ituri divide. 

Galago senegalensis moholi A. Smith 

Galago moholi A. Smith, Report Exped. for Exploring Central Africa, from Cape 

of Good Hope, June 23, 1834, under Dr. A. Smith, p. 42, June 1836. Banks 

of the Marikwa and Limpopo, Bechuanaland. 
Galago conspicillatus I. Geoffroy, Cat. Methodique Mamm. Mus. d'Hist. Nat., 

Paris, Primates, pt. 1, p. 81, 1851. 
Ololicnus galago var. australis Wagner, Schreber's Saugthiere, Suppl., 5: 158, 

1855. New name for Galago moholi A. Smith; according to Schwarz (Ann. 

Mag. Nat. Hist. (10) 7: 53, footnote, Jan. 1931) this is not a technical use 

of the name but adjectival merely. 
0[tolicniis\ mossambicus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 

p. 473, footnote, 1876. Tette, south bank of Zambesi River, Mozambique; 

based on the 0. senegalensis of the "Reise." 
Galago nyasx EUiot, Ann. Mag. Nat. Hist. (7) 20: 188, Sep. 1907. Mountains 

south of Lake Nyasa, Mozambique. 
Galago nyassse Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 

Hist., 1: 70, 1912 =15 June 1913 (lapsus cal.). 
Galago tumbolensis Monard, Bull. Soc. Neuchateloise Sci. Nat., Neuchatel, 55: 

68, 1931. Tumbole River, branch of the Kutato, about 40 km. east of 

Vila da Ponte, Angola. Probably a synonym of moholi. 
Range: Savanna country of South Africa, from Tanganyika Territory 

(Tabora) on the east to Angola in the west. 

Galago senegalensis sotik^ Hollister 

Galago sotikie Hollister, Smithsonian Misc. Coll., 72: no. 2, p. 1, 22 Jan. 1920; 

Bull. U. S. Nat. Mus., no. 99, pt. 3, p. 16, pi. 2 (skull), 1924. Telek River, 

Sotik, Kenya Colony. 

Galago senegalensis zanzibaricus Matschie 

Galago zanzibaricus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. Ill, 1893. 

Yambiani, western Zanzibar (not Muyuni as given in original description, 

see Schwarz, E., Ann. Mag. Nat. Hist. (10) 7: 55, Jan. 1931). 
Galago moholi cocos Heller, Smithsonian Misc. Coll., 60: no. 12, p. 1, 4 Nov. 

1912. Mazeras, Kenya Colony. 
Range: Zanzibar Island and adjacent mainland coast from northeast of 

Mombasa (Mazeras) to at least Dar-es-Salaam (Tanganyika Territory). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 121 

EuoTicus Gray 

Euoticus Gray, Proc. Zool. Soc. London, p. 140, text-f., Oct. 1863. As a sub- 
genus of Otogale Gray ( =Galago E. Geoffrey); type Otogale pallida Gray. 

Euoticus elegantulus elegantulus (Leconte) 

Microcebus elegantulus Leconte, Proc. Acad. Nat. Sci. Philadelphia, p. 10, 1857. 

"Western Africa" (Du Chaillu Collection); type locality fixed as Njola, 

Ogowe River, Gaboon (see Schwarz, E., Ann. Mag. Nat. Hist. (10) 7: 65, 

Jan. 1931). 
Ololicnus apicalis Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 361, Nov. 1860. 

Gaboon, "mountains of the interior near the equator," i.e., north of Njola, 

Ogowe River. 
Galago elegantulus tonsor Dollman, Ann. Mag. Nat. Hist. (8) 5: 94, Jan. 1910. 

Spanish Guinea, 15 miles from mouth of Benito River. 
Range: Forest region between the Sanaga, Ubangi and Congo Rivers on the 

east to the Cameroons, Muni, and Gaboon. 

Euoticus elegantulus pallidus (Gray) 

Otogale pallida Gray, Proc. Zool. Soc. London, p. 140, pi. 19, text-f., Oct. 1863. 
Island of Fernando Po. 

Galago talboti Dollman, Ann. Mag. Nat. Hist. (8) 5: 93, Jan. 1910. Nkami, 
southern Nigeria. 
Range: Island of Fernando Po and adjacent mainland of Nigeria and Came- 
roons from the lower Niger probably east to the Sanaga River. 



Family LEMURID^E. Typical Lemurs 

Review and key to genera: Schwarz, E. A revision of the genera and species 
of Madagascan Lemuridse. Proc. Zool. Soc. London, p. 399-428, 29 June 1931. 

Subfamily CHEIROGALEIN^. Mouse Lemurs 
Cheirogaleus E. Geoffroy 

Cheirogaleus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 172, 1812. Geno- 
type Cheirogaleus major E. Geoffroy (fixed by Elliot, D. G., Publ. Field 
Columbian Mus., Zool., 8: 548, 1907). 

Chirogaleus Oken, Lehrbuch d. Naturgesch., pt. 3, zool., sect. 2, p. xi, 1168, 
1816; emendation. Genotype Cheirogaleus major E. Geoffroy. 

Cebugale Lesson, Spec. Mamm., pp. 207, 219, 1840. Genotype, by monotypy, 
Cebugale commersonii Lesson = Cheirogaleus major E. Geoffroy. 

Mioxicebus Lesson, Spec. Mamm., pp. 207, 219, 1840. Genotype, by subse- 
quent designation, Mioxicebus griseus J^esson = Cheirogaleus major E. 
Geoffroy. 



122 bulletin: museum of comparative zoology 

Chirogale Gloger, Gemein. Hand- u. Hilfsbuch d. Naturgesch., 1: 44, 1841; 
emendation. 

Myspithecus F. Cuvier, Hist. Nat. d. Mammiferes, Tabl. Gen. Method., p. 2, 
1842; not of Blainville, 1839. Genotype by monotypy Le maki nain = 
Cheirogaleus major E. Geoffroy. 

Myoxicebus L. Agassiz, Nomenclator Zool., Index Univ., p. 243, 1845; emenda- 
tion of Mioxicebus. 

Myoxocebus L. Agassiz, Nomenclator Zool., Index Univ., p. 235, 1846; emenda- 
tion of Mioxicebus Lesson. 

Opolemur Gray, Proc. Zool. Soc. London, for 1872, p. 853, Apl. 1873. Genotype 
by monotypy Opolemur milii Gray (not E. Geoffroy) = Cheirogaleus medius 
samati (A. Grandidier). 

Altililemur Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 
Hist., 1: xlvii. 111, 1912 ( = 15 June 1913). Genotype Altililemur medius 
(E. Geoffroy) = Chirogaleus medius E. Geoffroy. 

Altilemur Weber, Die Saugetiere, ed. 2, 2: 736, 1928; lapsus for Altililemur. 

Cheirogaleus medius medius E. Geoffroy 

Cheirogaleus medius E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 172, pi. 10, 

f. 2, 1812. Ft. Dauphin, southeastern Madagascar (see Schwarz, E., 

Proc. Zool. Soc. London, p. 405, 29 June 1931). 
Opolemur thomasi Forsyth Major, Novitates Zool., 1: 20, pi. 1, f. 1; pi. 2, f. 2, 

11, 30 Jan. 1894. Ft. Dauphin, southeastern Madagascar. 

Cheirogaleus medius samati A. Grandidier 

Chirogalus samati A. Grandidier, Revue et Mag. de Zool., (2) 20: 49, Feb. 

1868. Tsidsibon River, western Madagascar. 
Cheirogaleus medius samati Schwarz, Proc. Zool. Soc. London, p. 405, 29 June 

1931. 

Cheirogaleus major major E. Geoffroy 

Chirogaleus major E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 172, pi. 10, 

f. 1, 1812. Ft. Dauphin, southeastern Madagascar. 
, Lemur commersonii Wolf, Abbild. u. Beschreib. merkwurd. naturgesch. 

Gegenstande, ed. 1, 2: 9, pi. 4, 1822. Based on Commerson's drawing 

made at Ft. Dauphin, southeastern Madagascar. 
Cheirogaleus milii E. Geoffroy, Cours de I'Hist. Nat. des Mammiferes, legon 11, 

p. 24, 1828. Madagascar. 
Myspithecus typus F. Cuvier, Hist. Nat. des Mammiferes, ed. 2, p. 228, pi. 

83. 1833. 
Cheirogaleus typicus A. Smith, South African Quart. Journ., 2: 50, 1833. 

Madagascar. 
Mioxicebus griseus Lesson, Spec. Mamm., p. 207, 219, 1840. Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 123 

Chirogalus adipicaudatus A. Grandidier, Ann. des Sci. Nat., Zool., (5) 10: 
378, 1868. Tullear, Madagascar. 
Range: Western and southeast coasts of Madagascar. 

Cheiroqaleus MAJOR CROSSLEYi A. Grandidier 

Chirogalus crossleyi A. Grandidier, Revue et Mag. de Zool., (2) 22: 49, Feb. 

1870. Forests east of Antsianak, Madagascar. 
Chirogale melanotis Forsyth Major, Novitates Zool., 1 : 25, pi. 2, f. 10, 30 Jan. 

1894. Vohemar or Vohima, northeast coast of Madagascar. 
Chirogale sibreei Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 325, Oct. 1896. 

Ankeramadinika, east of Antananarivo, Madagascar. 

Cheirogaleus trichotis Giinther 

Chirogaleus trichotis Giinther, Froc. Zool. Soc. London, p. 78, text-f. 1, 2, June 
1875. Between Tamatave and Morondava, Madagascar. 



MicROCEBUS E, Geoffroy 

Microcebus E. Geoffroy, Cours de I'Hist. Nat. des Mammiferes, le^on 11, p. 24, 

1828. Genotype by original designation Le Microcebe roux = Microcebus 

rufus (Wagner). 
According to Palmer (No. Amer. Fauna, no. 23, p. 420, 23 Jan. 1904) the 
work above mentioned was published in 1834, and the genotype is Lemur 
pusillus E. Geoffroy. 
Scartes Swainson, Nat. Hist, and Classif. Quadrupeds, p. 352, 1835. Genotype, 

by monotypy, Scartes murinus Swainson = Microcebus murinus J. F. 

Miller. 
Myscebus Lesson, Spec. Mamm., pp. 207, 214, 1840. Type by monotypy, 

Myscebus palmarum Ijesson = Microcebus murinus J. F. Miller. 
Gliscebus Lesson, Spec. Mamm., pp. 207, 216, 1840. Genotype Gliscebus 

murinus Lesson = Microcebus murinus J. F. Miller (as fixed by Schwarz, 

E., Froc. Zool. Soc. London, p. 401, 29 June 1931). 
Myocebus Wagner, Arch. f. Naturgesch., 7: sect. 2, p. 19, 1841. Error for 

Myscebus. 
Mirza Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., p. 131, 

1870. Genotype by monotypy Cheirogaleus coquereli A. Grandidier. 
Azema Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., p. 132, 

1870. Genotype by monotypy Cheirogaleus smithii Gray = Microcebus 

murinus rufus Wagner. 
Muriletnur Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., 

p. 132, 1870. Genotype by monotypy Microcebus murinus J. F. Miller. 



124 bulletin: museum of comparative zoology 

MiCROCEBUs MURiNTTS MURiNUS (J. F. Miller). Gray Mouse Lemur 

Lemur murinus J. F. Miller, Various Subjects of Nat. Hist., pi. 13, 1777; 
Cimelia Physica, pi. 13 and text p. 25, 1796. Madagascar but "obvi- 
ously the greyish southern and western" race. 

Prosimia minima Boddsert, Elenchus Animalium, p. 66, 1785. Based on 
Brown's "Little Maucauco," in New Illustr. Zool., p. 107, pi. 44, 1776 
(see Schwarz, E., Proc. Zool. Soc. London, p. 402, 29 June 1931). Mada- 
gascar. 

Lemur prehensilis Kerr, Linnseus's Animal Kingdom, p. 88, 1792. Based on 
same sources as Prosimia minima Boddsert. 

Lemur pusillus E. Geoffroy, Bull. Soc. Philomathique, Paris, 1: 90, 1795. 
Audebert's pi. 8 (Hist. Nat. d. Singes et Makis, 1800) "may be regarded 
as representing the type" (Schwarz, E., Proc. Zool. Soc. London, p. 402, 
29 June 1931). 

Galago madagascariensis E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 166, 
1812. Madagascar. 

Cheirogaleiis yninor E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 172, pi. 10, 
f. 3, 1812. Probably Fort Dauphin, southeastern Madagascar. 

Myocebus palmarum Lesson, Spec. Mamm., p. 214, 1840. 

Microcebus myoxinus Peters, Raise nach Mossambique, Zool., 1: Saugeth., 
p. 14, pis. 3, 4, f. 6-9, 1852. Bombatoka, St. Augustine's Bay, southwestern 
Madagascar. 

Chirogalus gliroides A. Grandidier, Ann. des Sci. Nat., Zool., (5) 10:378, 1868. 
Tulear, Madagascar. 

M[icrocebus] minor griseorufus Kollman, Bull. Mus. d'Hist. Nat. Paris, 16: 304, 
24 Nov. 1910. Southeast, south and southwest coasts of Madagascar. 
Range: Southeastern, eastern and western Madagascar as far north as the 
Betsiboka River. 

Microcebus murinus rufus Wagner 

Microcebus rufus Wagner, Schreber's Saugthiere, Suppl., 1 : 291, footnote, 1840; 
based on the Galago madagascariensis of Geoffroy (Ann. Mus. d'Hist. Nat., 
Paris, 19: 166, 1812) and of Desmarest (Encyclop. Methodique, Mam- 
malogie, 1: 103, 1820). Probably eastern Madagascar. 

Chirogaleus smithii Gray, Ann. Mag. Nat. Hist. (1) 10: 257, Dec. 1842. Mada- 
gascar; for F. A. Smith. 
Range: Eastern, east-central and northern Madagascar as far south as Ft. 
Dauphin in the east, and Bay of Bombetoka in the northwest. 

Microcebus coquereli (A. Grandidier) 

Cheirogalus coquereli A. Grandidier, Revue et Mag. de Zool., (2) 19: 85, Mar. 
1867. Morondova, southwestern Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 125 

Phaner Gray 

Phaner Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., p. 132, 

1870. Genotype by monotypy Lemur furcifer Blainville. 
Microcebus, Cheirogaleus, Lepilemur, of older authors, in part. 

Phaner furcifer (Blainville) 

Lemur furcifer Blainville, Osteographie, Mammiferes, Primates, Lemur, p. 35, 
pi. 7, f. 2, 1839. "Probably the region of the Bay of Antongil, N. E. Mada- 
gascar" (Schwarz, E., Proc. Zool. Soc. London, p. 407, 29 June 1931). 
Range : Northern and northwestern Madagascar. 



Subfamily LEMURIN^E 
Hapalemur I. Geoffroy 

Hapalemur I. Geofifroy, L' Institut, Paris, 19: 341, 22 Oct. 1851. Genotype by 

monotypy Lemur griseus E. Geoffroy. 
Hapalolemur Giebel, Die Saugethiere, p. 1018, 1855; emendation of Hapalemur 

I. Geoffroy. 
Prolemur Gray, Proc. Zool. Soc. London, for 1870, p. 828, pi. 52, text-f. 1-4, 

Apl. 1871. Type by monotypy Hapalemur si7nus Gray. 

Hapalemur simus Gray 

Hapalemur simus Gray, Proc. Zool. Soc. London, for 1870, p. 828, pi. 52, 
text-f. 1-4, Apl. 1871. Madagascar, without definite locality. 
Range : Forested region of eastern Madagascar. 

Hapalemur griseus griseus (Link) 

L[emur] griseus Link, Beytrage z. Naturgesch., 1: pt. 2, p. 65, 1795. Based on 

Le petit maki gris of Buffon, Hist. Nat., Suppl., 7: 121, pi. 34, 1789. 

Madagascar. 
Lemur cinereus Desmarest, Encyclop. Methodique, Mammalogie, 1: 101, 

1820, in error for griseus. Madagascar. 
Hapalemur schlegeli Pocock, Ann. Mag. Nat. Hist. (8) 19: 348, Mar. 1917. 

Madagascar, without definite locality, but "probably . . . the northwest 

coast." 
Range : Southern and western coasts and dry central plateau of Madagascar. 

Hapalemur griseus olivaceus I. Geoffroy 

Hapalemur olivaceus I. Geoffroy, Cat. Methodique Mamm. Mus. d'Hist. 

Nat. Paris, Primates, p. 75, 1851. Madagascar, without exact locality. 
Range: Moist wooded parts of eastern Madagascar probably as far south as 

Ft. Dauphin (Schwarz, E., Proc. Zool. Soc. London, p. 408, 29 June 1931). 



126 bulletin: museum of comparative zoology 

Lemur Linnseus 

Lemur Linnseus, Syst. Nat., ed. 10, 1: 29, 1758. Genotype Lemur catta Lin- 
nseus (see Thomas, O., Proc. Zool. Soc. London, p. 129, Mar. 1911). 

Prosimia Brisson, Regnum Anim. in Classes IX distrib., ed. 2, p. 13, 156, 1762. 
Genotype by subsequent designation. Lemur catta Linnseus (see Elliot, 
D. G., Review of the Primates, 1 : xxviii, 1912 =15 June 1913). 

Procebus Storr, Prodrom. Method. Mamm., p. 32, tabl. spec. A, 1780. Geno- 
type Lem.ur catta Linnaeus. 

Catta Link, Beschreib. Naturalien-Sammlung Univ. Rostock, 1: 7, 1806. 
Genotype by tautonymy. Lemur catta Linnseus. 

Maki Muirhead, Brewster's Edinburgh Encyclop., 13: 405, 1819. Genotype 
Maki mococo Muirhead =Lemur catta Linnseus (see Schwarz, E., Proc. 
Zool. Soc. London, p. 409, 29 June 1931). 

Varecia Gray, Proc. Zool. Soc. London, p. 135, text-f., Oct. 1863. Type by 
subsequent designation Lemur variegatus Kerr (see Elliot, D. G., Review 
of the Primates, Monographs Amer. Mus. Nat. Hist., 1: xxx, 1912 =15 
June 1913). 

Mococo Trouessart, Revue et Mag. de Zool., (3) 6: 163, 1878; quoted from 
Lesson. Genotype by monotypy Prosimia catta =Lemur catta Linnseus. 

Eulemur Haeckel, Syst. Phylogenie d. Wirbelthiere, 3: 600, 1895. Nomen 
nudum; no species mentioned. 

Review and key to species: Schwarz, E., Proc. Zool. Soc. London, p. 409-419, 
29 June 1931. 

Lemur catta Linnseus. Ring-tailed Lemur 

Lemur catta Linnteus, Syst. Nat., ed. 10, 1: 30, 1758. Madagascar. 

Maki mococo Muirhead, Brewster's Edinburgh Encycloped., 13: 405, 1819. 

No locahty. 
Range: Rocky open country from region of Ft. Dauphin in southeastern 

Madagascar to southern and southwestern parts near Bay of St. Augustin. 

Lemur macaco macaco Linnseus. Black Lemur 

Lemur macaco Linnseus, Syst. Nat., ed. 12, 1: 44, 1766. Madagascar, without 

exact locality. 
Lemur macaco niger Schreber, Saugthiere, 1 : (142), pi. 40 A, 1775. Madagascar. 
Lemur leucomystax Bartlett, Proc. Zool. Soc. London, for 1862, p. 347, pi. 41, 

Apl. 1863. Madagascar, without definite locality. 
Range: Forests of northwestern coasts of Madagascar and the coastal 

islands, north of Bay of Bombetoka. 

Lemur macaco albifrons E. Geoffroy 

Lemur albifrons E. Geoffroy, Mag. Encyclopedique, 1: 20, 1796. Madagascar, 
without exact locality. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 127 

Prosimia frederici Lesson, Spec. Mamm., p. 230, 1840. Madagascar, without 
exact locality. 

Lemur macaco collaris E. Geoffroy 

Lemur collaris E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 161, 181?. Mada- 
gascar, without exact locality. 

Prosimia melanocephala Gray, Proc. Zool. Soc. London, p. 137, pi. 18, Oct. 
1863. Madagascar. 

Prosimia xanthomystax Gray, Proc. Zool. Soc. London, p. 138, pi. 17, Oct. 1863. 
Madagascar, without exact locality. 

Lemur mongoz var. cinereiceps A. Grandidier and MUne-Edwards, Hist. Nat. 
Madagascar, Mamm., 5: atlas, 2: pi. 140; pi. 147, f. 2, 1890. Madagascar. 
Range: Coastal forests of southeastern Madagascar from Ft. Dauphin in the 
south, to "at least Masindrano in the north." 

Lemur macaco flavifrons (Gray) 

Prosimia flavifrons Gray, Proc. Zool. Soc. London, p. 596, pi. 31, Oct. 1867. 

Madagascar, without exact locality. Probably a synonym of Lemur macaco 

collaris. 
Lemur nigerrimus P. L. Sclater, Proc. Zool. Soc. London, p. 451, text-f. 2, Oct. 

1880. Madagascar, without exact locality. This is stated by Schwarz 

(Mammalia, 1:1,1 Sep. 1936) to be the female of L. m. flavifrons. 
Range: Maromandia, northwestern Madagascar. 

Lemur macaco fulvus E. Geoffroy 

Lemur fulvus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 161, 1812. Mada- 
gascar, without exact locality. As to subspecific status, see Schwarz, E. 
(Mammalia, 1: 1, 1 Sep. 1936). 

Prosimia macromongoz Lesson, Spec. Mamm., p. 226, 1840. No exact locality. 

Lemur bruneus van der Hoeven, Tijdschr. f. Natur. Geschied. Physiol., 11: 35, 
1844. New name for Lemur fulvus E. Geoffroy. 

Lemur macaco mayottensis Schlegel 

Lemur mayottensis Schlegel, Nederland. Tijdschr. Dierkunde, no. 3: 76, 1866. 
Island of Mayotte, Mascarene group. "There is little doubt about its 
being entitled to subspecific rank" (Schwarz, E., Proc. Zool. Soc. London, 
p. 414, 29 June 1931). 

Lemur macaco rufus Audebert 

Lemur rufus Audebert, Hist. Nat. des Singes et Makis, Makis, p. 12, pi. 2, 1799. 

Madagascar, without definite locality. 
Lemur rufifrons Bennett, Proc. Zool. Soc. London, p. 106, 18 Nov. 1833. 

Madagascar, without definite locality. 
Range: Central plateau of Madagascar and west coast. 



128 bulletin: museum of comparative zoology 

Lemur macaco sanfordi Archbold 

Lemur fulvus sanfordi Archbold, Amer. Mus. Novitates, no. 518, p. 1, 26 Mar. 
1932. Mt. D'Ambre, Madagascar. 

Lemur mongoz mongoz Linnseus 

Lemur mongoz Linnseus, Syst. Nat., ed. 12, 1: 44, 1766. Madagascar, based on 

Edwards's colored plate. Gleanings of Nat. Hist., 1 : 12, pi. 216, 1758 — in 

A Nat. Hist, of Birds. 
Lemur nigrifrons E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 160, 1812. 

Based on Petivier, Gazophylacii Naturae et Artis, dec. 1, p. 4, pi. 17, f. 5, 

1767. Madagascar. 
Lemur albimanus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 160, 1812. 

[Anjouan Id., Mascarene group.] 
[Lemur] dubius F. Cuvier, Hist. Nat. des Mamm., 1834. Anjouan Id., Masca- 
rene group. 
Prosimia micromongoz Lesson, Spec. Mamm. Bimanes et Quadrumanes, p. 226, 

1840. Based on E. Geoffroy's description of Lemur mongoz. 
Prosimia bugi Lesson, Spec. Mamm. Bimanes et Quadrumanes, p. 227, 1840. 

Based on Petivier, Gazophylacii Naturae et Artis, dec. 1, p. 4, pi. 17, f. 5, 

1767. 
Prosimia brissonii Lesson, Spec. Mamm. Bimanes et Quadrumanes, p. 230, 

1840. Based on Brisson's Prosimia pedibus albis (Regne Animale, ed. 2, 

p. 156, 1762). 
Prosimia ocularis Lesson, Spec. Mamm. Bimanes et Quadrumanes, p. 231, 1840. 

Madagascar, no locality. 
Lemur cuvieri Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 62: sect. 1, 

p. 646, 1870. Anjouan or Johanna Id., Comoro Ids., Mascarene group. 
Propithecus brissonianus Gray, Cat. Monkeys, Lemurs, and Fruit-eating Bats 

Brit. Mus., p. 75, 1870. Emendation of Prosimia brissonii Lesson. 
Range: Anjouan and Moheli Islands and on the mainland of Madagascar 

along the south bank of Betsiboka River. 

Lemur mongoz coronatus Gray 

Lemur coronatus Gray, Ann. Mag. Nat. Hist. (1) 10: 257, Dec. 1842. [Bay of 

Mahajamba, northwest coast of] Madagascar, 15° 14' S. 
Lemur chrysampyx Schuermans, Mem. Couronnes et des Savants Etrang. Acad. 

Roy. des Sci., Lettres et Beaux- Arts Belgique, Bruxelles, 4to, 22: 7th 

Mem. des Savants Etrang., p. 6, 1 pi., 1848. "Probablement Madagascar." 
Range: Northern Madagascar from Bombetoka Bay on the east to Bay of 

Antongil on the west. 

Lemur rubriventer I. Geoffroy 

Lemur rubriventer I. Geoffroy, Comptes Rend. Acad. Sci. Paris, 31: 876, 1850. 
Madagascar, without exact locality. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 129 

Lemur flaidvenler I. Geoffrey, Comptes Rend. Acad. Sci. Paris, 31: 876, 1850. 

Madagascar ("probably Tamatave," fide Schwarz). 
Lemur rufiventer Gray, Cat. Monkeys, Lemurs, and Fruit-eating Bats Brit. 

Mus., p. 74, 1870. Lapsus calami for Lemur rubriventer I. Geoffroy. 
Prosimia rufipes- Gray, Ann. Mag. Nat. Hist. (4) 7: 339, May 1871. Betsimi- 

saraka country, "probably west of Tamatave" (Schwarz), Madagascar. 
Range: Apparently "the greater part of Madagascar" (Schwarz, E., Proc. 

Zool. Soc. London, p. 417, 29 June 1931). 

Lemur variegatus Kerr. Ruffed Lemur 

Lemur macaco variegatus Kerr, Linnaius's Animal Kingdom, p. 86, 1792. Mada- 
gascar, without definite locality. 

Lemur ruber E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 159, 1812. Mada- 
gascar (based on Commerson's figure). 

M[aki] vari Muirhead, Edinburgh Encyclopedia, 13: 405, 1819. Based on Le 
Maki vari of Desmarest (Nouv. Dictionnaire d'Hist. Nat., ed. 2, 18: 437, 
1817). 

Prosimia erythromela Lesson, Spec. Mamm. Bimanes et Quadrumanes, p. 236, 
1840. 

Lemur varius I. Geoffroy, Cat. Methodique d. Mamm. Mus. d'Hist. Nat. 
Paris, Primates, p. 7, 1851. 
Range : Forests of northeastern Madagascar south to the region of Tamatave, 

about 18° S. 
As to the possibility that there are three local forms or races, see Schwarz, E., 

Proc. Zool. Soc. London, p. 418, 29 June 1931. 



Lepilemur I. Geoffroy 

Lepilemur I. Geoffroy, Cat. Methodique Mamm. Mus. d'Hist. Nat. Paris, Pri- 
mates, pt. 1, p. 75, 1851. Genotype Lepilemur mustelinus I. Geoffroy. 

Galeocebus Wagner, Schreber's Saugthiere, Suppl., 5: 147, 1855. Genotype 
Lepilemur mustelinus L Geoffroy. 

Lepidilemur Giebel, Saugethiere, p. 1018, 1859; emendation of Lepilemur. 

Mixocebus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 690, 1874. 
Genotype Mixocebus caniceps Peters = Lepilemur mustelinus L Geoffroy. 

Lepidolemur Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 690, footnote, 
1874; emendation of Lepilemur. 

Lepilemur mustelinus I. Geoffroy 

Lepilemur mustelinus I. Geoffroy, Cat. Methodique Mamm. Mus. d'Hist. Nat. 
Paris, p. 71, 1851. Madagascar, without exact locality, but fixed as "at 
or near Tamatave" (see Schwarz, E., Proc. Zool. Soc. London, p. 420, 29 
June 1931). 



130 bulletin: museum of comparative zoology 

Mixocebus caniceps Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 690, 

pi. 1 (col.), 2, 1874. Madagascar, probably on east coast between Tama- 

tave and Masindrano. 
Lepidolemur microdon Forsyth Major, in Forbes' Handbook to the Primates, 

1: 88, 1896. Eastern Betsileo Province, Madagascar. 
Range: Eastern and northeastern Madagascar from Vohemar south to 

Betsileo. 

Lepilemur rupicaudatus a. Grandidier 

Lepilemur ruficaudatus A. Grandidier, Revue et Mag. de Zool., (2) 19: 256, 

July 1867. [Morondava], Madagascar. 
Lepilemur dorsalis Gray, Cat. Monkeys, Lemurs, and Fruit-eating Bats Brit. 

Mus., p. 135, 1870. [Northwestern] Madagascar. 
Lepilemur pallidicauda Gray, Proc. Zool. Soc. London, for 1872, p. 850, Apl. 

1873. [Morondava, west coast of] Madagascar. 
Lepidolemur leucopus Forsyth Major, Ann. Mag. Nat. Hist. (6) 13: 211, Feb. 

1894. Fort Dauphin, southeastern Madagascar. 
Lepidolemur edwardsi Forsyth Major, in Forbes's Handbook to the Primates, 

1: 87, 1896. Betsako, Bay of Bombetoka, northwest coast of Madagascar. 
Lepidole?nur globiceps Forsyth Major, in Forbes's Handbook to the Primates, 

1: 89, 1896. Ambolisatra, 25 km. north of Tulear, southwest coast of 

Madagascar. 
Lepidolemur grandidieri Forsyth Major, in Forbes's Handbook to the Primates, 

1: 89, 1896. Northwestern Madagascar. 
L[epidolemur] mustelinus rufescens Lorenz, Abhandl. Senckenberg. Naturf. 

Ges., Frankfurt a. M., 21: 446, pi. 30; 31, f. 4, a, b, 1898. Ambundub6, 

north side of Bay of Bombetoka, northwestern Madagascar. 
Range: Entire west coast of Madagascar, but not the central plateau. 

Family INDRIID^ 
AvAHi Jourdan. WooUv Lemur 

Avahi Jourdan, L'Institut, Paris, 2: 231, 19 July 1834. Genotype Lemur 
laniger Gmelin, though not here given. 

Microrhynchus Jourdan, These Fac. Sci. Grenoble, 1834; not of Megerle, 1823, 
for Coleoptera. 

Avahis I. Geoffroy in Gervais, Resume Legons Mamm., p. 23, 1835; emenda- 
tion of Avahi Jourdan. 

Habrocebus Wagner, Schreber's Saugthiere, Suppl., 1: v, 1839. Genotype by 
subsequent designation Lemur lanatus Schreher = Lemur laniger Gmelin. 

Semnocebus Lesson, Species Mamm. Bimanes et Quadrumanes, p. 209, 1840. 
Genotype Semnocebus avahi Lesson = Lemur laniger Gmelin. 

Iropocus Gloger, Gemein. Hand.- u. Hilfsbuch Naturgesch., l:xxviii, 43, 1841. 
Genotype Lemur laniger Gmelin. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 131 

AvAHi LANiGER LANiGER (Gmelin). Woolly Lemur 

Lemur laniger Gmelin, Linne's Syst. Nat., ed. 13, 1: 44, 1788. Madagascar. 

Lemur brunneus Link, Beytrage z. Naturgesch., pt. 2, p. 165, 1795. Based on 

Sonnerat. 
Indris longicaudatus E. Geoffrey, Ann. Mus. d'Hist. Nat. Paris, 19: 158, 1812. 

Madagascar; based on Sonnerat. 
H[abrocebus] lanatus Wagner, Schreber's Saugth., Suppl., 1: 258, pi. 42A, 

1840. Madagascar; based on Sonnerat. 
Semnocebus avahi Lesson, Species Mamm. Bimanes et Quadrumanes, p. 210, 

1840; based on Sonnerat. 
A[vahis] laniger orientalis Lorenz, Abhandl. Senckenberg. Naturf. Ges., Frank- 
furt a. M., 21: 452, 1898. East coast of Madagascar. 
The eastern or typical race of A. Grandidier and Milne-Edwards, Hist. Nat. 
de Madagascar, Zool., Mamm., 1: 327, 1875. 

Range: Forest region of northeastern, eastern and south-central Madagascar. 

Avahi laniger occidentalis (Lorenz) 

Avahis laniger occidentalis Lorenz, Abhandl. Senckenberg. Naturf. Gea., 
Frankfurt a. M., 21: 452, 1898. Based on the Avahis laniger vari^te 
septentrionale of A. Grandidier and Milne-Edwards. Northwestern 
Madagascar, Ambundube. 
Range: Northwest coast of Madagascar south to the Bay of Bombetoka. 



Indri E. Geoffroy and G. Cuvier. The Indri 

Indri E. Geoffroy and G. Cuvier, Mag. Encyclop., 1 : 46, 1796, fide Palmer. 

Genotype by tautonymy Lemur indri Gmelin. 
Lichanotus IHiger, Prodromus Syst. Mamm. et Avium, p. 72, 1811. Genotype 

Lemur indri Gmelin. 
Indris G. Cuvier, Lemons d'Anat. Comp., 1: Tabl. Gen., 1805; emendation of 

Indri. 
Indrium Rafinesque, Analyse de la Nature, p. 54, 1815. New name for Indri 

E. Geoffroy and G. Cuvier. 
Lichanotes Temminck, Monographies de Mammalogie, 1 : xvi, 1827. Genotype 

Lemur indri Gmelin; lapsus for Lichanotus. 
Pithelemur Lesson, Species Mamm. Bimanes et Quadrumanes, pp. 207, 208, 

1840. Genotype Lemur indri Gmelin. 

Indri indri (Gmelin). Indri 

Lemur indri Gmehn, Linne's Syst. Nat., ed. 13, 1: 42, 1788. Madagascar, 

without exact locality. 
Indri brevicaudatus E. Geoffroy and G. Cuvier, Mag. Encyclop., p. 46, 1796. 

Based on Sonnerat's specimen (Voy. Ind. Or. Chine, 2: 142, pi. 88, 1782). 



132 bulletin: museum of comparative zoology 

Indris ater I. Geoffroy, Dictionnaire Class. d'Hist. Nat., 8: 534, 1825. Lapsus 

cal. for niger Lacepede. 
Indris variegatus Gray, Ann. Mag. Nat. Hist. (4) 10: 474, Dec. 1872. Probably 

Lalo River, east of Bay of Antongil, Madagascar (Schwarz). 
Range: Northeastern Madagascar to Lake Alaotra on the central plateau. 



Propithecus Bennett. The Sifakas 

Propithecus Bennett, Proc, Comm. Sci. and Correspondence Zool. Soc. London, 
p. 20, 29 Mar. 1832. Genotype by monotypy Propithecus diadema Bennett. 

Macromerus A. Smith, South African Quart. Journ., 2: pt. 2, p. 49, 1833. 
Genotype by monotypy Macromerus typicus A. Smith =Propithecus 
diadema Bennett. 

Propithecus diadema diadema Bennett 

Propithecus diadema Bennett, Proc, Comm. Sci. and Correspondence Zool. 

Soc. London, p. 20, 1832. Madagascar, without exact locality. 
Macromerus typicus A. Smith, South African Quart. Journ., 2: pt. 2, p. 49, 

1833. 
Indris albus Vinson, Comptes Rend. Acad. Sci. Paris, 55: 829, 1862. Alana- 

masoatrao, between Andevoranto and Tananarive, eastern Madagascar. 
Range: Mountain forests of the coast range of eastern Madagascar from 
Bay of Antongil south to Masora River. 

Propithecus diadema candidus A. Grandidier 

Propithecus candidus A. Grandidier, Comptes Rend. Acad. Sci. Paris, 72: 231, 

1871. Forests north of Bay of AntongU, (but according to Schwarz) 

Sahambavany, northeast coast of Madagascar. 
Propithecus sericeus Milne-Edwards and A. Grandidier, Revue et Mag. de 

Zool. (2) 23: 274, Aug. 1872. Sambava, northeast coast of Madagascar. 

Renaming of P. diadema candidus A. Grandidier. 
Range: Northeastern coast ranges of Madagascar, from Bemarivo (14° 16' 

30" S.) southward to Bay of Antongil. 

Propithecus diadema edwardsi A. Grandidier 

Propithecus edwardsi A. Grandidier, Comptes Rend. Acad. Sci. Paris, 72: 232, 
1871. Forests west of Mananzary, southeastern Madagascar. 

Propithecus bicolor Gray, Ann. Mag. Nat. Hist. (4) 10: 206, Sep. 1872. Prob- 
ably Masindrano, southeastern Madagascar (see Schwarz, E., Proc. 
Zool. Soc. London, p. 422, 29 June 1931). 
Range: Coast range of southeast Madagascar, between the Masora and 
Matitana Rivers. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 133 

Propithecxjs diadema holomelas Gimther 

Propithecus holomelas Giinther, Ann. Mag. Nat. Hist. (4) 16: 125, Aug. 1875. 
Fieneretora, probably Fianarantsoa, central Betsileo, Madagascar. 
Range: The inland mountain range in southeastern Madagascar. 

Propithecus verreatjxi verraeuxi A. Grandidier ! 

Propithecus verreauxi A. Grandidier, Revue et Mag. de Zool., (2) 19: 84, Mar. 

1867. Tsifanihy, Antandroy Province, southern Madagascar (Schwarz). 
Range: Southwestern Madagascar from Tsidsobon River southeast to the 

Ft. Dauphin region. 

Propithecus verreaxxxi coquereli (A. Grandidier) 

Cheirogalus coquereli A. Grandidier, Revue et Mag. de Zool., (2) 19: 85, Mar. 

1867. Morondova, west coast of Madagascar. 
Propithecus damoni Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. 

Mus., app., p. 137, 1870 (quoting Pollen). Madagascar. 
Propithecus perrieri Lavauden, Comptes Rend. Acad. Sci. Paris, 193: 77, 1931. 

Forest of Analamera, southeast of Diego Suarez, northern Madagascar. 
Probably a melanistic individual of P. v. coquereli. 
Range: Northwestern Madagascar from Antamba River south to River 

Betsiboka. 

Propithecus verreauxi coronatus Milne-Edwards 

P[ropithecus] coronatus Milne-Edwards, Revue Scientifique, (2) 1=8: 224, 

2 Sep. 1871. No locality; supposed to be Boeny Province, Bomb^toka 

Bay, Madagascar. 
Propithecus damanus Schlegel, ex Pollen, Mus. d'Hist. Nat. des Pays-Bas, 7: 

293, 1876. South shore of Bay of Bombetoka, Madagascar. 
Range: Between the Betsiboka and Mahavavy Rivers, northwestern 

Madagascar. 

Propithecus verreauxi deckenii Peters 

Propithecus deckenii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 421, 

1870. Kanatsy, northwestern Madagascar. 
Range: Northwestern Madagascar, between the Rivers Mahahavy and 

Mananbolo. 

Propithecus verreauxi majori W. Rothschild 

Propithecus majori W. Rothschild, Novitates Zool., 1 : 666, pi. 14, 20 Sep. 1894. 
Antinosy country, south-central Madagascar. 
Range: Known only from south-central Madagascar. 



134 bulletin: museum of comparative zoology 

Family DAUBENTONIID^ 
Daubentonia E. Geoffroy. Aye-aye 

Daubentonia E. Geoffroy, Decad. Philos. et Litt., p. 195, 1795. Genotype 

Sciurus madagascariensis Gmelin. 
Scolecophagus E. Geoffroy, Decad. Philos. et Litt., p. 196, 1795; alternative 

name for Daubentonia E. Geoffroy. Not Scolecophagus Swainson, 1831 

(Aves). 
Aye-aye Lacepede, Tabl. Mamm., p. 6, 1799. Genotype Sciurus madagas- 
cariensis Gmelin. 
Cheyromys E. Geoffroy, Cat. Mamm. Mus. d'Hist. Nat. Paris, p. 181, 1803; 

misprint for Cheiromys G. Cuvier. 
Chiromys Illiger, Prodromus Syst. Mamm. et Avium, p. 75, 1811; emendation 

of Cheiromys G. Cuvier. Genotype Sciurus madagascariensis Gmelin. 
Psilodactylus Oken, Lehrb. d. Naturgesch., pt. 3, zool., sect. 2, p. xi, 1164, 

1816. Genotype Lemur -psilodactylus Shaw =<Saurus madagascariensis 

GmeUn. 
Cheiromys G. Cuvier, Regne Animal, ed. 1, 1: 207, 1817. Genotype Sciurus 

madagascariensis Gmelin. 
Myspithecus Blainville, Osteograph. des Vertebres Recents et Foss., Mamm., 

1: pt. 3, r Aye-aye, p. 34, 1839. Genotype Sciurus madagascariensis 

Gmelin; alternative name for Cheiromys G. Cuvier. 
Myslemur Blainville, Dictionnaire Univ. d'Hist. Nat., 8: 559, 1846. New 

name for Myspithecus Blainville, not G. Cuvier. 

Daubentonlv madagascariensis (Gmelin). The Aye-aye 

Sciurus madagascariensis Gmelin, Linne's Syst. Nat., ed. 13, 1: 152, 1788. 

Western Madagascar, based on the Aye-aye of Sonnerat's Voyage. 
Lemur psilodactylus Shaw, Gen. Zool., Mamm., 1: 109, 1800. Madagascar. 
Cheiromys madagascariensis var. laniger G. Grandidier, Bull. Acad. Malgache, 

1: 106, pi., 1930. Madagascar. Probably based on a specimen of D. 

?nadagascariensis (Gmelin) in molting pelage. 
Chiromys robustus Lamberton, Mem. Acad. Malgache, 17: 40, pi. 10a, 1934. 

Tairave, 15 km. from Beroroha, south-central Madagascar. Subfossil(?); 

and probably same as Daubentonia madagascariensis. 
Range: Entire forested part of eastern and northwestern Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 135 

Suborder ANTHROPOIDEA. Monkeys, Apes 

Family CERCOPITHECID^ 

Allenopithecus Lang 

Allenopithecus Lang, Amer. Mus. Novitates, no. 87, p. 1, 12 Sep. 1923. Geno- 
type Cercopithecus nigroviridis Pocock. 

Allenopithecus nigroviridis (Pocock) 

Cercopithecus nigroviridis Pocock, Proc. Zool. Soc. London, p. 739, pi. 42, f. 5, 

8 0ct. 1907. "Upper Congo." 
Lasiopyga nigriviridis Elliot, Review of the Primates, Monographs Amer. 

Mus. Nat. Hist., 2: 348, 1912 ( = 15 June 1913). 
Allenopithecus nigroviridis Lang, Amer. Mus. Novitates, no. 87, p. 4, text-f. 

1-3 (skull and teeth), 12 Sep. 1923. 

Cercocebus E. Geoffroy. Mangabeys 

Cercocebus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 97, 1812. Genotype 
Cercocebus fuliginosus E. Geoffroy =Simia atys Audebert. 

Aethiops Martin, Gen. Introduct. Nat. Hist. Mammiferous Anim., p. 506, 1841. 
Genotype not specified; based on "the three White-eyelid Monkeys" of 
Africa, hence =Cercocebus. 

Semnocebus Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., 
p. 27, 1870; not of Lesson, 1840, for a genus of Lemurs. Genotype Presbylis 
albigena Gray. 

Lophocebus Palmer, Science, n.s., 17: 873, 29 May 1903. To replace Semnocebus 
Gray, not Lesson; type Presbytis albigena Gray. 

Cercolophocebus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 341, July 
1914. Subgenus of Cercocebus; type Cercocebus aterrimus Oudemans. 

Leptocebus Trouessart, Cat. Mamm. Viv. Foss., pt. 1, p. 15, 1904. As a sub- 
genus of Cercocebus to include C. galerilus, agilis, congicus and albigena; no 
type specified. 

Simia, Pithecus, Cercopithecus, Presbylis, in part, of older authors. 

Review: Schwarz, E. The species of the genus Cercocebus. Ann. Mag. Nat. 
Hist. (10) 1: 644-670, June 1928. 

Cercocebus albigena albigena (Gray) 

Presbytis albigena Gray, Proc. Zool. Soc. London, p. 77, Mamm., pi. 16, 12 Nov. 
1850. West Africa; "assumed to be the Lower Congo and Gaboon" 

(Schwarz). 



136 bulletin: museum of comparative zoology 

Cercocebus albigena johnstoni (Lydekker) 

Semnocebus albigena johnstoni Lydekker, Novitates Zool., 7: 594, 29 Dec. 1900. 

"Near Lake Tanganyika," but probably from the Semliki or Ituri Forest 

(fide Lorenz). 
Cercocebus jamrachi Pocock, Ann. Mag. Nat. Hist. (7) 18: 454, pi. 11, Dec. 

1906. Molinga, Lake Mweru, Congo Beige. 
Cercocebus (Leptocebus) albigena ituricus Matschie, Revue Zool. Africaine, 2: 

208, 15 Feb. 1913. Between Beni and Irumu, Semliki-Ituri Forest, Congo 

Beige. 
Cercocebus albigena ugandse Matschie, Revue Zool. Africaine, 2: 210, 15 Feb. 

1913. Chagwe, Uganda. 
C[ercocebus] albigena maioambicus Lorenz, Ann. Naturh. Hofmus. Wien, 31 : 

233, 1917. Mawambi, Ituri River, Congo Beige. 

Cercocebus albigena zenkeri Schwarz 

Cercocebus albigena zenkeri Schwarz, Sitzb. Ges. Naturf . Freunde, Berlin, p. 456, 

1910. Bipindi, Lokundje River, Cameroons. 
Cercocebus {Leptocebus) albigena weynsi Matschie, Revue Zool. Africaine, 2: 

211, 15 Feb. 1913. Sanga, French Congo. 

Cercocebus aterrimus (Oudemans). Black Mangabey 

Cercopithecus aterrimus Oudemans, Zool. Garten, 31 : 267, Sep. 1890. Stanley 

Falls, Congo Beige. 
Cercocebus congicus P. L. Sclater, Proc. Zool. Soc. London, for 1899, p. 827, 

text-f., 1 Apl. 1900. Stanley Falls district, Congo Beige. 
Semnocebus albigena rothschildi Lydekker, Novitates Zool., 7: 595, 29 Dec. 1900. 

No locality, but probably "the Congo." 
Cercocebus hamlyni Pocock, Ann. Mag. Nat. Hist. (7) 18: 208, pi. 7, Sep. 1906. 

"Upper Congo." 
Cercolophocebus ccelognathus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 342, July 1914. Kindu, Lualaba River, Congo Beige. 
Range: Whole of the forest area south of the Congo, but not north or east 

of it. 

Cercocebus galeritus galeritus Peters 

Cercocebus galeritus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 830, 
pi. IB, 3, 1879. Mitole, mouth of Tana and Osi Rivers, Kenya Colony. 

Cercocebus galeritus agilis Milne-Edwards 

Cercocebus agilis Milne-Edwards in Riviere, Revue Scientifique, (3) 12 =38: 

15, 3 July 1886. French Congo; "Poste des Ouaddes" (junction of Ubangi 

and Congo) (fide Schwarz). 
Cercocebus hagenbecki Lydekker, Novitates Zool., 7: 594, 29 Dec. 1900. Upper 

Congo River, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 137 

Cercocebus fumosus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 341, July 

1914. Northwest of Beni, Semliki, Congo Beige. 
Cercocebus oberlaenderi Lorenz, Anzeiger K. Akad. Wiss. Wien, math.-nat. KI., 

52: 172, 1915; Ann. K. K. Naturh. Hofmus., Wien, 31 : 230, pi. 15, f. 5, 6, 

1917. Mawambi, Ituri Forest, Congo Beige. 

Cercocebus galeritus chrysogaster Lydekker 

Cercocebus chrysogaster Lydekker, Novitates ZooL, 7: 279, pi. 3, 20 Aug. 1900. 
Upper Congo. 
Range: Southern and central Congo Basin (Schwarz). 

Cercocebus torquatus torquatus (Kerr) 

Simia (Cercopithecus) sethiops torquahis Kerr, Linnseus's Animal Kingdom, 
p. 67, 1792. Based on Buffon's (Hist. Nat., 14: 244, pi. 33, 1770) "Le 
Mangabey a collier blanc;" "I'Ethiopie," i.e.. West Africa. 

[Cercocebus] sethiopicus F. Cuvier, Hist. Nat. des Mamm., 2: pt. 35, pi. and text- 
leaf, Dec. 1821. West Africa, south of Cape Verde. 

Cercocebus collaris Gray, List Spec. Mamm. Brit. Mus., p. 7, 13 May 1843. 
"Africa." 

Cercocebus crossii Gray, ex Clark MS., List Spec. Mamm. Brit. Mus., p. 7, 13 
May 1843. "Africa." 
Range: Lower Niger to the Gaboon (Schwarz). 

Cercocebus torquatus atys (Audebert) 

Simia atys Audebert, Hist. Nat. des Singes et Makis, fam. 4, sect. 2, p. 13, pi. 8, 

1797. "Indes Orientales" but the type is an albino, labelled "Afrique occi- 

dentale" in Paris Mus. (Schwarz). 
Cercocebus fuliginosus E. Geoffroy, Ann Mus. d'.Hist. Nat. Paris, 19: 97, 1812. 

No locality, but based on "Le Mangabey" of Buffon (Hist. Nat., 14: 119, 

pi. 32, 1770) as shown by Schwarz. 
Range : French Guinea to Liberia. 

Cercocebus torquatus lunulatus (Temminck) 

Cercopithecus lunulatus Temminck, Esquisses Zool. sur la Cote de Guine, p. 37, 
1853. Rio Boutry, Gold Coast. 



Cercopithecus Linnaeus. Guenons 

Cercopitheci Linnaeus, Syst. Nat., ed. 10, 1: 26, 1758. As a subgroup of Simia, 
containing 17 species; no type designated but fixed by Stiles and Orleman 
(Journ. Mammalogy, 7: 52, 15 Feb. 1926) as Simia diana Linnseus. These 
authors show that although used only in the plural, the name is a subgenus 
in the modern sense, so that they "see no reason for rejecting the sub- 
generic names given under Simia." 



138 bulletin: museum of comparative zoology 

Cercopithecus Briinnich, Zoologiae Fundamenta, pp. 34, 40, 1772. Genotype 

Simia mona Schreber, fide Palmer, 1904. 
For a full discussion as to the use of this name instead of Lasiopyga for the 

African guenons, see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 373- 

379, 6 Feb. 1925. 
Lasiopyga Illiger, Prodromus Syst. Mamm. et Avium, p. 68, 1811. No type 

specified; included Simia nemsea Linnaeus and S. nictitans Linnaeus. 

Adopted by Elliot (Review of the Primates, Monographs Amer. Mus. 

Nat. Hist., 2: 275, 1912 = 15 June 1913) in place of Cercopithecus; type by 

elimination, Simia nictitans Linnaeus. 
Monichus Oken, Lehrbuch d. Naturgesch., pt. 3, zool., sect. 2, pp. .xi, 1208, 

1816. As a subgenus of Cercopithecus, to include three species; no type 

designated, but fixed as Simia mona Schreber by J. A. Allen (Bull. Amer. 

Mus. Nat. Hist., 47: 350, 6 Feb. 1925) thus antedating the selection of 

Simia diana as type by Stiles and Orleman (Journ. Mammalogy, 7: 52, 

15 Feb. 1926). 
Miopithecus I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 15: 720, 1037, 1842. 

Genotype Simia talapoin Schreber. 

See Pocock, R. I., Notes on the Cercopithecine genera Rhinostigma and 

Miopithecus Ann. Mag. Nat. Hist. (9) 16: 264-268, text-ff. A, B, Aug. 

1925. 
Cercocephalus Temminck, Esquisses Zool. sur la Cote de Guine, p. 31, 1853. 

Genotype Cercocephalus cephus Temminck =»Simm cephus Linnaeus; a 

lapsus calami for Cercopithecus. 
Callithrix Reichenbach, Vollstand. Naturgesch. d. Affen, p. 105, 1862. As a 

subgenus of Cercopithecus; type by tautonymy, Cercopithecus callithrix 

L Geoff roy = <Simzo sabaea Linnaeus. Not Callithrix Erxleben, 1777. 
Petaurista Reichenbach, Vollstand. Naturgesch. d. Affen, p. 105, 1862. As a 

subgenus of Cercopithecus; type Simia petaurista Schreber. Preoccupied 

by Petaurista Link, 1795, a rodent. 
Diademia Reichenbach, Vollstand. Naturgesch. d. Affen, p. 107, 1862. As a 

subgenus of Cercopithecus; type not specified but fixed by Pocock (Proc. 

Zool. Soc. London, p. 678, 8 Oct. 1907) as Simia leucampyx Fischer = 

Cercopithecus mitis Wolf, thus rendering invalid the later designation of 

Simia diana Linnaeus as type by Stiles and Orleman (Journ. Mammalogy, 

7: 52, 15 Feb. 1926). 
Mona Reichenbach, Vollstand. Naturgesch. d. Affen, p. 109, 1862. As a sub- 
genus of Cercopithecus; type Simia' mona Schreber. 
Chlorocebus Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., 

pp. 5, 24, 1870 (in part). No type specified, but includes six species, of 

which Pocock (Proc. Zool. Soc. London, p. 678, 8 Oct. 1907) designated 

Simia sabaea Linnaeus as genotype. 
Cynocebus Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., 

p. 26, 1870. As a subgenus of Chlorocebus; type Cercopithecus cynosurus 

Geoffroy. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 139 

Diana Trouessart, Revue et Mag. de Zool., (3) 6: 124, 1878. As a subgenus of 
Cercopithecus; type C. diana (Linnaeus). Preoccupied by Diana Risso, 
1826, in Pisces. 

Rhinostictus Trouessart, Cat. Mamm. Viv. Foss., new ed., pt. 1, p. 17, 1897. 
As a subgenus of Cercopithecus for the nictitans group to replace Pelaurista 
Reichenbach, preoccupied; type Simia petaurista Schreber. 

Otopithecus Trouessart, Cat. Mamm. Viv. Foss., new ed., pt. 1, p. 22, 1897. 
As a subgenus of Cercopithecus; no type specified, but fixed by Pocock 
(Proc. Zool. Soc. London, p. 678, 8 Oct. 1907) as Cercopithecus pogonias 
Bennett. 

Pogonocebus Trouessart, Cat. Mamm. Viv. Foss., Suppl., pt. 1, p. 14, 1904. 
As a subgenus of Cercopithecus for the Diana and Brazza Guenons; no type 
specified; but fixed by Pocock (Proc. Zool. Soc. London, p. 678, 8 Oct. 
1907) and by Stiles and Orleman (Journ. Mammalogy, 7: 52, 15 Feb. 1926) 
as Simia diana Linnseus. 

Allochrocebus Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 
Hist., 1 : p. xl; 2: 296, 1912 ( = 15 June 1913). As a subgenus of Lasiopyga; 
type Cercopithecus Vhoesti Sclater. 

Insignicebus Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 
Hist., 1 : p. xl; 2: 296, 1912 ( = 15 June 1913). As a subgenus of Lasiopyga; 
type Cercopithecus albogularis Sykes. 

Melanocebus Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 
Hist., 2: 296, 1912 ( = 15 June 1913). As a subgenus of Lasiopyga: no type 
specified, but several members of the rnitis and nictitans groups are in- 
cluded. A substitute for Diademia Reichenbach. 

Neocebus Elliot, Review of the Primates, Monographs Amer. Mus. Nat. Hist., 
1 : p. xl; 2: 296, 1912 ( = 15 June 1913). As a subgenus of Lasiopyga; type 
Simia cephus Linnaeus. 

Rhinostigma Elliot, Review of the Primates, Monographs Amer. Mus. Nat. 
Hist., 1: xl; 2: 273, pi. 10, 1912 ( = 15 June 1913). Genotype Cercopithe- 
cus hamlyni Pocock. For commentary see Pocock, R. I., Notes on the 
Cercopithecine genera Rhinostigma and Miopithecus, Ann. Mag. Nat. 
Hist. (9) 16: 264-267, Aug. 1925. Elliot has confused the type skull of 
Cercocebus hamlyni Pocock ( = Cercocebus aterrimus) with that of Cerco- 
pithecus hamlyni Pocock. 

oethiops group 

Review: Schwarz, Ernst. Die Meerkatzen der Cercopithecus sethiops- 
Gruppe. Zeitschr. f. Saugetierkunde, 1: 28-47, 31 Aug. 1926. 

Cercopithecus ^thiops .^thiops (Linnaeus) 

Simia aethiops Linnaeus, Syst. Nat., ed. 10, 1: 28, 1758. Based on account in 
Hasselquist, Iter Palest., p. 190, 1757; Ethiopia, Upper Egypt, i.e., Sen- 
naar (see Schwarz, E., Zeitschr. f. Saugetierkunde, 1: 35, 31 Aug. 1926). 



140 bulletin: museum of comparative zoology 

Simia engylitthia Hermann, Observationes Zool., 1: 1, 1804. No locality. 

Placed as a synonym of C. xthiops by Schwarz, 1926. 
Cercopithecus griseo-viridis Desmarest, Encyclop. Methodique, Mamma- 

logie, 1 : 61, 1820. Based on "Le Grivet" of Cuvier (Hist. Nat. des Mamm., 

pt. 7, pi. 39, 1819). "Afrique." 
Simia subviridis F. Cuvier, Dictionnaire des Sci. Nat., 20: 27, 1821. (Africa). 
Cercopithecus griseus F. Cuvier, Hist. Nat. des Mamm., Tabl. Gen. Methodique, 

p. 1, 1824. Based on "Le Grivet" of 1819. 
C[ercopithecus] cano-viridis Gray, List Spec. Mamm. Brit. Mus., p. 5, 13 May 

1843. Wrongly attributed to Riippell. 
C[ercopithecus] cinereo-viridis Gray, List Spec. Mamm. Brit. Mus., p. 5, 13 

May 1843. Wrongly attributed to Temminck. 
Cercopithecus (Chlorocebus) toldti Wettstein, Anzeiger K. Akad. Wiss. Wien, 

53: 189, 1916; Denkschr. K. Akad. Wiss. Wien, math.-nat. KL, 94: 645, 

1918. Jebel Riha, near Kadugli, Kordofan. 
Cercopithecus (Chlorocebus) cailliaudi Wettstein, Denkschr. K. Akad. Wiss. 

Wien, math.-nat. KL, 94: 643, 1918. Blue Nile, Anglo-Egyptian Sudan. 
Lasiopyga {Cercopithecus) weidholzi Lorenz, Ann. Naturh. Mus., Wien, 36: 1, 

pi. 1, Sep. 1922. Egypt. 



Cercopithecus ^thiops arenarius (Heller) 

Lasiopyga pygeryihra arenaria Heller, Smithsonian Misc. Coll., 61 : no. 17, p. 11, 
21 Oct. 1913. Merelle waterholes, Marsabit Road, Kenya Colony. 

Cercopithecus ^thiops callidus (Hollister) 

Lasiopyga pygerythra callida Hollister, Smithsonian Misc. Coll., 59: no. 3, p. 1, 
2 Mar. 1912. South side of Lake Naivasha, Kenya Colony. 

Cercopithecus ^thiops centralis Neumann 

Cercopithecus centralis Neumann, Zool. Jahrb., Syst., 13: 533, 10 Oct. 1900. 

Bukoba, Lake Victoria, Tanganyika Territory. 
Cercopithecus tantalus budgetti Pocock, Proc. Zool. Soc. London, p. 733, 8 Oct. 

1907. Butiaba ("Bathyaba"), Lake Albert, Uganda. 
Cercopithecus tantalus griseistictus Elliot, Ann. Mag. Nat. Hist. (8) 4: 259, 

Sep. 1909. Bambara, Uele River, Congo Beige. 
Cercopithecus (Chlorocebus) cynosurus itimbiriensis Matschie and Dubois, 

Revue Zool. Africaine, 1 : 435, 31 Mar. 1912. Itimbiri River, middle Congo, 

Congo Beige. 
Lasiopyga tantalus beniana Lorenz, Anzeiger K. Akad. Wiss. Wien, 51 : 358, 

1914. Beni Semhki, Congo Beige. 
Lasiopyga tantalus graueri Lorenz, Anzeiger K. Akad. Wiss. Wien, 51 : 358, 

1914. Northwestern Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 141 

Cercopithecus ^thiops cloetei Roberts 

Cercopithecus sethiops cloetei Roberts, Ann. Transvaal Mus., 14: 223, pis. 3, 4, 5, 
figs. 15a, 15b, 15c, 28 July 1931. Mariepskop, Pilgrimsrest district, Trans- 
vaal. 

Cercopithecus ^thiops cynosuros (Scopoli) 

Simia cynosuros Scopoli, Delicise Faunae et Flor. Insubricse, 1 : 44, pi. 19, 1786. 
No locality; but fixed as Banana, lower Congo, Congo Beige (see Schwarz, 
E., Zeitschr. f. Saugetierkunde, 1: 46, 31 Aug. 1926). 

Cercopithecus tephrops Bennett, Proc. Zool. Soc. London, p. 109, 18 Nov. 1833. 
No locality. 

Cercopithecus silaceus Elliot, Ann. Mag. Nat. Hist. (8) 4: 263, Sep. 1909. South 
bank of the Loangwa, Angoniland, northeastern Rhodesia. 

Cercopithecus (Chlorocebus) cynosurus weynsi Matschie and Dubois, Revue 
Zool. Africaine, 1 : 435, 31 Mar. 1912. Banana, mouth of Congo, Congo 
Beige. 

Cercopithecus (Chlorocebus) sethiops lukonzolwae Matschie, Revue Zool. Afri- 
caine, 1: 438, text-f. 2, 31 Mar. 1912. Lukonzolwa, Lake Mweru, Tan- 
ganyika Territory. Probable synonym of C. sethiops cynosuros. 

Cercopithecus (Chlorocebus) cynosurus tholloni Matschie, Revue Zool. Afri- 
caine, 1: 438, 31 Mar. 1912. Brazzaville, Stanley Pool, lower Congo, 
Congo Beige. 

Cercopithecus pygerythrus katangensis Lonnberg, Revue Zool. Africaine, 7: 
141, 1 Oct. 1919. Funda Biabo and Kinda, Katanga, Congo Beige. 
Probable synonym of C. sethiops cynosuros. 

Cercopithecus ^thiops excubitor Schwarz 

Chlorocebus voeltzkowi Matschie, in Voeltzkow's- Reise in Ostafrika, 1: no. 1, 
p. 47, 1923; a nomen nudum. 

Cercopithecus sethiops excubitor Schwarz, Zeitschr. f. Saugetierkunde, 1 : 43, 31 
Aug. 1926. Manda Island, Witu Island, off coast of Tanganyika Terri- 
tory. 

Cercopithecus ^thiops hilgerti Neumann 

Cercopithecus hilgerti Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 50, 

1902. Gobele River, Webi Shebeli, eastern Abyssinia. 
Cercopithecus ellenbecki Neumann, Sjtzb. Ges. Naturf. Freunde, Berlin, p. 50, 

1902. Suksuki and Maki Rivers, Lake Zwai, Abyssinia. 
Cercopithecus djamdjamensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 51, 1902. Bamboo forest near Abera, east of Lake Abaja, Abyssinia. 

Cercopithecus ^thiops johnstoni Pocock 

[Cercopithecus pygerythrus] johnstoni Pocock, Proc. Zool. Soc. London, p. 738, 
8 Oct. 1907. Moshi, Kilimanjaro district, Tanganyika Territory. 



142 bulletin: museum of comparative zoology 

Cercopithecus rubellus Elliot, Ann. Mag. Nat. Hist. (8) 4: 260, Sep. 1909. 

Fort Hall, Kenya Colony. 
Cercopithecus centralis luteus Elliot, Smithsonian Misc. Coll., 56: no. 7, p. 1, 

11 June 1910. Wambugu, southwest of Mt. Kenya, Kenya Colony. 
Lasiopyga pygerythra turnhili Heller, Smithsonian Misc. Coll., 61: no. 17, p. 10, 

21 Oct. 1913. Ndi, Taita Hills, Kenya Colony. 
Lasiopyga pygerythra contigua Hollister, Smithsonian Misc. Coll., 72: no. 2, 

p. 2, 22 Jan. 1920. Changamwe, inland from Mombasa, Kenya Colony. 

Cercopithecus ^thiops marjori^ Bradfield 

Cerpithecus (sic) aeihiops marjorias Bradfield, in a privately printed leaflet, 
dated Benoni, South Africa, 26 Sep. 1935; reprinted in The Auk, 53: 
131, Jan. 1936. Zoetvlei, near Kuruman, Bechuanaland. 

Cercopithecus ^thiops marrensis Thomas and Hinton 
Cercopithecus tantalus marrensis Thomas and Hinton, Proc. Zool. Soc. London, 
p. 248, 6 July 1923. Foothills of Jebel Marra, central Darfur, Anglo- 
Egyptian Sudan. 

Cercopithecus ^thiops matschiei Neumann 

Cercopithecus matschiei Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 51, 

1902. Malo, Omo River, southern Abyssinia. 
Cercopithecus sethiops matschiei Schwarz, Zeitschr. f. Saugetierkunde, 1: 38, 31 

Aug. 1926. 

Cercopithecus ^thiops nesiotes Schwarz 

Chlorocebus pemhae Matschie in Voeltzkow, Reise in Ostafrika, 1: no. 1, p. 184, 
1923; a nomen nudum. Pemba Island. 

Cercopithecus sethiops nesiotes Schwarz, Zeitschr. f. Saugetierkunde, 1: 42, 31 
Aug. 1926. Chake Chake, Pemba Island, off coast of Tanganyika Terri- 
tory. 

Cercopithecus ^thiops ngamiensis Roberts 

Cercopithecus sethiops ngamiensis Roberts, Ann. Transvaal Mus., 15: 19, 1 Oct. 
1932. Toten-Maun road, Ngamiland, Bechuanaland. 

Cercopithecus ^thiops pygerythrus (F. Cuvier) 

Cercopithecus glaucus Lichtenstein, Reisen in Sudlich. Afrika, 1: 645, 1811; 

nomen nudum. 
[Simia] pygerythra F. Cuvier, Hist. Nat. des Mammiferes, pt. 24, pi. [139] and 

text, p. 2, Jan. 1821. "Africa." 
Cercopithecus pygerithrseus Desmarest, Encyclopedic M^thodique, Mam- 

malogie, 2: 534, 1822; errorim. 
Cercopithecus pusillus Desmoulins, Dictionnaire Classique d'Hist. Nat., 7: 

568, 1825. Keiskama, near Great Fish River, Caff raria = Natal. 



ALLEN: CHECKLIST OF AFRICAN AIAMMALS 143 

S[imia] erythropyga G. Cuvier, Regne Anim., ed. 2, 1: 92, 1829. New name for 

pygerythrus. 
Cercopithecus lalandii I. Geoffroy, Arch. Mus. d' Hist. Nat. Paris, (1) 2: 561, 

1841. New name for pusillus. 

Cercopithecus ^thiops rufoviridis I. Geoffroy 

Cercopithecus rufo-viridis I. Geoffroy, Comptes Rend. Acad. Sci. Paris, 15: 

1038, 1842; Arch. Mus. d' Hist. Nat. Paris, (1) 2: 564, 1841. Africa. 
Cercopithecus flavidus Peters, Reise nach Mossambique, Saugeth., p. 3, pi. 

IB, 1852. Quitangonha, Mozambique, 15° S. 
Cercopithecus circumcinctus Reichenbach, VoUstand. Naturgesch. d. Affen, 

p. 123, pi. 21, f. 310, 1862. "W. Afrika" probably a synonym of this race. 
Cercopithecus rufoniger Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats 

Brit. Mus., p. 26, 1870. Lapsus for rufoviridis 1. Geoffroy, to whose plate 

32 (Arch. Mus. d' Hist. Nat. Paris, (1) 2: pi. 32, 1841) his page reference 

evidently refers. 
Cercopithecus pygerythrus whytei Pocock, Proc. Zool. Soc. London, p. 738, 

8 Oct. 1907. Mt. Chiradgula, Nyasaland. 
Cercopithecus pygerythrus helvescens Thomas, Proc. Zool. Soc. London, p. 286, 

29 Apl. 1926. Cunene Falls, Southwest Africa. Probably synonym of C. 

sethiops rufoviridis. 

Cercopithecus ^thiops sab^us (Linnaeus) 

Simia sabsea Linnaeus, Syst. Nat., ed. 12, 1: 38, 1766. Based on Edwards' St. 

Jago, Cape Verde Islands, monkey; doubtless from Senegal (see J. A. 

Allen, Bull. Amer. Mus. Nat. Hist., 47: 352, 6 Feb. 1925). 
Cercopithecus werneri 1. Geoffroy, Compt. Rend. Acad. Sci. Paris, 31 : 874, 1850. 

No locality — Africa. 
Cercopithecus callitrichus 1. Geoffroy, Cat. Methodiqued. Mamm. Mus. d'Hist. 

Nat. Paris, p. 23, 1851 (ex Cuvier). 

Cercopithecus ^thiops tantalus Ogilby 

Cercopithecus tantalus Ogilby, Proc. Zool. Soc. London, p. 33, Oct. 1841. No 

locality. 
Cercopithecus chrysurus Blyth, Ann. Mag. Nat. Hist. (1) 15: 461, July 1845. 

No locality. 
Cercopithecus passargei Matschie, Sjtzb. Ges. Naturf. Freunde, Berlin, p. 74, 

1897. Yola, Benue River, northern Nigeria. 
Cercopithecus pousarguei Mitchell, Proc. Zool. Soc. London, 1: 1, 1 June 1905; 

error for passargei. 
Cercopithecus tantalus alexandri Pocock, Proc. Zool. Soc. London, p. 545, 19 

Oct. 1909. Lake Chad. 
Cercopithecus viridis Schultze, Das Sultanat Bornu, p. 62, 1910. Bornu, Nigeria. 

A nomen nudum, according to Schwarz. 



144 bulletin: museum of comparative zoology 

cephus group 

Cercopithecus cephus cephus (Linnseus) 

Simla cephus Linnaeus, Syst. Nat., ed. 10, 1: 27, 1758. "America" = West 

Africa. 
Cercopithecus buccalis Leconte, Proc. Acad. Nat. Sci. Philadelphia, p. 10, Feb. 

1857. Gaboon. 
[Cercopithecus cephus] cephodes Pocock, Proc. Zool. Soc. London, p. 724, 8 Oct. 

1907. Gaboon. 
Cercopithecus inobservatus ElHot, Ann. Mag. Nat. Hist. (8) 5: 81, Jan. 1910. 

"West Africa," exact locality unknown. See Schwarz, E., Ann. Mag. Nat. 

Hist. (9) 19: 153, Jan. 1927. 
Cercopithecus pulcher Lorenz, Anzeiger K. Akad. Wiss. Wien, math.-nat. Kl., 

52: 171, 1915. Cameroons. 

Cercopithecus cephus erythrotis Waterhouse 

Cercopithecus erythrotis Waterhouse, Proc. Zool. Soc. London, p. 59, July 1838. 

Island of Fernando Po, West Africa. 
Cercopithecus sclateri Pocock, Abstract Proc. Zool. Soc. London, no. 5, p. 18, 

22 Mar. 1904; Proc. Zool. Soc. London, 1: 433, text-f. 87, 2 Aug. 1904, 

Benin, Nigeria. 

diana group 

Cercopithecus diana diana (Linnaeus) 

Simia diana Linnseus, Syst. Nat., ed. 10, 1: 26, 1758. "Guinea" i.e., Liberia 

(see Jentink, Notes Leyden Mus., 20: 233, 1898). 
Simia faunus Linnseus, Syst. Nat., ed. 12, 1: 36, 1766. No locality. 
Cercopithecus diana var. ignita Gray, Cat. Monkeys, Lemurs and Fruit-eating 

Bats Brit. Mus., p. 22, 1870. "West Africa." 

Cercopithecus diana roloway (Schreber) 

Simia roloway Schreber, Siiugthiere, 1: 109, 186 (Latin name), pi. 25, 1774. 

Guinea. 
C[ercopithecus] palatinus Wagner, Schreber's Saugthiere, Suppl., 5: 47, 1855. 

Guinea. 

Cercopithecus dryas Schwarz 

Cercopithecus dryas Schwarz, Revue Zool. et Bot. Africaines, 21: 251, 15 Mar. 
1932. Lomela, on Lomela River, Sankuru district, Congo Beige. 

Vhoesti group 

Cercopithecus hamlyni Pocock 

Cercopithecus hamlyni Pocock, Ann. Mag. Nat. Hist. (7) 20: 521, Dec. 1907; 

Proc. Zool. Soc. London, p. 160, pi. 10, f. 3 (colored), 17 Sep. 1908. Ituri 

Forest, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 145 

Cercopithecus leucampyx aurora Thomas and Wroughton, Trans. Zool. Soc. 

London, 19: 485, Mar. 1910. South end of Lake Kivu, Congo Beige. 
Rhinostigma hamlyni Elliot, Review of the Primates, Monographs Amer. Mus. 

Nat. Hist., 2: 273, pi. 10, 30, 1912 ( = 15 June 1913). 

Cercopithecus l'hoesti l'hoesti P. L. Sclater 

Cercopithecus l'hoesti P. L. Sclater, Proc. Zool. Soc. London, for 1898, p. 586, pi. 

48, 1 Apl. 1899. Congo. 
Cercopithecus thomasi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 262, 

1905. East shore. Lake Kivu, Congo Beige. 
Cercopithecus thomasi rutschuricus Lorenz, Anzeiger K. Akad. Wiss. Wien, 

math.-nat. Kl., 52: 172, 1915. Mountains bounding east side of Rut- 

schuru plains, eastern Congo Beige. 

Cercopithecus l'hoesti preussi Matschie 

Cercopithecus preussi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 76, 
1898. Victoria and Cameroon Mt., Cameroons. 

Cercopithecus crossi Forbes, Nature (London), 72: 630, 26 Oct. 1905. Cam- 
eroons. 

Cercopithecus preussi insularis Thomas, Ann. Mag. Nat. Hist. (8) 5: 191, Feb. 
1910. Northern Bantabiri, Island of Fernando Po, West Africa ("the 
differences described for the Fernando Po race appear to be only individ- 
ual" fide Schwarz, E., Ann. Mag. Nat. Hist. (10) 1: 658, June 1928). 

C. mitis group; C. leucampyx group of Schwarz (192S); subgenera 
Melanocebus and Insignicebus of Elliot 

Cercopithecus mitis mitis Wolf 

Cercopithecus mitis Wolf, Abbildungen u. Beschreibungen merkwiirdiger natur- 

gesch.' Gegenstande, 2: 145, pi. 34, 1822 (Nlirnberg). Identified as the 

Angola form of Diademed Monkey by Schwarz; based on a menagerie 

animal. 

See Schwarz, E., Cercopithecus mitis Wolf fur Simia leucampyx Fischer. 

Zeitschr. f. Saugetierkunde, 8: 279, 20 Dec. 1933. 

Simia leucampyx J. B. Fischer, Synopsis Mamm., p. 20, 1829. "In Guinea", 
but "perhaps, after all, Angola" (see Schwarz, E., Ann. Mag. Nat. Hist. 
(10) 1:653, June 1928). 

Cercopithecus diadematus I. Geoffroy, Belanger's Voyage aux Indes-Orientales, 
Zool., p. 51, 1834. West Coast of Africa. 

Cercopithecus dilophos Ogilby, Library of Entertaining Knowledge, The Me- 
nageries, Nat. Hist. Monkeys, Opossums and Lemurs, 1: 343, 1838, based 
on Cuvier's description of the Diana Monkey. Temminck, Esquisses Zool. 
sur la Cote de Guine, p. 36, 1853. No locality. 

Cercopithecus pluto Gray, Proc. Zool. Soc. London, p. 56, text-f., pi. 3, 2 Dec. 
1848. Angola. 



146 bulletin: museum of comparative zoology 

[Cercopithecus stuhlmnnni] nigrigenis Pocock, Proc. Zool. Soc. London, p. 692, 
pi. 39, f. 1, 9 Oct. 1907. West Africa. 

Cercopithecus mitis albogularis (Sykes) 

Semn[opithecus] ?albogularis Sykes, Proc. Committee of Sci. and Correspon- 
dence Zool. Soc. London, pt. 1, p. 106, 5 Aug. 1831. Type locality fixed as 
Zanzibar (Schwarz, E., Ann. Mag. Nat. Hist. (9) 19: 152, Jan. 1927). 

Semnopithecus alhigularis Lesson, Spec. Mamm., p. 64, 1840. 

Cercopithecus mitis albotorquatus Pousargues 

Cercopithecus albotorquatus Pousargues, Bull. Mus. d'Hist. Nat., Paris, 2: 55, 

1896. No locality. 
Cercopithecus rufotinctus Pocock, Proc. Zool. Soc. London, p. 706, 9 Oct. 1907. 

?Mombasa, Kenya Colony; but see Ann. Mag. Nat. Hist. (9) 19: 153, 

Jan. 1927, where it is said to be the form of the Juba River forests. 
Cercopithecus (Insignicebus) albogularis zammaranoi de Beaux, Atti Soc. Ital. 

Sci. Nat. e Mus. Civ. Stor. Nat. Milano, 62: 248, Feb. 1924. Bidi Scionde, 

lower Juba, Italian Somaliland. 

Cercopithecus mitis boutourlinii Giglioli 

Cercopithecus boutourlinii Giglioli, Zool. Anzeiger, 10: 510, 26 Sep. 1887. 

Kaffa, Abyssinia. 
Cercopithecus omensis Thomas, Proc. Zool. Soc. London, for 1900, p. 801, 

1 Apl. 1901. Mursu, Omo River, 40 miles north of Lake Rudolf. 

Cercopithecus mitis doggetti Pocock 

[Cercopithecus stuhlmanni] doggetti Pocock, Proc. Zool. Soc. London, p. 691, 

9 Oct. 1907. Southwestern Ankole, between Lakes Victoria and Albert 

Edward, Uganda. 
Lasiopyga leucampyx sibatoi Lorenz, Anzeiger K. Akad. Wiss. Wien, math.- 

nat. Kl., 50: 439, 1913. Mountain forest, northwest of Lake Tanganyika, 

Congo Beige, 2000 meters. 

Cercopithecus mitis erythrarchus Peters 

Cercopithecus erythrarchus Peters, Reise nach Mossambique, Siiugeth., p. 1, pi- 

1, 1852. Inhambane, south of the Zambesi, Mozambique (see also Schwarz, 

E., Ann. Mag. Nat. Hist. (9) 19: 151, Jan. 1927). 
Cercopithecus stairsi P. L. Sclater, Proc. Zool. Soc. London, p. 580, pi. 40, 1892. 

Delta of the Zambesi, Mozambique. 
Cercopithecus albogularis beirensis Pocock, Proc. Zool. Soc. London, p. 701, 

9 Oct. 1907. Beira, Mozambique. 
[Cercopithecus stairsi] mossambicus Pocock, Proc. Zool. Soc. London, p. 705, 

9 Oct. 1907. Mozambique. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 147 

Cercopithecus mitis kandti Matschie 

Cercopithecus kandti Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 264, 

1905. Volcanoes north of Lake Kivu, between Lakes Tanganyika and 

Albert Edward, Congo Beige. 
Cercopithecus insignis Elliot, Ann. Mag. Nat. Hist. (8) 4: 274, Sep. 1909. 

Congo Forest, West Africa. 

Cercopithecus mitis kibonotensis Lonnberg. Blue Monkey 

Cercopithecus albogularis kibonotensis Lonnberg, Wiss. Ergebn. d. Schwed. Zool. 

Exped. Kilimandjaro 1905-6 u. Leitung v. Sjostedt, Upsala, 2: Mamm., 

p. 5, 1908. Kibonoto, near Mt. Kilimanjaro, Tanganyika Territory. 
Lasiopyga albogularis maritima Heller, Smithsonian Misc. Coll., 61: no. 17, 

p. 8, 21 Oct. 1913. Mazeras, Kenya Colony. 
Lasiopyga albogularis kima Heller, Smithsonian Misc. Coll., 61: no. 17, p. 9, 

21 Oct. 1913. Mt. Mbololo, Taita district, Kenya Colony. 

Cercopithecus mitis kolbi Neumann 

Cercopithecus kolbi Neumann, Proc. Zool. Soc. London, 2: 144, Oct. 1902. 

Kedong escarpment, east side of Mt. Kenya, Kenya Colony. 
Cercopithecus kolbi hindei Pocock, Proc. Zool. Soc. London, p. 703, pi. 39, f. 3, 

9 Oct. 1907. Tutha, Kenya district, Kenya Colony, 8000 feet. 
Cercopithecus kolbi nubilus Dollman, Ann. Mag. Nat. Hist. (8) 5: 202, Feb. 1910. 

Nairobi Forest, Kenya Colony. 

Cercopithecus mitis labiatus L Geoff roy 

Cercopithecus labiatus I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 15: 1038, 

1842; Arch. Mus. d'Hist. Nat. Paris, (1) 2: 555, 1841. [South] Africa. 
Cercopithecus samango Wahlberg in Sundevall, Ofvers. Kongl. Svenska Vet.- 

Akad. Forhandl., Stockholm, for 1844, 1 : 160, 1845. Inland of Port Natal, 

Natal. 
Cercopithecus chimango Temminck, Esquisses Zool. sur la Cote de Guin6, p. 32, 

1853; emendation of samango Wahlberg. 

Cercopithecus mitis m^si Lonnberg 

Cercopithecus leucampyx msesi Lonnberg, Revue Zool. Africaine, 7: 133, 1 Oct. 
1919. Kutu, Lake Leopold H district, Congo Beige. 

Cercopithecus mitis moloneyi P. L. Sclater 

Cercopithecus moloneyi P. L. Sclater, Proc. Zool. Soc. London, p. 252, pi. 17, 

Aug. 1893. Karonga, north end of Lake Nyasa, Mozambique. 
Cercopithecus francescse Thomas, Ann. Mag. Nat. Hist. (7) 10: 243, Sep. 1902. 

Near Mt. Waller, west side of Lake Nyasa, Nyasaland. 



148 bulletin: museum of comparative zoology 

Cercopithecus mitis monoides I. Geoffrey 

C[ercopithecuSi monoides I. Geoffrey, Arch. Mus. d'Hist. Nat. Paris, (1) 2: 
558, pi. 31, 1841. Lectotype, Rufiji River, 8° S., Tanganyika Territory 
(Schwarz, E., Ann. Mag. Nat. Hist. (10) 1: 656, June 1928). 

[Cercopithecus albogularis] rufilatus Pocock, Proc. Zool. Soc. London, p. 702, 
9 Oct. 1907. Rufiji River (south of Zanzibar), Tanganyika Territory. 

Cercopithecus mitis neumanni Matschie 

Cercopithecus neumanni Matschie, Sitzb. Ges. Naturf. Freunde, BerHn, p. 266, 

1905. Kwa Kitoto, northern Kavirondo, Tanganyika Territory. 
Lasiopyga leucampax (sic) mause Heller, Smithsonian Misc. Coll., 61: no. 17, 

p. 7, 21 Oct. 1913. Summit of Mau escarpment, between Londiani and 

Sirgoit, Kenya Colony. 

Cercopithecus mitis nyas^e Schwarz 

Cercopithecus leucampyx nyasse Schwarz, Ann. Mag. Nat. Hist. (10) 1: 656, 
footnote, June 1928. Fort Lister, Mlanje, southern Nyasaland, 3500 ft. 

Cercopithecus mitis opisthostictus P. L. Sclater 

Cercopithecus opisthostictus P. L. Sclater, Proc. Zool. Soc. London, for 1893, 
p. 725, Apl. 1894. Lake Mweru, Northern Rhodesia. 

Cercopithecus mitis phylax Schwarz 

Cercopithecus leucampyx phylax Schwarz, Sitzb. Ges. Naturf. Freunde, Berlin, 
for 1926, p. 32, 1927. Patta Island, Witu group, off coast of Kenya Colony. 

Cercopithecus mitis schoutedeni Schwarz 

Cercopithecus leucampyx schoutedeni Schwarz, Revue Zool. et Bet. Africaines, 
16: 126, 15 Sep. 1928. Kwidjwi Island in Lake Kivu, Congo Beige. 

Cercopithecus mitis stuhlmanni Matschie 

Cercopithecus stuhlmanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 225, 

1893. North of Kinyawanga, northwest of Lake Albert, Congo Beige. 
Cercopithecus otoleucus P. L. Sclater, Proc. Zool. Soc. London, 1 : 237, pi. 25, 

1 Aug. 1902. Latuka Mts., northern Uganda. 
[Cercopithecus stuhlmanni] carruthersi Pocock, Proc. Zool. Soc. London, p. 691, 

9 Oct. 1907. Ruwenzori East, Uganda, 10,000 feet. 
Cercopithecus princeps Elliot, Ann. Mag. Nat. Hist. (8) 4: 304, Sep. 1909. Fort 

Portal, Mpanga Forest, Toro, Uganda. 
Cercopithecus {Mono) leucampyx schubotzi Matschie, Ann. Soc. Roy. Zool. et 

Malacol. Belgique, Bruxelles, 47: 78, Dec. 1913. Mawambi, Ituri River, 

Congo Beige. 
Cercopithecus leucampyx elgonis Lonnberg, Revue Zool. Africaine, 7: 133, 1 Oct. 

1919. Mt. Elgon, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 149 

Cercopithecus mitis schwarzi Roberts 

Cercopithecus leucampyx schwarzi Roberts, Ann. Transvaal Mus., 14: 222, pis. 

3, 4, 5, f. 14a, 14b, 14c, 28 July 1931. Mariepskop, Pilgrimsrest district, 

Transvaal. 

mo7ia group 

Cercopithecus mona mona (Schreber). Mona Monkey 

Simia mona Schreber, Saugthiere, 1: 97, 185 (Latin name), pi. 15, 1775. "Bar- 
bary." 

Simia monacha Schreber, Saugthiere, 5: pi. 15B, 1804. Africa. Wagner, who 
completed the Suppl. vol. 1, credits this plate and name to Schreber, as a 
synonym of Cercopithecus mona, in Suppl. 1: 128, 1834. 

Cercopithecus mona, var. monella Gray, Cat. Monkeys, Lemurs and Fruit- 
eating Bats Brit. Mus., p. 22, 1870. Senegal. 

Cercopithecus mona campbelli Waterhouse 

Cercopithecus campbelli Waterhouse, Proc. Zool. Soc. London, p. 61, July 1838. 
Sierra Leone. 

Cercopithecus temminckii Ogilby, Library of Entertaining Knowledge, The 
Menageries, Nat. Hist. Monkeys, Opossums, and Lemurs, 1: 345, 1838. 
No locality — coast of Guinea. Probably unidentifiable but may be camp- 
belli. 

Cercopithecus burnettii Gray, Ann. Mag. Nat. Hist. (1) 10: 256, Dec. 1842. 
Island of Fernando Po. 

Cercopithecus cambelli Temminck, Esquisses Zool. sur la Cote de Guin6, p. 33, 
1853; errorim. 

Cercopithecus mona denti Thomas 

Cercopithecus denti Thomas, Abstract Proc. Zool. Soc. London, no. 38, p. 1, 
22 Jan. 1907; Proc. Zool. Soc. London, p. 2, pi. 1, 12 June 1907. Ituri 
Forest between Mawambi and Avakubi, Congo Beige. 

Cercopithecus (Otopithecus) denti liebrechtsi Dubois and Matschie, Revue Zool. 
Africaine, 1: 439, text-f. 3, 31 Mar. 1912. Stanley Falls, Congo Beige. 

Cercopithecus mona elegans Dubois and Matschie 

Cercopithecus (Otopithecus) elegans Dubois and Matschie, Revue Zool. Africaine, 
1: 440, text-f. 4, 31 Mar. 1912. J*robably Lomani, Congo Beige. 

Cercopithecus mona grayi Fraser 

Cercopithecus grayi Fraser, Cat. Knowsley Coll., p. 8, 1850. 

Cercopithecus erxlehenii Dallbet and Pucheran, Revue et Mag. de Zool. (2) 8: 

96, Feb. 1856. ?West Africa. 
Cercopithecus pogonias pallidus Elliot, Ann. Mag. Nat. Hist. (8) 4: 261, Sep. 

1909. Gaboon. 



150 bulletin: museum of compakative zoology 

Cercopithecus petronellse Biittikofer, Notes Ley den Mus., 34: 1, 1 Dec. 1911. 
"Upper Congo." 

Cercopithecus mona lowei Thomas 

Cercopithecus campbelli lowei Thomas, Ann. Mag. Nat. Hist. (9) 11: 608, May 
1923. Bandama, Ivory Coast. 

Cercopithecus mona nigripes Du Chaillu 

Cercopithecus nigripes Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 360, Nov. 
1860. Gaboon, banks of the Ofoubour River, in the mountains. 

Cercopithecus mona pogonias Bennett 

Cercopithecus pogonias Bennett, Proc. Zool. Soc. London, p. 67, 20 Sep. 1833. 
Island of Fernando Po, West Africa. 

Cercopithecus mona pyrogaster Lonnberg 

Cercopithecus pyrogaster Lonnberg, Revue Zool. Africaine, 7: 137, 1 Oct. 1919. 
Atene, Kwango, Congo Beige. 

Cercopithecus mona wolfi A. B. Meyer 

Cercopithecus wolfi A. B. Meyer, Notes Leyden Mus., 13: 63, Jan. 1891. "Cen- 
tral West Africa." 



neglectus group 

Cercopithecus neglectus Schlegel 

Cercopithecus leucampyx Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats 

Brit. Mus., p. 22, 1870, not Simia leucampyx J. B. Fischer, 1829. White 

Nile. 
Cercopithecus neglectus Schlegel, Mus. du Pays-Bas, Simiae, p. 70, 1876. Type 

obtained by Petherick on "White Nile." 
Cercopithecus hrazzse A. Milne-Edwards, Revue Scientifique, (3) 12=38: 15, 

3 July 1886; P. L. Sclater, Proc. Zool. Soc. London, pp. 225, 443, pi. 33 

(col.), 1893. French Congo. 
■C{ercopithecus] neglectus hrazziformis Pocock, Proc. Zool. Soc. London, p. 687, 

9 Oct. 1907. Locality uncertain, said to be from "French Congo." 
Cercopithecus ezrse Pocock, Abstract Proc. Zool. Soc. London, no. 54, p. 10, 

10 Mar. 1908; Proc. Zool. Soc. London, p. 158, pi. 10, f. 2, 17 Sep. 1908. 
Locality unknown; "probably Upper Congo." 

Cercopithecus hrazzse uelensis Lonnberg, Revue Zool. Africaine, 7: 130, 1 Oct. 
1919. Poko, Uele district, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 151 



nictitans group 

Cercopithecus nictitans nictitans (Linnaeus). White-nosed Monkey 
Simia nictitans Linnaeus, Syst. Nat., ed. 12, 1: 40, 1766. Guinea. 
[Cercopithecus nictitans] laglaizei Pocock, Proc. Zool. Soc. London, p. 698, 9 Oct. 

1907. Gaboon. 
Cercopithecus sticticeps Elliot, Ann. Mag. Nat. Hist. (8) 4: 263, Sep. 1909. 

N'dongo-leti, upper Ubangi River, Congo Beige. 

Cercopithecus nictitans ascanius (Audebert) 

Simia ascanius Audebert, Hist. Nat. des Singes et Makis, fam. 4, sect. 2, fig. 13, 

1799. No locality. 
Cercopithecus melanogenys Gray, Ann. Mag. Nat. Hist. (1) 16: 212, Sep. 1845. 

"Africa." 
Cercopithecus histrio Reichenbach, Vollstand. Naturgesch. d. Affen, p. 106, 

ff. 256-259, 1863. 
Cercopithecus picturatus Santos, Jorn. Sci. Math., Phys. e Nat., Lisboa, (1) 

11: 98, June 1886. West Africa. 
Cercopithecus (Rhinostictus) ascanius cirrhorhinus Matschie, Ann. Soc. Roy. 

Zool. et Malacol. Belgique, Bruxelles, 47: 70, Dec. 1913. Province of 

Stanley Falls, probably from the lower Lomani, Congo Beige. 
Cercopithecus (Rhinostictus) ascanius kassaicus Matschie, Ann. Soc. Roy. Zool. 

et Malacol. Belgique, Bruxelles, 47: 74, Dec. 1913. Pogge Falls, Kasai, 

Congo Beige. 
Cercopithecus (Rhinostictus) ascanius pelorhinus Matschie, Ann. Soc. Roy. Zool, 

et Malacol. Belgique, Bruxelles, 47: 76, Dec. 1913. Bought at Yambuya, 

Congo Beige. 

Cercopithecus nictitans BtJTTiKOFERi Jentink 

Cercopithecus biittikoferi Jentink, Notes Leyden Mus., 8: 56, Jan. 1886. Liberia. 

Cercopithecus nictitans erythrogaster Gray 

Cercopithecus erythrogaster Gray, Proc. Zool. Soc. London, p. 169, pi. 16 (col.), 
Sep. 1866. [?Lagos, Nigeria], West Africa. 

Cercopithecus nictitans katang^ Lonnberg 

Cercopithecus ascanius katangse Lonnberg, Revue Zool. Africaine, 7: 122, 1 Oct. 
1919. Kinda, Katanga, Congo Beige. 

Cercopithecus nictitans martini Waterhouse 

Cercopithecus martini Waterhouse, Proc. Zool. Soc. London, p. 58, July 1838. 

Island of Fernando Po, West Africa. 
Cercopithecus ludio Gray, Proc. Zool. Soc. London, p. 8, Mamm., pi. 9, f. 1 

(where name is misspelled lupio), 11 Dec. 1849. "Said to have been 

brought from the west Africa." 



152 bulletin: museum of comparative zoology 

Cercopithecus stampflii Jentink, Notes Leyden Mus., 10: 10, Apl. 1888. Pessi 

Country, Liberia. 
Cercopithecus insolitus Elliot, Ann. Mag. Nat. Hist. (8) 4: 258, Sep. 1909. 

Northern Nigeria. 

Cercopithecus nictitans montanus Lorenz 

Cercopithecus schmidti montanus Lorenz, Anzeiger K. Akad. Wiss. Wien, math.- 

nat. Kl., 51 : 357, 2 July 1914. Wabembe, northwestern Lake Tanganyika, 

Congo Beige. 

Cercopithecus nictitans mpang^ Matschie 

Cercopithecus (Rhinostictus) schmidti mpangx Matschie, Ann. Soc. Roy. Zool. 

et Malacol. Belgique, Bruxelles, 47: 67, Dec. 1913. Mpanga Forest, 

Uganda. 
Cercopithecus (Rhinostictus) schmidti enkamer Matschie, Ann. Soc. Roy. Zool. 

et Malacol. Belgique, Bruxelles, 47: 73, Dec. 1913. Chima Kilima, north 

of Mawambi, upper Ituri, Congo Beige. 
Lasiopyga schmidti ituriensis Lorenz, Anzeiger K. Akad. Wiss. Wien, math.- 

nat. Kl., 51: 357, 2 July 1914. Ituri Forest, Congo Beige. 

Cercopithecus nictitans petaurista (Schreber) 

Simia petaurista Schreber, Saugthiere, 1 : 103, 186 (Latin name), pi. 19B, 1774. 

"Guinea." 
Simia albinasus Reichenbach, Vollstand. Naturgesch. d. Affen, 1863. 
Cercopithecus fantiensis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, pp. 64, 

98, 1893. Fanti, Rio Boutry, Gold Coast. 
Cercopithecus hiittikoferi pygrius Thomas, Ann. Mag. Nat. Hist. (9) 11: 607, 

May 1923. Bandama, Ivory Coast. 

Cercopithecus nictitans schmidti Matschie 

Cercopithecus schmidti Matschie, Zool. Anzeiger, 15: 161, 2 May 1892. Man- 
yema, west of north end of Lake Tanganyika. (No type locality is desig- 
nated, but this is given as the origin of Schmidt's specimen.) 

Lasiopyga ascanius kaimosx Heller, Smithsonian Misc. Coll., 61: no. 17, p. 10, 
21 Oct. 1913. Upper Lukosa River, near Kaimosi mission station, Kenya 
Colony. 

Cercopithecus (Rhinostictus) schmidti sassse Matschie, Ann. Soc. RoJ^ Zool. et 
Malacol. Belgique, Bruxelles, 47: 72, Dec. 1913. Sassa, west of Lake 
Albert Edward, Congo Beige. 

Cercopithecus schmidti rutschuricus Lorenz, Ann. K. K. Naturhist. Hofmus. 
Wien, 31: 228, pi. 15, f. 2, 1917. Mountains on east border of Rutschuru 
plains, Congo Beige. 

Cercopithecus ascanius orientalis Lonnberg, Revue Zool. Africaine, 7: 125, 1 Oct. 
1919. Camp Simba, Zuwani River, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 153 

Cercopithecxjs nictitans signatus Jentink 

Cercopithecus signatus Jentink, Notes Leyden Mus., 8: 55, Jan. 1886. West 
Africa, "perhaps from Banana," Congo. 

Cercopithecus nictitans whitesidei Thomas 

Cercopithecus ascanius whitesidei Thomas, Ann. Mag. Nat. Hist. (8) 4: 542, 

Dec. 1909. Nsoli, Ikau, upper Lulanga River, Congo Beige, about 1° N., 

22° E. 
Cercopithecus {Rhinostictus) ascanius omissus Matschie, Ann. Soc. Roy. Zool. 

et Malacol. Belgique, Bruxelles, 47: 68, Dec. 1913. Locality unknown; 

brought by a caravan coming from Manyema, west of the north end of 

Lake Tanganyika, Congo Beige. 

talapoin group 

Cercopithecus talapoin talapoin (Schreber) 

Simia talapoin Schreber, Saugthiere, 1: 101, 186 (Latin name), pi. 17, 1774. 

No locality. 
S[im,ia] (Cercopithecus) niger Kerr, Linnseus's Animal Kingdom, p. 71, 1792. 

No locality, but refers to Bosman's voyage, hence probably West Africa. 

Possibly a synonym of Cercopithecus talapoin (Schreber). 
Cercopithecus pileatus I. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 110, 1812. 

No locality. 
Miopithecus capillatus I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 15: 720, 

1842. No locality; on a later page (1037) the author retracted this sup- 
posed species. 
Cercopithecus melarhinus Schinz, Systemat. Verzeichn. Saugeth., 1: 47, 1844. 

"Africa." 

Cercopithecus talapoin ansorgei Pocock 

[Cercopithecus talapoin] ansorgei Pocock, Proc. Zool. Soc. London, p. 742, 9 
Oct. 1907. Canhoca (not "Cambaca"), Angola. 

Cercopithecus talapoin pilettei (Lonnberg) 

Miopithecus talapoin pilettei Lonnberg, Revue Zool. Africaine, 7: 119, 1 Oct. 
1919. Kabawaki, Ruwenzori, Congo Beige, 2500 m. 

Erythrocebus Trpuessart. Red Monkeys 

Erythrocebus Trouessart, Cat. Mamm. Viv. Foss., new ed., pt. 1, p. 19, 1897. 
As a subgenus of Cercopithecus; no type designated, but of the four species 
included, Cercopithecus patas (Schreber) is here made the type. 

Simia, Cercopithecus, Cercocebus, Chlorocebus, in part, of older authors. 

Review: Schwarz, Ernst. Die Formen der Gattung Erythrocebus. Sitzb. 
Ges. Naturf. Freunde, Berlin, for 1926, p. 24-31, 1927. 



154 bulletin: museum of comparative zoology 

Erythrocebtjs patas patas (Schreber) 

Simia patas Schreber, Siiugthiere, 1 : 98, 185 (Latin name), pi. 16, 1774. Senegal. 

Simia rubra Gmelin, Linnseus's Syst. Nat., ed. 13, 1: pt. 1, p. 34, 1788. No 

locality; based on Buff on. 
Simia (Cercopithecus) ruber nigro-fasciatus Kerr, Linnseus's Animal Kingdom, 

p. 71, 1792. Senegal, Congo. 
Simia (Cercopithecus) ruber albo-fasciatus Kerr, Linnseus's Animal Kingdom, 

p. 71, 1792. Senegal, Congo. 
Simia rufa Schreber, Siiugthiere, Suppl., pi. 16B, 1804?. 
Cercopithecus circu?ncinctus Reichenbach, Die Vollstand. Naturgesch. d. 

Affen, p. 123, pi. 21, f. 310, 1863. ?West Africa. 
Cercopithecus patas sannio Thomas, Ann. Mag. Nat. Hist. (7) 17: 173, 31 Jan. 

1906 (see Schwarz, E., Sitzb. Ges. Naturf. Freunde, Berlin, for 1926, 

p. 29, 1927). Yo, Lake Chad. 
Cercocebus kerstingi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, for 1905, 

p. 274. 2 Feb. 1906 (see Schwarz, E., Sitzb. Ges. Naturf. Freunde, Berlin, 

for 1926, p. 29, 1927). Sokode, Togo, Nigeria. 
Erythrocebus langheldi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, for 

1905, p. 276, 2 Feb. 1906 (see Schwarz, E., Sitzb. Ges. Naturf. Freunde, 

Berlin for 1926, p. 29, 1927). Garua, upper Benua River, Cameroons. 
Erythrocebus zechi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, for 1905, 

p. 274, 2 Feb. 1906 (see Schwarz, E., Sitzb. Ges. Naturf. Freunde, Berlin, 

for 1926, p. 29, 1927). Kete Kradji, western Togoland, Nigeria. 
Range: Senegal to Lake Chad. 

Erythrocebus patas baumstarki Matschie. 

Erythrocebus baumstarki Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 273, 
1905. Ikoma, southeast end of Lake Victoria, Tanganyika Territory. 
Probably a recognizable race (see Schwarz, E., Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1926, p. 28, 1927). 

Erythrocebus patas pyrrhonotus (Hemprich and Ehrenberg) 

Cercopithecus pyrrhonotus Hemprich and Ehrenberg, Symbolse Physicae, 
Mamm., dec. 1, pi. 10 and folio hh, kk, Aug. 1832. Darfur, Anglo- 
Egyptian Sudan. 

Cercopithecus poliophseus Reichenbach, Vollstand. Naturgesch. d. Affen, 
p. 122, pi. 21, f. 309, 1863. Fazogli, Blue Nile, eastern border of Anglo- 
Egyptian Sudan. 

Cercopithecus poliolophus Heuglin, Reise in Nordost-Afrika, 2: 6, 1877. Pro- 
posed as a substitute for Cercopithecus poliophseus Reichenbach. Fazogli, 
Blue Nile, eastern Anglo-Egyptian Sudan. 

Erythrocebus formosus Elliot, Ann. Mag. Nat. Hist. (8) 4: 264, Sep. 1909. 
Uganda. 
A synonym of pyrrhonotus Hemprich and Ehrenberg (see Schwarz, E., 

Sitzb. Ges. Naturf. Freunde, Berlin, for 1926, p. 29, 1927). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 155 

Cercopithecus (Erythrocebus) patas albosignatus Matschie, Revue Zool. Afri- 
caine, 1: 433, text-f. 1, 31 Mar. 1912. Mbomu River, Uele basin, Congo 
Beige. 
Erythrocebus albigenus Elliot, Ann. Mag. Nat. Hist. (8) 4: 265, Sep. 1909. 
Egyptian Sudan. 
A synonym of E. patas pyrrhonotus (see Schwarz, E., Sitzb. Ges. Naturf. 
Freunde, Berlin, for 1926, p. 30, 1927). 

Erythrocebus whilei Hollister, Smithsonian Misc. Coll., 56: no. 2, p. 11, pi. 2, 
31 Mar. 1910. Nzoia River, Guas Ngishu plateau, Kenya Colony. 
A synonym of E. patas pyrrhonotus (see Schwarz, E., Sitzb. Ges. Naturf. 
Freunde, Berlin, for 1926, p. 30, 1927). 

Cercopithecus (Erythrocebus) patas poliomystax Matschie, Revue Zool. Afri- 
caine, 1: 434, 31 Mar. 1912. Upper Congo. 
Range: Sudan, south to northern Uganda and the Guas Ngishu plateau. 

CoMOPiTHECUS J. A. Allen. Hamadryas Baboon 

Comopithecus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 312, 6 Feb. 1925. 
Genotype Simia hamadryas Linnaeus; to replace Hamadryas Lesson, pre- 
occupied in insects. 

Hamadryas Lesson, Spec. Mamm., p. 107, 1840. Genotype Hamadryas 
choeropithecus Lesson = Simia hamadryas Linnseus; preoccupied by 
Hamadryas Hiibner, 1806, for a genus of insects. 

Simia, Cercopithecus, Cynocephalus, Papio, in part, of older authors. 

Comopithecus brockmani (Elliot) 

Papio brockmani Elliot, Ann. Mag. Nat. Hist. (8) 4: 248, Sep. 1909. Dirre 
Dawa, Somaliland, 3500 feet. 
Elliot insists that this is a species distinct from Comopithecus hamadryas 
(Linnseus) . 

Comopithecus hamadryas (Linnseus). Hamadryas 

Si7nia hamadryas Linnaeus, Syst. Nat., ed. 10, 1: 27, 1758. Egypt; based on 

Alpius. 
Simia cynamolgos Linnseus, Syst. Nat., ed. 10, 1: 28, 1758. Based on Simia 

asgyptiaca Linnseus, in Hasselquist, 1757, hence Upper Egypt. 
Cynocephalus wagleri L. Agassiz, Oken's Isis, 21 : 862, pi. 11, 1828. No locality; 

probably same as Comopithecus-hamadryas (Linnseus). 
Hamadryas chosropithecus Lesson, Spec. Mamm., p. 109, 1840. "L'Abyssinie, 

I'Arabie, et I'Egypte." 
Hamadryas segyptiaca Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats 

Brit. Mus., p. 34, 1870. New name for Simia hamadryas Linnseus and 

Simia aegyptiaca Hasselquist. Egypt, Arabia, Abyssinia. 
Range: Eastern Ethiopia and eastern Sudan, mainly in the lowlands; to 

Arabia. 



156 bulletin: museum of compakative zoology 



Macaca Lacepede 

Simia Linnaeus, Syst. Nat., ed. 10, 1: 25, 1758. Genotype Simia sylvanus 

Linnaeus. 
On status of the generic name Simia, see Thomas, O., Proc. Zool. Soc. 
London, p. 125, 22 Mar. 191 1 ; Stiles, C, Science, n. ser., 64 : 138, 6 Aug. 1926; also 
Stiles, C. W., and Orleman, M. B., The nomenclature for Man, the Chim- 
panzee, the Orang-Utan and the Barbary Ape, Bull. Hygienic Lab., U. S. 
Pub. Health Service, Washington, no. 145, p. 1-60, 1927. 

By suspension of rules, the names Simia, Simia satyrus and Pithecus are 
suppressed, see Opinion 114, of Internat. Comm. on Nomenclature, Smith- 
sonian Misc. Coll., 73: no. 6, p. 25, 8 June 1929. 
Macaca Lacepede, Tabl. Mamm., p. 4, 1799. Genotype Simia inuus Linnaeus = 

Simia sylvanus Linnaeus. 
Inuus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 100, 1812. Genotype 

Inuus ecaudatus E. Geofiroy = Simia sylvanus Linnaeus. 
Sylvanus Oken, Lehrbuch d. Naturgesch., pt. 3: zool., sect. 2, p. 1223, 1816. 

Genotype Inuus ecaudatus E. Geoffroy =*Simia sylvanus Linnaeus. Not 

Sylvanus Rafinesque, 1815, but synonymous with Sylvanus Virey, 1819, 

genotype Simia sylvanus Linnaeus. Preoccupied by Sylvanus Latreille, 

1807, in Coleoptera. 
Pithes Burnett, Quart. Journ. Sci., Lit. and Art, 26: 307, 1828. Genotype 

Pithes sylvanus = Simia sylvanus Linnaeus. 
Perhaps a misprint for Pithecus. 
Pithecus Gervais, Hist. Nat. des Mamm., 1: 94, 1854, in part. 
Innuus Encyclop. Britannica, Mamm., ed. 8, 14: 141, 1857; misspelling of 

Inuus. 

MAdACA sylvanus (Linnaeus). Barbary Ape 

Simia sylvanus Linnaeus, Syst. Nat., ed. 10, 1 : 25, 1758. "In Africa, Ceylona" = 

Barbary Coast. 
Simia inuus Linnaeus, Syst. Nat., ed. 12, 1 : 35, 1766. "Africa." 
Simia pithecus Schreber, Saugthiere, 1: pi. 4B, 1799. From Buffon. 
Inuus ecaudatus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 100, 1812. 

Mediterranean coast of Africa, and Gibraltar. 
Inuus pithecus Lesson, Spec. Mamm., p. 99, 1840. 
Pithecus pygmaeus Reichenbach, Vollstand. Naturgesch. d. Affen, p. 145, 1863. 

Range: Morocco and Algeria; introduced on the rock of Gibraltar. 

See Joleaud, L. Le magot de Berberie. Bull. Soc. d'Acclimatation, Paris, 
78: 153-155, 1931. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 157 

Mandrillus Ritgen. Mandrills 

Mandrillus Ritgen, Naturl. Eintheilung d. Saugeth., p. 33, 1824, where the 
name is credited to Cuvier, who, however, used only the French form. 
Based on "Les Mandrills" of Cuvier, i.e., Si77iia maimon Linnaeus and 
Simla mormon Alstromer, both synonyms of Siviia sphinx Linnaeus; hence 
genotype Simia sphinx Linnaeus. See Allen, J. A., Bull. Amer. Mus. Nat. 
Hist., 47: 311, 6 Feb. 1925. 

Mandril Voigt, Cuvier's Thierreich, 1: 88, 1831. As a subgenus of Simia; in- 
cluded two species, neither designated as type. 

Mormon Wagner, Schreber's Saugthiere, Suppl., 1 : 164, 1839. As a subgenus 
of Cynocephalus; type by tautonymy Simia mormon Alstromer = Simia 
sphinx Linnaeus, 1758. Preoccupied by Mormon Illiger, 1811, in birds. 

Drill Reichenbach, VollstJind. Naturgesch. d. Affen, p. 162, 1862. As a sub- 
genus of Mormon; type by monotypy, Simia leucophsea F. Cuvier. 

Chseropithecus Gray, Cat. Monkeys, Lemurs, and Fruit-eating Bats Brit. Mus., 
p. 35, 1870. Genotype Simia leucophsea F. Cuvier. Not Chseropithecus 
Blainville, 1839. 

Maimon Trouessart, Cat. Mamm. Viv. Foss., Suppl., pt. 1, p. 21, 1904. As a 
subgenus of Papio; type by tautonymy Simia maimon Linna?us = 5mta 
sphinx Linnaeus. Not Maimon Wagner, 1839, for a wholly different group. 

Mandrillus sphinx (Linnaeus). The Mandrill 

Simia sphinx Linnajus, Syst. Nat., ed. 10, 1 : 25, 1758. "Borneo" = West Africa. 

Simia maimon Linnaeus, Syst. Nat., ed. 12, 1: 35, 1766. "Zeylona" =West 
AfrifTa, based on an immature specimen of Simia sphinx Linnteus. 

Simia mormon Alstromer, Kongl. Svenska Vet.-Akad. Handl., Stockholm, p. 
144, 1766. 

Simia madarogaster Zimmermann, Geographische Geschichte, 2: 176, 1780. No 
locality. A synonym of Simia maimon Linnaeus = Mandrillus sphinx (Lin- 
na;us), see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 16: 21, 1 Feb. 1902. 

Simia suilla Kerr, Linnaeus's Animal Kingdom, p. 59, 1792. Based on Pennant's 
figure; no locality. 

Papio planirostris Elliot, Ann. Mag. Nat. Hist. (8) 4: 304, Sep. 1909. Fan, 
southern Cameroons. Possibly a synonym of Mandrillus sphinx (Linnaeus). 

Mandrillus schreberi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 344, 25 
Sep. 1917. New name for Simia maimon Schreher = Simia sphinx Lin- 
naeus. 

Mandrillus leucoph^us (F. Cuvier). Drill 

S[imia] (Papio) cinerea Kerr, Linnaeus's Animal Kingdom, p. 62, 1792. Based 
on the Cinereous Baboon of Pennant; unidentifiable, but "strongly sug- 
gestive" of the later-described Simia leucophsea F. Cuvier (see Allen, J. A., 
Bull. Amer. Mus. Nat. Hist., 7: 185, 1895). "Probably . . . Africa." 



158 bulletin: museum of comparative zoology 

Simia leucophsea F. Cuvier, Ann. Mus. d' Hist. Nat. Paris, 9: 477, pi. 37, 1807. 

No locality but "probablement des cotes d'Afrique." 
Cynocephalus drill Lesson, Complements de Buffon, ed. 2, Paris, 1 : 259, 1838. 

New name for Simia leucophsea F. Cuvier. 
Papio mundamensis Hilzheimer, Zool. Anzeiger, 30: 109, 17 Apl. 1906. Mukonje 

Farm, Mundame, Cameroons. A synonym of Mandrillus leucophxus (F. 

Cuvier) according to Elliot, who examined the type. 

Probably the following described forms are not entitled to recognition : 
Mandrillus burlacei (W. Rothschild) 

Maimon burlacei W. Rothschild, Ann. Mag. Nat. Hist. (9) 10: 232, Aug. 1922. 
Bitye, Ja River, Cameroons. Status uncertain. 

Mandrillus escherichi Matschie and Zukowsky 

Mandrillus escherichi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 478, 25 Oct. 1917. Ekododo, on the Temboni, near southern 

border of Spanish Guinea. 

Mandrillus hagenbecki Matschie and Zukowsky 

Mandrillus hagenbecki Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, p. 488, 25 Oct. 1917. Probably coast region near Lagos, Nigeria. 

Mandrillus poensis Zukowsky 

Mandrillus poensis Zukowsky, Arch. f. Naturgesch., 88: sect. A, pt. 3, p. 184, 
1 pi., June 1922. Island of Fernando Po, West Africa. 

Mandrillus insidaris Zukowsky, Arch. f. Naturgesch., for 1925, 91: sect. A, no. 
5, p. 132, text-f., July 1926. Island of Fernando Po, West Africa. Pre- 
sumably = M. poensis Zukowsky. 

Mandrillus tessmanni Matschie and Zukowsky 

Mandrillus tessmanni Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 470, 25 Oct. 1917. Okak-land, on middle Benito, not far from 

Alen (Nkolentengan), Spanish Guinea. 

Mandrillus zenkeri Matschie and Zukowsky 

Mandrillus zenkeri Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 479, 25 Oct. 1917. Probably between Bipindi on the Lokundje, 

and Yaunde, Cameroons. 

Mandrillus zenkeri ebolow^ Matschie and Zukowsky 
[Mandrillus zenkeri] var. eboloivse? Matschie and Zukowsky, Sitzb. Ges. Naturf. 
Freunde, Berlin, p. 493, 25 Oct. 1917. Near Ebolowa, Cameroons. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 159 



Papio Erxleben. Baboons 

Papio Erxleben, Syst. Regni Animalis, p. 15, 1777. Genotype Papio sphinx 
Erxleben (not of Linnaeus, q.e. Cynocephalus papio Desmarest, 1820). 
On the status of Papio "Brisson" (Haak) see Allen, J. A., Bull. Amer. 
Mus. Nat. Hist., 47: 306, footnote, 6 Feb. 1925. 

Cynocephalus Cuvier and Geoffroy, Mag. Encyclop., 3: 462, 1795. Genotype 
Sitnia cynocephalus Linnaeus. Preoccupied by Cynocephalus Boddaert, 
1768, in Insectivora. 

Chseropithecus Gervais, Dictionnaire Pittoresque d'Hist. Nat., 8: 90, 1839. 
As a subgenus of Simia; type Simia cynocephalus Linnaeus. 

Choiropithecus Reichenbach, Vollstand. Naturgesch. d. Affen, p. 151, 1862. 
As a subgenus of Cynocephalus; type Simia porcaria Boddaert. An emenda- 
tion of Chseropithecus Gervais. 

Papio comatus comatus E. Geoffroy. Chacma Baboon 

Simia porcaria Boddaert, Naturforscher, 22: 17, pi. 1, f. 1-2, 1787. Africa. 

Not Simia porcaria Brtinnich, Dyrenes Hist., 1: 13, pi. 1, 1782 = Macaca 

nemestrina (Linnaeus); see Schwarz, E., Ann. Mag. Nat. Hist. (10) 14: 

258, Aug. 1934. Probably Simia porcaria Zimmermann (Geogr. Geschich., 

2: 197, 1780) is unidentifiable (see J. A. Allen, Bull. Amer. Mus. Nat. 

Hist., 16: 22, 1 Feb. 1902). 
Simia sphingiola Hermann, Observationes Zool., 1 : 2, 1804. The type locality, 

Sumatra, precludes this being an African species (see Schwarz, E., Ann. 

Mag. Nat. Hist. (10) 14: 258, Aug. 1934); but no locality is given in above 

citation. 
S[imia] (Cercopithecus) hamadryas ursinus Kerr, Linnseus's Animal Kingdom, 

p. 39, 63, 1792. Based on Pennant, Hist. Quadrupeds, p. 179, no. 86, 1781; 

Cape of Good Hope. 
Papio comatus E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 103, 1812. 

Cape of Good Hope. For the use of this name in place of Papio porcarius 

and on the synonymy see Schwarz, E., Ann. Mag. Nat. Hist. (10) 14: 

258, Aug. 1934. 
C[ynocephalus] ursinus Wagner, Schreber's Saugthiere, Suppl., 1: 162, .pi. 

8B, 1840; 5: 65, 1855. Cape of Good Hope. 
Papio porcarius orientalis Goldblatt, South African Journ. Sci., 23: 782, pi. 16, 

f. la, 2a, 2c, Dec. 1926. Eastern South Africa. 
Papio porcarius occidentalis Goldblatt, South African Journ. Sci., 23: 782, 

pi. 16, f. lb, 2b, 2d, Dec 1926. Western Transvaal and Cape of Good 

Hope. 
Choiropithecus porcarius transvaalensis Zukowsky, Carl Hagenbeck's Illus- 

trierte Tier- u. Menschenwelt (Hamburg), 2: pt. 3, p. 57, Sept. 1927. 

Northern Transvaal. 
Papio cynocephalus porcarius Schwarz, Zeitschr. f. Saugetierkunde, 3: 212, 

11 June 1928. 



160 bulletin: museum of comparative zoology 

Papio cynocephalus jubilseus Schwarz, Zeitschr. f. Saugetierkunde, 3: 211, 11 
June 1928. Missale, 14° S., 33° 10' E., northeastern Rhodesia. 

Papio comatus chobiensis Roberts 

Papio porcarius chobiensis Roberts, Ann. Transvaal Mus., 15: 18, 1 Oct. 1932. 
Kabulabula, Chobe River, Bechuanaland. 

Papio comatus griseihes Pocock 

Papio porcarius griseipes Pocock, Abstract Proc. Zool. Soc. London, no. 93, 

p. 17, 28 Mar. 1911; Proc. Zool. Soc. London, p. 558, 15 Sep. 1911. Pot- 

chefstroom, Transvaal. 
Although Shortridge (Mamm. South West Africa, 1: 9, 1934) regards this 
as a species distinct from Papio comatus, Schwarz (Ann. Mag. Nat. Hist. (10) 
14: 258, Aug. 1934) beheves it is a race of the Chacma, and thinks it probable 
that the type may have come from the northern Transvaal instead of 
from Potchefstroom. 
Papio porcarius nigripes Roberts, Ann. Transvaal Mus., 15: 19, 1 Oct. 1932. 

Magalakuin River, northern Transvaal. Probably a synonym of Papio 

comatus griseipes Pocock. 

Papio comatus ngamiensis Roberts 

Papio porcarius ngamiensis Roberts, Ann. Transvaal Mus., 15: 18, 1 Oct. 1932. 
Maun, Ngamiland, Bechuanaland. 

Papio cynocephalus cynocephalus (Linnaeus). Yellow Baboon 

Simia cynocephalus Linnaeus, Syst. Nat., ed. 12, 1: 38, 1766. "Africa," re- 
garded by Anderson (Zool. of Egypt, Mamm., p. 64, 1902) as inland from 
Mombasa, Kenya Colony, a decision which "may well be accepted as 
frnal" (Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 315, 6 Feb. 1925). 

Papio sphinx Erxleben, Syst. Regni Animalis, p. 15, 1777. Tropical Africa 
("in calidissimis Africae Indiaque"). Not Simia sphinx Linnaeus, 1758, = 
Mandrillus sphinx (Linnaeus). 

S[i7nia] (Papio) variegata Kerr, Linnaeus's Anim. Kingdom, p. 62, 1792. Based 
on the Yellow Baboon of Pennant; "Africa." 

Simia sylvicola Shaw, General Zool., 1: Mamm., pt. 1, p. 22, pi. 12, 1800. No 
locality; based on the Wood Baboon of Pennant, from Guinea. 

Simia sublutea Shaw, General Zool., 1 : Mamm., pt. 1, p. 23, 1800. No locality; 
based on the Yellow Baboon of Pennant. 

Cynocephalus babouin Desmarest, Encyclop. Methodique, Mammalogie, 1 : 68, 
1820. No locality, except northern Africa. 

Cynocephalus antiquorum Schinz, in Cuvier's Thierreich (Schinz ed.), 1: 115, 
1821. 

Cynocephalus thoth Ogilby, Proc. Zool. Soc. London, p. 11, July 1843. Locality 
unknown. Type regarded by Anderson (see Mamm. of Egypt, p. 57-64, 
1902) as unquestionably Papio cynocephalus. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 161 

Cercopithecus ochraceus Peters, Reise nach Mossambique, Saugeth., p. 2, pi. la, 
1853. Querimba, Mozambique, 10° - 13° S. 
This is a young baboon, probably of the cynocephalus group according to 

Elliot (Review of the Primates, Monographs Amer. Mus. Nat. Hist., 2: 

121, 1912 = 15 June 1913). 

Cynocephalus langheldi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 233, 
1892. Eastern slope of Unguru Mts., Tanga, Ukami, Usukuma, Tang- 
anyika Territory. 

Papio cynocephalus kind^ Lonnberg 

Papio kindas Lonnberg, Revue Zool. Africaine, 7: 147, 1 Oct. 1919. Kinda, 
Lulua district, Congo Beige. 

Papio cynocephalus rhodesle (Haagner) 

Choir opithecus rhodesix Haagner, South African Journ. Nat. Hist., 1: 83, 
pi. 3, May 1918. Central Rhodesia. 

Papio cynocephalus strepitus Elliot 

Papio strepitus Elliot, Ann. Mag. Nat. Hist. (7) 20: 194, Sep. 1907. Fort 

Johnston, south end of Lake Nyasa, Nyasaland. Possibly same as P. 

cynocephalus cynocephalus. 

Papio doguera doguera (Pucheran). Anubis Baboon 

S[imia] anubis J. B. Fischer, Synopsis Mamm., p. 33, 1829. Africa (according 
to Anderson, perhaps Upper Nile, Meroe Id.), but regarded as "abso- 
lutely undeterminable" by J. A. Allen (Bull. Amer. Mus. Nat. Hist., 47: 
315, 6 Feb. 1925), and hence to be dropped. 

Cynocephalus doguera Pucheran, Revue et Mag. de Zool., (2) 8: 96, Feb. 1856; 
(2) 9: 250, June 1857. Abyssinia. 
On the use of this name rather than the unidentifiable anubis, see Allen, 
J. A., Bull Amer. Mus. Nat. Hist., 47: 315-316, 6 Feb. 1925. 

Papio doguera furax Elliot 

Papio furax Elliot, Ann. Mag. Nat. Hist. (7) 20: 498, Dec. 1907. Baringo, 
northwest of Mt. Kenya, Kenya Colony. 
Range: Northern Guaso Nyiro to Lake Naivasha, Kenya Colony. 

Papio doguera heuglini Matschie 

Papio heuglini Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 81, 1898. 

Near Shilluk Islands, White Nile, Anglo-Egyptian Sudan. 
On status and synonymy see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 
47: 320, 6 Feb. 1925. 
Papio lydekkeri W. Rothschild, Novitates Zool., 9: 140, 10 Apl. 1902. Upper 

Blue NUe. 



162 bulletin: museum of comparative zoology 

P[apio] anubis, subsp. olivaceus De Winton, in Anderson's Zool. Egypt, Mamm., 

p. 53, 1902, in part; not Cynocephalus olivaceus I. Geoffroy = Papzo papio 

Desmarest. 
Papio nigerias Elliot, Ann. Mag. Nat. Hist. (8) 4: 247, Sep. 1909. Ibi, northern 

Nigeria. 
Papio werneri Wettstein, Anzeiger K. Akad. Wiss. Wien, math.-nat. Kl., 53: 

190, 1916. Gebel Talodi, near Talodi, Nuba Mts., southern Kordofan. 

Papio doguera ibeanus Thomas 

Papio thoth ibeanus Thomas, Ann. Mag. Nat. Hist. (6) 11: 47, Jan. 1893. 
Lamu, Kenya Colony. 

Papio doguera neumanni Matschie 

Papio neumanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 161, 1897. 

Donyo Ngai and Wadi Natron, Masailand, Tanganyika Territory. 
Papio anubis testes Heller, Smithsonian Misc. Coll., 61: no. 19, p. 10, 8 Nov. 

1913. Ulukenia Hills, Athi Plains, Kenya Colony. Probably same as 

P. doguera neumanni Matschie. 
Papio anubis vigilis Heller, Smithsonian Misc. Coll., 61: no. 19, p. 11, 8 Nov. 

1913. Lakiundu River, near its junction with Northern Guaso Nyiro, 

Kenya Colony. Probably a synonym of Papio doguera neumanni Matschie. 

Papio doguera tessellatus Elliot 

Papio tessellatum Elliot, Ann. Mag. Nat. Hist. (8) 4: 247, Sep. 1909. Mulema, 

Ankole, Uganda. 
On status and synonymy see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 

317, 6 Feb. 1925. 
Papio silvestris Lorenz, Anzeiger K. Akad. Wiss. Wien, math.-nat. Kl., 52: 

173, 16 June 1915. Mawambi, Congo Beige. 
Papio graueri Lorenz, Ann. K. K. Naturhist. Hofmus. Wien, 31: 236, 1917. 

Rutschuru Plains, Congo Beige. 

Papio papio (Desmarest). Western Baboon 

Cynocephalus papio Desmarest, Encyclop. Methodique, Mammalogie, 1 : 69, 

1820. West Africa, "la cote de Guin^e." 
C[ynocephalus] choras Ogilby, Proc. Zool. Soc. London, p. 12, July 1843. West 

Africa, "brought from the Niger Expedition." 
Cynocephalus olivaceus I. Geoffroy, Arch. Mus. d'Hist. Nat. Paris, 5:531 (name 

only), 543, footnote, 1851. Gulf of Benin, Guinea, West Africa. 
Papio rubescens Temminck, Esquisses Zool. sur la Cote de Guine, p. 39, 1853. 

West Africa, exact locality of the coast unknown. 
Papio yokoensis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 89, 1900. 

Yoko, on the Sanaga River, Cameroons. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 163 

Papio pruinosus Thomas 

Papio pruinosus Thomas, Proc. Zool. Soc. London, for 1896, p. 789, pi. 38, 

Apl. 1897. Fort Johnston = Lesumbwe, Monkey Bay, south end of Lake 

Nyasa, Nyasaland. 
Probably an individual variation. 

Theropithecus I. Geoffroy. Gelada Baboon 

Theropithecus I. Geoffroy, Arch. Mus. d'Hist. Nat. Paris for 1841, 2: 576, 1843. 
Genotype Macacus gelada Rlippell. 

Gelada Gray, List Spec. Mamm. Brit. Mus., pp. xvii, 9, 13 May 1843. Geno- 
type Gelada riippelln Gray = Macacus gelada Rtippell. 
"Although published in the same year," Theropithecus is universally given 

recognition. 

Theropithecus gelada (Rlippell). Gelada Baboon 

Macacus gelada Rlippell, Neue Wirbelthiere zu d. Fauna v. Abyssinien ge- 

horig, Saugeth., p. 5, pi. 2, 1835. Mountains of southern Abyssinia, in 

provinces of Haremat, Simien, and Godjam. 
Theropithecus senex Pucheran (ex Schimper MS.), Revue et Mag. de Zool., 

(2) 9: 243, June 1857. Abyssinia. 
Elliot (Review of the Primates, Monographs Amer. Mus. Nat. Hist., 2: 156, 
1912, =15 June 1913) shows that this is doubtless a synonym of T. gelada 
(Rtippell). 
Theropithecus nedjo ReichenbachJ Vollstand. Naturgesch. d. Affen, p. 204, 

1863. New name for Theropithecus obscurus Heuglin = Theropithecus 

gelada (Rlippell). 
Theropithecus obscurus Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Dresden, 

30: Abhandl., no. 2, suppl., p. 10, 1863. Sources of Takazza River, Galla 

country, southern Abyssinia, 6-10,000 feet. 
Gelada ruppellii Gray, List Spec. Mamm. Brit. Mus., p. 9, 13 May 1843; Cat. 

Monkeys, Lemurs, and Fruit-eating Bats Brit. Mus., p. 33, 1870. New 

name for Macacus gelada Rlippell. 
Range: Mountainous parts of Abyssinia, mainly above 6000 feet. 

Family COLOBID.E. Leaf-eating Monkeys 

Regarded by Allen, J. A. (Bull. Amer. Mus. Nat. Hist., 47: 431, 6 Feb. 
1925) as a subfamily of Lasiopygidae ( = Cercopithecidae) but by Hollister, N. 
(Bull. U. S. Nat. Mus., no. 99, pt. 3, p. 37, 1924) as a distinct family, Colo- 
bidae. 

CoLOBUS lUiger. Colobus or Guereza Monkeys 

Colobus lUiger, Prodromus Syst. Mamm. et Avium, p. 69, 1811. Genotype 

Simia polycornos Schreber. 
Colobolus Gray, London Medical Repository, 15: 298, 1 Apl. 1821. 



164 bulletin: museum of comparative zoology 

Guereza Gray, Cat. Monkeys, Lemurs and Fruit-eating Bats Brit. Mus., 

pp. V, 19, 1870. Genotype Guereza ruppellii Gray = Colobus polykomos 

abyssinicus (Oken). 
Procolobus Rochebrune, Faune de la S^n^gambie, Suppl., Vert^br^s, pt. 1, 

p. 95, pi. 1, 1886-87. Genotype Colobus verus Van Beneden. 
Tropicolobus Rochebrune, Faune de la Senegambie, Suppl., Vertebres, pt. 1, 

p. 96, pis. 2, 28, 1886-87. Genotype Colobus rufomilratus Peters. 
Piliocolobus Rochebrune, Faune de la Senegambie, Suppl., Vertebres, pt. 1, 

pp. 96, 105, pi. 3-6, 1886-87. Included four species; genotype Simia {Cer- 

copithecus) badius Kerr (see Allen, J. A., Journ. Mammalogy, 1 : 97, Feb. = 

2 Mar. 1920). 
Stachycolobus Rochebrune, Faune de la Senegambie, Suppl., Vertebres, pt. 1, 

pp. 96, 114, 1886-87. Genotype Colobus satanas Waterhouse. 
Pterycolobus Rochebrune, Faune de la Senegambie, Suppl., Vertebres, pt. 1, 

pp. 96, 125, pi. 10, 1886-87. Genotype Colobus vellerosus I. Geoffroy. 
Lophocolobus Pousargues, Bull. Mus. d'Hist. Nat. Paris, 1: 98, text-f. 1, 

[20 Apl.] 1895. As a subgenus of Colobus; type Colobus verus Van Beneden. 

Review: Schwarz, E. On the local races and distribution of the Black and 
White Colobus Monkeys. Proc. Zool. Soc. London, p. 585-598, map, 16 Oct. 
1929. 

Schwarz, E. Bemerkungen liber die roten Stummelaffen. Zeitschr. f. Sauge- 
tierkunde, 3: 92-97, 11 June 1928. 

See also Allen, J. A. Primates collected by the American Museum Congo 
Expedition. Bull. Amer. Mus. Nat. Hist., 47: 283-499, pi. 79-167, text-f. 1-3. 
map, 6 Feb. 1925. 



polykomos section 

Colobus polykomos polykomos (Zimmermann). Black and White Colobus 
Cebus polykomos Zimmermann, Geogr. Geschichte, 2: 202, 1780. Based on 

Pennant's account, in litt. Sierra Leone. 
Simia (Cercopithecus) regalis Kerr, Linnseus's Animal Kingdom, p. 74, 1792. 

Sierra Leone. 
Simia tetradactyla Link, Beytrage Naturgesch., p. 62, 1795. Based on Buffon. 
Simia polycomos Schreber, Saugthiere, pi. 10 D, ?1800 (see Sherborn, Proc. Zool. 

Soc. London, for 1891, p. 590, Apl. 1892). Sierra Leone and Guinea. 
Simia comosa Shaw, Gen. Zoology, Mamm., 1: pt. 1, p. 59, 1800. Based on 

Pennant, hence Sierra Leone. 
Col[obus] ursinus Ogilby, Proc. Zool. Soc. London, p. 98, 9 Oct. 1835. "Algoa 

Bay." 
Range: Forests from French Guinea, through Sierra Leone and Liberia to 

western part of Ivory Coast. 



N 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 165 

COLOBUS POLYKOMOS DOLLMANI Schwarz 

Colobus polykomos dollmani Schwarz, Ann. Mag. Nat. Hist. (9) 19: 155, Jan. 
1927. Bandama, Bandama River, Ivory Coast. 

Colobus polykomos vellerosxjs (I. Geoffroy) 

Semnopithecus vellerosus I. Geoffroy, in Belanger's Voy. aux Indes-Orientales, 

Zool., p. 37, 1834. No locality — "Afrique." 
Semnopithecus bicolor Wesmael, Bull. Acad. Roy. Sci. et Belles-lettres Belg., 

Bruxelles, 2: 236, 1835. "Cotes d'Afrique." 
Colobus leucomeros Ogilby, Proc. Zool. Soc. London, for 1837, p. 69, 22 Jan. 

1838. "Africa;" Gambia River. 
Range: Forests of Gold Coast and Togo. 



satanas section 

Colobus polykomos satanas Waterhouse. Black Colobus 

Colobus satanas Waterhouse, Proc. Zool. Soc. London, p. 58, July 1838. Island 

of Fernando Po, West Africa. 
Semnopithecus anthracinus Leconte, Proc. Acad. Nat. Sci., Philadelphia, p. 10, 

1857. Gaboon. 
Stachycolobus municus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 155, 

16 July 1917. Wurminsog, south of Rio Muni, and west of Noya, French 

Congo. 
Stachycolobus limbarenicus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 

156, 16 July 1917. Limbareni, Ogowe River, Gaboon. 
Stachycolobus zenkeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 157, 

16 July 1917. Bipindi, Lokundje River, southern Cameroons. 
Range: Island of Fernando Po, and adjacent coasts of Gaboon and southern 

Cameroons. 

ahyssinicus section 

Colobus polykomos abyssinicus (Oken) 

Leynur abyssinicus Oken, Lehrb. d. Naturgesch., pt. 3: sect. 2, p. 1182, 1816. 

Abyssinia. 
Colobus guereza Riippell, Neue Wirbelthiere zu d. Fauna v. Abyssinien gehorig, 

1: Saugeth., p. 1, pi. 1, 1835. Province of Godjam, central Abyssinia. 
Guereza ruppellii Gray, Cat. Monkeys, Lemurs, and Fruit-eating Bats Brit. 

Mus., p. 19, 1870. New name for Colobus guereza Riippell. 
Colobus abyssinicus poliurus Thomas, Proc. Zool. Soc. London, for 1900, p. 800, 

1 Apl. 1901. Omo River, southern Abyssinia. 
Colobus (Guereza) poliurus managaschas Matschie, Ann. Soc. Roy. Zool. et 

Malacol. Belgique, Bruxelles, for 1912, 47: 54, Dec. 1913. Managasha 

forest, west of Addis Ababa, upper Hawash, Abyssinia. 
Range: Mountain forests of Abyssinia, west of the Hawash River. 



166 bulletin: museum of comparative zoology 

CoLOBUS POLYKOMOS CAUDATUS Thomas 

Colobus guereza caudatus Thomas, Proc. Zool. Soc. London, p. 219, pi. 12, 

1 Aug. 1885. Useri, northeast slope of Mt. Kilimanjaro, Tanganyika 

Territory, 3000 feet. 
Colobus alhocaudatus Lydekker, Proc. Zool. Soc. London, for 1905, 2: 328, 5 Apl. 

1906. Substitute for caudatus. 

Colobus polykomos doding^ Matschie 

Colobus (Guereza) matschiei dodingse Matschie, Ann. Soc. Roy. Zool. et Malacol. 
Belgique, Bruxelles, for 1912, 47: 52, Dec. 1913. Southwestern Dodinga 
Mts., about 4° 10' N., 33° 42' E., southern Mongalla, Anglo-Egyptian 
Sudan. 

Colobus polykomos gallarum Neumann 

Colobus gallarum Neumann, Sitzb. Ges. Naturf. T'reunde, Berlin, p. 49, 1902. 

Mountains near headwaters of Webi Shebeli: Gara Mulata near Harar and 

Djaffa Mts. in Arussi Land, Abyssinia. 
Range: Mountain forests east of the Hawash River, Abyssinia. 

Colobus polykomos kikuyuensis Lonnberg 

Colobus abyssinicus kikuyuensis Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 63, 

Jan. 1912. Escarpment Station, Kenya Colony. 
Colobus (Guereza) caudatus thikse Matschie, Ann. Soc. Roy. Zool. et Malacol. 

Belgique, Bruxelles, for 1912, 47: 56, Dec. 1913. West slope of Mt. Kenya, 

Kenya Colony. 
Colobus (Guereza) caudatus laticeps Matschie, Ann. Soc. Roy. Zool. et Malacol. 

Belgique, Bruxelles, for 1912, 47: 57, Dec. 1913. West slope of Mt. Kenya, 

Kenya Colony. 
Range: Mt. Kenya and uplands of Kenya Colony east of Rift Valley. 

Colobus polykomos matschiei Neumann 

Colobus matschiei Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 15, 1899. 

Kwa Kitoto, near Kisumu, Kavirondo, Kenya Colony. 
Colobus abyssinicus roosevelti Heller, Smithsonian Misc. Coll., 61: no. 17, p.5, 

21 Oct. 1913. Mau forest, near Njoro, Kenya Colony. 
Colobus abyssinicus elgonis Granvik, Acta LTniv. Lundensis, (2) 21: no. 3, p. 5, 

pi. 1, 1925. East slope of Mt. Elgon, Kenya Colony. 
Range: Mountain forest west of Rift Valley, from Elgeyo Escarpment and 

Mau forest throughout the Nandi and Elgon region to the small forest 

near Kisumu, Lake Victoria, Kenya Colony. 

Colobus polykomos occidentalis (Rochebrune) 

Guereza occidentalis Rochebrune, Faune de Senegambie, Suppl., Vertebres, p. 
140, pi. 13, 1886-87. Noki, upper Congo. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS . 167 

Colobus (Guereza) escherichi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 342, July 1914. Gombe, below Ikelemba, Sanga River, French Congo. 
Range : From Ubangi and upper Shari district to western slope of Cameroons 
plateau. 

Colobus polykomos percivali Heller 

Colobus abyssinicus percivali Heller, Smithsonian Misc. Coll., 61: no. 17, p. 6, 
21 Oct. 1913. Mt. Gargues, Mathews Range, Kenya Colony. 

Colobus polykomos uellensis Matschie 

Colobus (Guereza) matschiei uellensis Matschie, Ann. Soc. Roy. Zool. et Mala- 
col. Belgique, Bruxelles, for 1912, 47:47, Dec. 1913. Uele River, Congo 
Beige. 

Colobus (Guereza) matschiei ituricus Matschie, Ann. Soc. Roy. Zool. et Malacol. 
Belgique, Bruxelles, for 1912, 47: 48, Dec. 1913. Ituri, Congo Beige. 

Colobus (Guereza) matschiei dianx Matschie, Ann. Soc. Roy. Zool. et Malacol. 
Belgique, Bruxelles, for 1912, 47: 49, Dec. 1913. Kissenge, northeast shore 
of Lake Albert Edward, Congo Beige. 

Colobus (Guereza) matschiei brachychaites Matschie, Ann. Soc. Roy. Zool. et 
Malacol. Belgique, Bruxelles, for 1912, 47:53, 1913. Modi, between Kaya 
and Dufile, Lado. 

Colobus abyssinicus terrestris Heller, Smithsonian Misc. Coll., 61: no. 17, p. 7, 
21 Oct. 1913. Rhino Camp, Lado Enclave. 

Colobus occidentalis ituricus Lorenz, Anzeiger K. Akad. Wiss. Wien, math.-nat. 
Kl., 51 : 508, 1914. Mawambi, Ituri River, Congo Beige. 

Colobus occidentalis rulschuricus Lorenz, Anzeiger K. Akad. Wiss. Wien, math.- 
nat. Kl., 51 : 508, 1914. Sassa River, southeast of Lake Albert Edward, 
Congo Beige. 
Range: Congo forest, north of the Congo, from Lado Enclave and Lake Vic- 
toria west probably to the great bend of the Ubangi, Congo Beige. 

angolensis section 

Colobus polykomos AooLFi-FRrEDERici Matschie 

Colobus adolfi-friederici Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 337, 

July 1914. Rugege forest, northeast of Lake Kivu, Ruanda, Congo Beige. 
Range: Mountain ranges east of Lake Kivu and northern end of Lake 

Tanganyika. 

Colobus polykomos angolensis P. L. Sclater 

Colobus angolensis P. L. Sclater, Proc. Zool. Soc. London, p. 245, May-Aug. 

1860. Inland from Bembe, northern Angola. 
Colobiis angolensis sandbergi Lonnberg, Arkiv f. Zool., 4: no. 15, p. 1, 2 text-f., 

28 Apl. 1908. Lufizi River, near junction with upper Zambesi River, 

Angola. 



168 bulletin: museum of comparative zoology 

Colobus (Colobus) palliatus weynsi Matschie, Revue Zool. Africaine, 2: 207, 

15 Feb. 1913. Lower Congo. 
Colobus maniemse Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 336, July 

1914. Between Fienimulamba and Kibombo, between Stanleyville and 

Kassongo, middle Luala, Congo Beige. 
Colobus benamakimx Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 337, 

July 1914. Bena Makima, Sankuru River, Congo Beige. 
Range: Forest area south of the Congo. 

Colobus polykomos cottoni Lydekker 

Colobus palliatus cottoni Lydekker, Ann. Mag. Nat. Hist. (7) 16: 432, Oct. 

1905. Zokwa (misprinted Qugo), between Mahagi, Lake Albert, and 

Irumu, Ituri River, Congo Beige. 
Colobus (Colobus) palliatus mawambicus Matschie, Revue Zool. Africaine, 2: 

205, 15 Feb. 1913. Northern Pemba, between Irumu and Mawambi, 

Ituri, Congo Beige. 
Colobus mawambicus nahani Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 335, July 1914. Panga, on the Aruwimi, Congo Beige. 
Range : Eastern Congo basin, in the Uele, Ituri and Aruwimi districts. 

Colobus polykomos palliatus Peters 

Colobus palliatus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 637, 1868. 

East Africa, opposite Zanzibar, i.e. Pangani River, Tanganyika Territory. 
Range: Coast ranges of northern Tanganyika Territory and southern 

Kenya Colony. 

Colobus polykomos ruwenzorii Thomas 

Colobus ruwenzorii Thomas, Proc. Zool. Soc. London, 2: 85, 1 Oct. 1901. 
Buamba country, northwest flank of Mt. Ruwenzori, Congo Beige. 

Colobus polykomos sharpei Thomas 

Colobus sharpei Thomas, Proc. Zool. Soc. London, 1: 118, June 1, 1902. Fort 

Hill, Nyasa-Tanganyika plateau, Nyasaland. 
Colobus langheldi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 337, July 

1914. No exact locality; skin brought from Ujiji, Tanganyika Territory. 



badius section 

(Pocock, Proc. Zool. Soc. London, for 1935, p. 939, 10 Jan. 1936, provision- 
ally assigns the red colobus monkeys to a separate genus, Procolobus.) 

Colobus badius badius (Kerr). Red Colobus 

Simia (Cercopithecus) badius Kerr, Linnajus's Animal Kingdom, p. 74, 1792. 
Based on the Bay Monkey of Pennant; Sierra Leone. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 169 

Simia ferruginea Shaw, General Zool., Mamm., 1 : pt. 1, p. 59, 1800. Based on 

the Bay Monkey of Pennant; Sierra Leone. 
Colobus ferruginosus E. GeofFroy, Ann. Mus. d'Hist. Nat. Paris, 19; 92, 1812. 

New name for Simia ferruginea Shaw. "La Guinee." 
Colobus rufoniger Ogilby, Library of Entertaining Knowledge, The Menageries, 

Nat. Hist, of Monkeys, Opossums and Lemurs, 1 : 273, 1838. Sierra Leone. 
Range : Sierra Leone and Liberia. 

Colobus badius pennantii Waterhouse 

Colobus pennantii Waterhouse, Proc. Zool. Soc. London, p. 57, July 1838. 
Island of Fernando Po, West Africa. 
Range : Confined to Fernando Po. 

Colobus badius preussi (Matschie) 

Piliocolobus preussi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 183, 
1900. Barombi, Elephant Lake, northern Cameroons. 

Colobus badius temminckii (Kuhl) 

Colobus temminkii (sic) Kuhl, Beitrage Zool., p. 7, 1820. No locality. 

Colobus fuliginosus Ogilby, Proc. Zool. Soc. London, p. 97, 9 Oct. 1835. Gambia. 

Colobus riifo-fuliginus Ogilby, Library of Entertaining Knowledge, The Men- 
ageries, 1: 269, 270, f. 3, 1838. Substitute for C. fuliginosus Ogilby. 

Cercopithecus temminckii Ogilby, Library of Entertaining Knowledge, The 
Menageries, 1: 345, 1838, "is obviously . . . but a misquotation (gen- 
erally repeated) of Colobus temminckii, Kuhl = Colobus badius temminckii, 
Kuhl" (see Schwarz, E., Ann. Mag. Nat. Hist. (10) 1: 650, footnote, 
June 1928). "Coast of Guinea." 
Range: Upper Senegal, and Gambia to French Guinea. 

Colobus badius verus Van Beneden 

Colobus verus Van Beneden, Bull. Acad. Roy. Sci. et Belles-Lettres Belgique, 

Bruxelles, 5: 347, pi., 1838. "Afrique." 
Semnopithecus (Colobus) olivaceus Wagner, Schreber's Saugthiere, Suppl., 1: 

309, 1840. Substitute for Colobus verus Van Beneden. 
Colobusf? chrysurus Gray, Ann. Mag. Nat. Hist. (3) 17: 77, Jan. 1866. West 

Africa (type a trade skin in British Museum; may be a synonym of C. 

badius verus Van Beneden). ^ 
Colobus cristatus Gray, Ann. Mag. Nat. Hist. (3) 17: 77, Jan. 1866; Proc. Zool. 

Soc. London, pi. 15, fig., 1868. West Africa. 
Range: Gold Coast and Togo. 

Colobus badius waldroni Hayman 

Colobus badius waldroni Hayman, Proc. Zool. Soc. London, for 1935, p. 915, 
pi. 1 (col.), 10 Jan. 1936. Goaso, Ashanti, Gold Coast. 



170 bulletin: museum of comparative zoology 



ihoUoni section 

CoLOBUS BADius BOUViERi (Rochebrune) 

Piliocolobus houvieri Rochebrune, Faune de la Senegambie, Suppl., Vert6br6s, 

pt. 1, p. 108, pi. 4, 1887. Gambia, whence came Bouvier's specimen, may 

be selected as type locality. 
Colobus (Piliocolobus) likualae Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 346, July 1914. Sanga River, opposite mouth of Likuala River, French 

Congo. 

Colobus badius tholloni Milne-Edwards 

Colobus tholloni Milne-Edwards in Riviere, Revue Scientifique, (3) 12=38: 15, 

3 July 1886. Mission Brazza, Lower Congo. 
Colobus (Piliocolobus) lovizettii Matschie, Revue Zool. Africaine, 2: 207, 15 

Feb. 1913. Kutu, Lake Leopold II, Congo Beige. 

rufomitratus section 

Colobus badius ellioti Dollman 

Colobus ellioti Dollman, Ann. Mag. Nat. Hist. (8) 4: 475, Nov. 1909. Ninety 

km. west of south end of Lake Albert Edward, probably near Oso River, 

Congo Beige. 
Piliocolobus anzeliusi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 339, 

July 1914. Upper Ituri Province, Congo Beige. 
Piliocolobus ellioti melanochir Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 339, July 1914. Between Beni and Irumu, Upper Ituri River, Congo 

Beige. 
Colobus variabilis Lorenz, Anzeiger K. Akad. Wiss. Wien, math.-nat. Kl., 51 : 

383, 1914. Ituri forest, Congo Beige. 
Colobus multicolor Lorenz, Anzeiger K. Akad. Wiss. Wien, math.-nat. KI., 51: 

385, 1914. Mawambi, Ituri district, Congo Beige. 
Range: Rainforest in region of the Lindi, Semliki, and Ituri-Aruwimi 

Rivers, south to the Lowa, Congo Beige. 

Colobus badius foai Pousargues 

Colobus foai Pousargues, Bull. Mus. d'Hist. Nat., Paris, 5: 278, 1899. Ouroua 

country, between southwest of Lake Tanganyika and upper Congo, 

Congo Beige. 
Piliocolobus kabambarei Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 338, 

July 1914. Near Kabambare, between Baraka and Kasongo, Lualaba 

River, west of Lake Tanganyika, Congo Beige. 
Piliocolobus lulindicus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 338, 

July 1914. Lulindi River, near Kasongo, Congo Beige. 
Range: Forest between northern Tanganyika and the Congo. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 171 

CoLOBUS BADius GRAUERi Dollman 

Colobus graueri Dollman, Ann. Mag. Nat. Hist. (8) 4: 474, Nov. 1909. Wabem- 
beland, 80 kilom. west of the north end of Lake Tanganyika, Congo Beige. 
Range: Mountain forest northwest of Lake Tanganyika. 

Colobus badius gordonorum (Matschie) 

Piliocolobus gordonorum Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 186, 
1900. LTzungwe Mts., Uhehe, Tanganyika Territory. 

Colobus badius gudoviusi (Matschie) 

Tropicolobus gudoviusi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 340, 
July 1914. Between Ussuwi and Ihangiro, west of Lake Victoria, Tan- 
ganyika Territory. 

Colobus badius langi J. A. Allen 

Colobus langi J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 443, pi. 102-103, 6 
Feb. 1925. Risimu, between Stanley vUle and Bafwaboli, Congo Beige. 

Colobus badius nigrimanus Trouessart 

Colobus nigrimanus Trouessart, Bull. Mus. d'Hist. Nat., Paris, 12: 444, 1906. 
Liranga, left bank of Congo, French Equatorial Africa. 

Colobus badius oustaleti Trouessart 

Colobus oustaleti Trouessart, Bull. Mus. d'Hist. Nat., Paris, 12: 443, 1906. 

Youmba country, Ubangi basin, Congo Beige. 
Colobus (Tropicolobus) umbrinus Matschie, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 343, July 1914. Bungi, Sanga River, between Wesso and 

Ikelemba, Congo Beige. 

Colobus badius powelli Matschie 

Colobus (Piliocolobus) powelli Matschie, Ann. Soc. Roy. Zool. et Malacol. 

Belgique, Bruxelles, for 1912, 47: 61, Dec. 1913. Zokwa, between Mahagi 

and Irumu, west of Lake Albert, upper Ituri, Congo Beige, 5350 feet. 
Colobus (Tropicolobus) schubotzi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 345, July 1914. Near Koloka, between the Likati and Bima Rivers, 

Congo Beige. 
Colobus (Piliocolobus) brunneus Lonnberg, Revue Zool. Africaine, 7: 112, 1 

Oct. 1919. Sili, district of Upper Uele, Congo Beige. 
Range: Gallery forest in region of Lake Albert, the Uele, Kibali, Rubi, 

Likati Rivers, Congo Beige. 

Colobus badius rufomitratus Peters 

Colobus rufomitratus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 829, 
pis. lA, 2, 1879. Muniuni, Tana River, Kenya Colony. 
Range: Confined to the gallery forests of the lower Tana River. 



172 bulletin: museum of comparative zoology 

COLOBUS BADIUS TEPHROSCELES Elliot 

Colobus tephrosceles Elliot, Ann. Mag. Nat. Hist. (7) 20: 195, Sep. 1907. 
Ruahara River, Toro, east side of Mt. Ruwenzori, Uganda, 4000 feet. 

kirkii section 

Colobus badius kirkii Gray 

Colobus kirkii Gray, Proc. Zool. See. London, p. 180, pi. 15, May 1868. Zanzi- 
bar Island. 



Family PONGID.E. Great Apes 
Pan Oken. Chimpanzees 

Simia of various authors, in part, not of Linnaeus, 1758, of which the type is 
Simia sylvanus. On the suppression of Simia from zoological nomenclature 
see Opinion 114, International Commission on Zool. Nomenclature (Smith- 
sonian Misc. Coll., 73: no. 6, p. 25, 8 June 1929). 

Troglodytes E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 87, 1812. Genotype 
Troglodytes niger E. Geoffroy = Simia satyr us Linnaeus. Preoccupied by 
Troglodytes Vieillot, 1806, in birds. 

P[an] Oken, Lehrb. d. Naturgesch., pt. 3, zool., sect. 2, pp. xi, 1230, 1816. 
Genotype Pan africanus Oken = Simia satyrus Linnaeus. 

Mimetes Leach, Thomson's Ann. Philos., 16: 104, Aug. 1820. Genotype Simia 
satyrus Linnaeus; preoccupied by Mimetes Hlibner, 1816, in insects. 

Theranthropus Brookes, Cat. Anat. and Zool. Mus. of Joshua Brookes, London, 
p. 28, 1828 (previous to July 14). Genotype Theranthropus niger — Troglo- 
dytes niger E. Geoffroy. 

Anthropopithecus Blainville, Ann. Franq:. et Etrang. d'Anat. et Physiol., Paris, 
2: 360, 1838. Genotype Anthropopithecus troglodytes = Simia troglodytes 
Gmelin. 

Hylanthropus Gloger, Hand- u. Hilfsbuch d. Naturgesch., 1 : pp. xxvii, 34, 1841. 
Genotype Hylanthropus troglodytes = Simia troglodytes Gmelin. 

Satyrus Mayer, Arch. f. Naturgesch., 22: sect. 1, p. 281, 1856. Genotype not 
specified; included Orang, Gorilla, and Chimpanzee. Not Satyrus Lin- 
naeus, 1760, nor of Oken, 1816. 

Pseudanthropos Reichenbach, Fortsetzung Vollstand. Naturgesch. d. Affen, 
1860 (see Palmer, T. S., No. Amer. Fauna, no. 23, p. 589, 1904). New name 
to replace Troglodytes E. Geoffroy, 1812, preoccupied. 

Pongo Haeckel, Gen. Morphol. d. Organismen, 2: cl, footnote, 1866. New name 
for Troglodytes E. Geoffroy, preoccupied; not Pongo Lacepede, 1799. 

Engeco Haeckel, Gen. Morphol. d. Organismen, 2: cl and footnote, clx, 1866. 
Genotype Simia troglodytes Gmelin. 

Andropithecus Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 286, Oct. 1868. 
Nomen nudum for the Chimpanzee and the Gorilla. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 173 

Anthropithecus Haeckel, Syst. Phylogenie d. Wirbelthiere, 3: 600, 1895. Emen- 
dation of Anthropopithecus. 

Fsihego De Pauw, Notes sur la Solidification et le Montage des grands Mammi- 
feres (Belgium), p. 13, 1905. Based on Chimpanzee of the Ituri; geno- 
type by monotypy Fsihego ituriensis Matschie. 

Pseudanthropus Elliot, Review of the Primates, Monographs 'Amer. Mus. Nat. 
Hist., 1: xxxvii, 1912 = 15 June 1913, emendation. 

Chimpanza Haime. 

Review: Schwarz, Ernst. On the local races of the Chimpanzee. Ann. Mag. 
Nat. Hist. (10) 13: 576-583, June 1934. See also, for critical notes, Allen, J. A., 
Bull. Amer. Mus. Nat. Hist., 47: 477-497, pi. 152-165, 166, f. 2, 6 Feb. 1925. 

Pan troglodytes troglodytes (Blumenbach) Chimpanzee 

Simia satyrus Linnseus, Syst. Nat., ed. 10, 1: 25, 1758. Africa; locality re- 
stricted to Mayumba, French Congo, by Schwarz (Ann. Mag. Nat. Hist. 
(10) 13: 580, June 1934), who adds, "The original specimen described by 
Tulpius upon which Simia satyrus was founded is said to have come from 
the coast of Angola; there is, however, no proof that it really came from 
there." Suppressed, see Smithsonian Misc. Coll., 73: no. 6, 8 June 1929. 

Simia troglodytes Blumenbach, Handb. d. Naturgesch., p. 65, 1799. Not Homo 
troglodytes Linnaeus (Syst. Nat., ed. 10, 1 : 24, 1758), which was based upon 
an account of an abnormally hairy woman and albino negroes (see 
Thomas, O., Proc. Zool. Soc. London, p. 125, Mar. 1911). "Angola." 

Simia satyrus pongo Kerr, Linnseus's Animal Kingdom, p. 56, 1792. "Java 
and interior parts of Guinea." 

Simia satyrus jocko Kerr, Linnaeus's Animal Kingdom, p. 56, 1792. No locality. 

Troglodytes niger E. Geoffroy, Ann. Mus. d'Hist. Nat. Paris, 19: 87, 1812. 
Coast of Angola (but probably this is erroneous). 

Pan africanus Oken, Lehrb. d. Naturgesch., pt. 3, zool., sect. 2, p. 1231, 1816. 
"Congo, Angola, Guinea, Sierra Leona." 

Troglodytes leucoprymnus Lesson, lUustr. Zool., pi. 32, 1831. Coast of "Guin^e.' 

Anthropopithecus pan Lesson, Spec. Mamm. Bimanes et Quadrumanes, p. ?, 1840. 

Troglodytes tschego Duvernoy, Arch. Mus. d'Hist. Nat., Paris, 8: 1, pi. 1, 1855; 
the name appears as legend to pi. 1 and in title. West coast of Africa. 

Satyrus lagaros Mayer, Arch. f. Naturgesch., 22: sect. 1, p. 282, 1856. No 
locality. 

Satyrus chimpanse Mayer, Arch. f. Naturgesch., 22: sect. 1, p. 282, 1856. No 
locality. 

Troglodytes calvus Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 296, July 1860. 
Interior of Gaboon, south of Cape Lopez. 

Troglodytes kooloo-kamba Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 358, 
Nov. 1860. Ashankolo Mts., upper Ovenga River, French Congo. 

Troglodytes vellerosus Gray, Proc. Zool. Soc. London, p. 181, Sep. 1862. Came- 
roon Mountains, northern Ca'meroons. 



174 bulletin: museum of comparative zoology 

Troglodytes aubryi Gratiolet and Alix, Nouv. Arch. Mus. d'Hist. Nat. Paris, 

(1) 2: 1-258, pi. 1-9, 1866. Probably Gaboon. 
Pseudanthropus fuliginosus Schaufuss, Nunquam Otiosus, Zool. MittheU., p. 

345, 1870. French Congo, exact locahty doubtful. 
Troglodytes angustimanus Brehm, 1871 (not seen). 
Anthropopithecus fuscus A. B. Meyer, Abhandl. u. Ber. K. Zool. Anthrop.- 

Ethnogr. Mus., Dresden, 5: no. 14, p. 7, footnote, pi., 12 Sep. 1895. No 

locality; provisional name. 
Troglodytes livingstonii Selenka, Studien liber Entwicklungsgesch. d. Tiere, pt. 

7, art. 2, p. 104, 1899; nomen nudum. 
Simia pygmaeus raripilosus W. Rothschild, Proc. Zool. Soc. London, for 1904, 

2: 422, 430, text-f. Ill, 18 Apl. 1905. Interior of French Congo, no 

exact locality. 
A [nthropopithecus] ellioti Matschie, Sitzb. Ges. Naturf . Freunde, Berlin, p. 327, 

July 1914. Near Bascho, northern Cameroons. 
Anthropopithecus oertzeni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 327, 

July 1914. Near Bascho, northern Cameroons. 
Anthropopithecus reuteri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 328, 

July 1914. Mouth of Dume River, in the Kadei, southern Cameroons. 
Anthropopithecus ochroleucus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 329, July 1914. North of Sangmelima, upper Lobo River, southern 

Cameroons. 
Anthropopithecus schneideri Matschie, Zeitschr. f. Ethnol., 51 : 75, text-f. 4-5, 

1919. Fernan Vaz district, French Congo. Probably the hinterland south 

of Eliva Nkomi. 
Anthropopithecus pusillus Matschie, Zeitschr. f. Ethnol., 51 : 76, 1919. Probably 

lower Ogowe, between Cape Lopez and Lake Anengue, Gaboon. 
Anthropopithecus papio Matschie, Zeitschr. f. Ethnol., 51: 80, text-f. 7, 1919. 

Barombi, Elephant Lake, Cameroons. 
Anthropopithecus maior and minor Stiles, Bull. Hygien. Lab., LT. S. Dept. Publ. 

Health, no. 115, p. 53, 1927. 
According to Schwarz, E., (Ann. Mag. Nat. Hist. (10) 13: 580, footnote, 
June 1934) these names "must stand to the credit of Stiles" since "Erxleben 
never used them in a technical sense, but only as mere descriptions." 
Anthropopithecus hecki Koch, Zool. Garten, (2) 5: 100, 105, text-f. 1, Aug. 1932. 

Hinterland of Cameroons. 

Pan troglodytes schweinfurthii (Giglioli). Long-haired Chimpanzee 
Troglodytes schweinfurthii Giglioli, Ann. Mus. Civ. Stor. Nat., Genova, 3: 114, 
footnote, 135, 1872. Upper Uele drainage, Niam-niam country, north- 
eastern Congo Beige. 
Troglodytes niger var. mariingensis Noack, Zool. Jahrb., Syst., 2: 291, pi. 10, 
f. 38-41, 7 May 1887. Marungu, west of Lake Tanganyika, Tanganyika 
Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 175 

Fsihego ituriensis De Pauw, Notes sur la Solidification et le Montage des grands 

Mammiferes, St. Nicolas, Belgium, p. 13, pi. 1, 1905. Ituri and central 

Congo Beige. 
Simia (Anthropopithecus) nahani Matschie, Revue Zool. Africaine, 2: 118, 31 

Sep. 1912. Banalia, Aruwimi River, Congo Beige. 
Simia (Anthropopithecus) ituricus Matschie, Revue Zool. Africaine, 2: 121, 31 

Sep. 1912. Makala-Avakubi road, Ituri basin, Congo Beige. 
Simia {Anthropithecus} (sic) coitoni Matschie, Revue Zool. Africaine, 2: 124, 

31 Sep. 1912. Sassa (Ishasha) River, southeast of Lake Albert Edward, 

western border of Uganda. 
Simia (Anthropopithecus) adolfi-friederici Matschie, Ann. Soc. Roy. Zool. et 

Malacol. Belgique, Bruxelles, for 1912, 47: 46, Dec. 1913. Bugoie forest, 

northeast of Lake Kivu, western border of Tanganyika Territory. 
Simia (Anthropopithecus) kooloo-kamba yamhuyse Matschie, Ann. Soc. Roy. 

Zool. et Malacol. Belgique, Bruxelles, for 1912, 47: 46, Dec. 1913. Yam- 

buya, lower Aruwimi, Congo Beige. 
Anthropopithecus purschei Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 

332, July 1914. Tschingogo forest, between Lakes Kivu and Luhondo, 

western border of Tanganyika Territory. 
Anthropopithecus pfeifferi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 333, 

July 1914. East of Russissi on border of Urundi, near sources of the Akan- 

jaru, western border of Tanganyika Territory. 
Anthropopithecus graueri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 333, 

July 1914. West shore of Lake Tanganyika, 80 km. northwest of Boko, 

Wabembe country, Congo Beige. 
Anthropopithecus calvescens Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 334, July 1914. Baraka-Kasongo road, between Niembo and Kabam- 

bare, on the Luama River, west of Lake Tanganyika, Congo Beige. 
Anthropopithecus castanomale Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 334, July 1914. Northeast shore of Lake Tanganyika, border of Urundi, 

Tanganyika Territory. 
Anthropopithecus schubotzi Matschie, Sitzb. Ges. Naturf. Freunde, Berhn, 

p. 335, July 1914. Between Kilo and Irumu, upper Ituri River, west of 

Lake Albert Edward, Congo Beige. 
Anthropopithecus steindachneri Lorenz, Anzeiger K. Akad. Wiss. Wien, math.- 

nat. Kl., 51 : 550, Dec. 1914. Ituri forest near Moera, north of Beni, Congo 

Beige. 
Range: Forested parts of central Africa, north and east of the Congo, from 

Ubangi River to Lake Albert and Lake Victoria. 

Pan troglodytes verus Schwarz. Western Chimpanzee 

Pan satyrus verus Schwarz, Ann. Mag. Nat. Hist. (10) 13: 578, June 1934. 

Sanda Magbolonto chiefdom, Karima district, Sierra Leone. 
Simia schimpanse Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 67, 1904; 

not Satyrus chimpanse Mayer, 1856. No locality. 



176 bulletin: museum of comparative zoology 

Pan paniscus Schwarz. Lesser Chimpanzee 

Pan satyrus paniscus Schwarz, Revue Zool. Africaine, 16: 425, 1 Apl. 1929. 

South of the upper Maringa River, 30 km. south of Befall, south bank of 

the Congo, Congo Beige. 
Pan paniscus Coolidge, Amer. Journ. Phys. Anthropol., 18: 1-57, Sep. 1933. 

Schwarz (Ann. Mag. Nat. Hist. (10) 13: 582, June 1934) disagrees with the 

view that this is a species distinct from Pan satyrus (Linnaeus). 

Gorilla I. GeoflFroy. Gorillas 

Gorilla I. Geoffroy, Comptes Rend. Acad. Sci. Paris, 34: 84, 1852; 36: 933, 1853. 
Genotype Troglodytes gorilla Savage and Wyman. 

Pseudogorilla EUiot, Review of the Primates, Monographs Amer. Mus. Nat. 
Hist., 3: 224, pi. 33, 1912 ( = 15 June 1913). Genotype Gorilla mayema 
Alix and Bouvier = Gon7Za gorilla (Savage and Wyman). 

Review: Coohdge, H. J., Jr. A revision of the genus Gorilla. Mem. Musi 
Comp. Zool., 50: 281-381, pi. 1-21, maps 1, 2, Aug. 1929. 

Gorilla gorilla gorilla (Savage and Wyman). Coast Gorilla 

Troglodytes gorilla Savage and Wyman, Boston Journ. Nat. Hist., 5: 417, 4 pis., 
Dec. 1847; Wyman, Proc. Boston Soc. Nat. Hist., 2: 245, Aug. 1847. 
Mpongwe, Gaboon River. 

Troglodytes savagei Owen, Proc. Zool. Soc. London, p. 27, 2 text-f., 1 Aug. 1848. 
Mpongwe, Gaboon River. 

Gorilla gina I. Geoffroy, Arch. Mus. d'Hist. Nat., Paris, 8: 1, pi. 2-4, 1855. 
West coast of Africa. 

Satyrus adrotes Mayer, Arch. f. Naturgesch., 22: sect. 1, p. 282, 1856. Substi- 
tute name for Troglodytes gorilla Savage and Wyman. 

Sat[yrus] africanus Mayer, Arch. f. Naturgesch., 22: sect. 1, p. 281, 1856. 
Substitute name for Troglodytes gorilla Savage and Wyman. 

Gorilla castaneiceps Slack, Proc. Acad. Nat. Sci. Philadelphia, p. 159, May 1862. 
[Gaboon?]. 

Pithecus gesilla Blainville, Osteographie, pi. 2, 5 bis, 1839-64. Misprint for 
gorilla, corrected in explan. of pis. 

Gorilla mayema Alix and Bouvier, Bull. Soc. Zool. de France, 2: 488, 1877. 
Banks of the Quilo, near village of King Mayema, 4° 35' S., Congo. 

Gorilla diehli Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 52, 1904. 
Region of the Mun-Aya, which flows into the Cross River, near Dakbe, 
Menome River, 6° 6' N., 9° 20' E., near Cameroons boundary. 

Gorilla gorilla matschiei W. Rothschild, Proc. Zool. Soc. London, for 1904, 2: 
415, text-f. 101, 102, 18 Apl. 1905. Southern Cameroons. 

Gorilla jacobi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 283, 1905. 
Lobo-mouth Station, near junction of the Ja River and the Njong, Came- 
roons. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 177 

Gorilla gorilla schwarzi Matschie in Fritze, Jahrb. d. Provinzial-Mus., Hann- 
over, p. 113, pi. 8, 1912; for description see Auerbach, Verh. d. Naturw. 
Ver. Karlsruhe, 25: Abhandl., p. 5, pi. 1, f. 2; pi. 2, f. 3, 1913. Near Sogema 
farm, Ja River, southern Cameroons. 

Gorilla hansmeyeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 325, July 
1914. Assobam road, between Mensima and Bimba, west of Mokbe, south 
of Dum6 River, southern Cameroons. 

Gorilla zenkeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 325, July 1914. 
Mbiawe on the Lokundje, six hours upstream from Bipindi, Cameroons. 

Gorilla gorilla halli W. Rothschild, Ann. Mag. Nat. Hist. (9) 19: 271, Feb. 1927. 
Punta Mbouda, Spanish Guinea. 
Range: Southeastern Nigeria (Ikom) eastward to the Sanga River to near 
Mayombe, at about 5° S., 13° E., thus wholly to the north and northwest 
of the Congo River. 

Gorilla gorilla beringei Matschie. Mountain Gorilla 

Gorilla beringeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 257, 1903; 
p. 51, 1904. Kirunga, Ya Sabinyo Volcano, Tanganyika Territory. Named 
for Capt. von Beringe; the spelling of the specific name is an obvious 
misprint. 

Gorilla beringei Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 279, 1905. 

Gorilla graueri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 324, July 1914. 
West shore of Lake Tanganyika, 80 km. northwest of Boko, Congo Beige. 

Gorilla beringei mikenensis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 
Stockholm, (2) 58: no. 2, p. 7, pi. 1-4, 1 Sep. 1917. Mt. Mikeno, Virunga 
(or Birunga) Volcanoes, Congo Beige. 

Gorilla gorilla rex-pygmaeorum Schwarz, Revue ZooL. Africaine, 14 : 333, pi. 1 1-13, 
1 Feb. 1927. Luofu, west of Lake Albert Edward, Congo Beige. 

G[orilla] uellensis Schouteden, in Schwarz, Revue Zool. Africaine, 14: 335, foot- 
note, 1 Feb. 1927. Bondo region, Uele district, Congo Beige. In his note 
Schouteden attributes the name to Matschie, who apparently never pub- 
lished it. 
Range: Mountain forests of eastern Congo Beige, from Mulu, 0° 10' S., to 
Baraka on Lake Tanganyika, 4° 19' S. in a narrow strip, nearly limited 
eastward by Kigezi, Uganda, and on the west by Walikale, Congo Beige. 
Schultz, A. H. (On some distinguishing characters of the Mountain Gorilla, 

Journ. Mammalogy, 15: 51-61, Feb. 1934), has pointed out various characters 

of this gorilla, and regards it as sufficiently different to be treated as a species 

distinct from G. gorilla gorilla, of which it is nevertheless the representative 

form in the eastern part of the forest area. 



178 bulletin: museum of comparative zoology 

Family HOMINID^E. Human Race 
Homo Linnseus. Man 

Homo Linnaeus, Syst. Nat., ed. 10, 1: 20, 1758. Genotype Homo sapiens 
Linnaeus. 

Homo sapiens sapiens Linnseus. White Man 

Homo sapiens Linnseus, Syst. Nat., ed. 10, 1: 20, 1758. No locality; five adjec- 
tival names are given for as many varieties, of which europaeus may be 
taken to denote the Nordic race. Type locality taken as Sweden. 

Homo . Congo Pygmy 

Congo Pygmy of Congo forests and its desert race, the Bushman, of South- 
west Africa form a basic species. 

The Bantu and other Negro races of eastern and South Africa probably 
represent hybrid types between the Pygmy of the forest and Asiatic races. 

Order CARNIVORA 

Family URSID^. Bears 
Ursus LinniBus 

Ursus Linnseus, Syst. Nat., ed. 10, 1: 47, 1758. Genotype Ursus arctos Lin- 
nseus. 

Ursus arctos crowtheri Schinz. Crowther's Bear 

Ursus croivtheri Schinz, Syst. Verzeichniss Saugeth., 1: 302, 1844. Atlas 
Mountains near Tetuan, Spanish Morocco. 
On the status of this supposed bear see: Lataste, F., Actes Soc. Linn. Bor- 
deaux (4) 9: 235, 1885; Blyth, E., Proc. Zool. Soc. London, for 1841, p. 65, 
1842; Lydekker, R., Proc. Zool. Soc. London, p. 425, Aug. 1897; G. Busk, 
Trans. Zool. Soc. London, 10: 73, 1877. 

Family MUSTELID.*:. Weasels, etc. 

Subfamily MUSTELINE 

IcTONYX Kaup. African Polecats 

Zorilla Oken, Lehrbuch d. Naturgesch., sect. 3, zool., pt. 2, p. xi, 1816. Geno- 
type Viverra zorilla. (This name is improperly diagnosed and hence un- 
available; see A. H. Howell, The generic name Zorilla, Proc. Biol. Soc. 
Washington, 19: 46, 26 Feb. 1906.) 

Ictonyx Kaup, Das Thierreich, 1: 352, 1835. Genotype Ictonyx capensis 
Kaup = Viverra zorilla Erxleben. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 179 

Rhabdogale Miiller, Arch, f . Naturgesch. for 1838, 1 : 278, footnote, 1838. No 

genotype named, but includes the zorillas of Africa. 
Ictidonyx Agassiz, Nomenclator Zool., Index Univ., p. 194, 1846 (emendation). 
Iclomys Roberts, Aim. Transvaal Mus., 18: 228, 21 Yeh. 1936. Misprint for 

Ictonyx, with I. striahis maximus Roberts. 

ICTONYX KALAHARICUS KALAHARICUS Roberts 

Ictonyx kalaharicus Roberts, Ann. Transvaal Mus., 15: 8, 1 Oct. 1932. Kuke 
Pan, Central Kalahari, Bechuanaland, South Africa. 

Ictonyx kalaharicus giganteus Roberts 

Ictonyx kalaharicus giganteus Roberts, Ann. Transvaal Mus., 15: 9, 1 Oct. 

1932. Shorobe, 25 miles north of Maun, Ngamiland, Bechuanaland, 

South Africa. 

Ictonyx kalaharicus nigricaudus Roberts 

Ictonyx kalaharicus nigricaudus Roberts, Ann. Transvaal Mus., 15: 9, 1 Oct. 
1932. Gemsbok Pan, Bechuanaland, South Africa. 

Ictonyx orangi^e orangi^ Roberts 

Ictonyx orangise Roberts, Ann. Transvaal Mus., 10: 67, 31 Jan. 1924. Angra 
Pequina, south of Bothaville, Orange Free State. 

Ictonyx orangi^ arenarius Roberts 

Ictonyx orangise arenarius Roberts, Ann. Transvaal Mus., 10: 67, 31 Jan. 1924. 
Lambert's Bay, Cape Colony. 

Ictonyx orangi^ ghansiensis Roberts 

Ictonyx orangise ghansiensis Roberts, Ann. Transvaal Mus., 15: 8, 1 Oct. 1932. 
Gemsbok Pan, Bechuanaland, South Africa. 

Ictonyx orangi^ Pretoria Roberts 

Ictonyx orangise pretorise Roberts, Ann. Transvaal Mus., 10: 67, 31 Jan. 1924. 
Bcekenhoutfontein, Pretoria district, Transvaal. 

Ictonyx striatus striatus (Perry) 

Bradypns striatus Geo. Perry, Arcaila or the Museum of Natural History, 

pt. 11, pi. (41) and text, 1810. Cape of Good Hope. (Not Viverra striata 

Shaw, which is a Spilogale, see HoUister, Proc. Biol. Soc. Washington, 28: 

184, 29 Nov. 1915.) 
Mephitis capensis A. Smith, Descriptive Cat. South Air. Mus., p. 20, 1826. 

Cape of Good Hope. 
Mustela zorilla a capensis J. B. Fischer, Synopsis Mamm., p. 218, 1829. Cape 

of Good Hope. 



180 bulletin: museum of comparative zoology 

M[ephitis] ofricana Lichtenstein, Abhandl. K. Akad. Wiss., Berlin, p. 284, 

pi. 1, f. 3, 1836. "Per Africam totam," Cape of Good Hope, Senegambia, 

Abyssinia, Barbary. 
Rhabdogale mmtelina Wagner, Schreber's Saugthiere, Suppl., 2: 219, pi. 133 A, 

1841. Cape of Good Hope. 
Zorilla variegata Lesson, Nouv. Tabl. Regne Anim., Mammif., p. 7, Nov. 1842. 

Cape of Good Hope, Senegal, Abyssinia. 
Zorilla zorilla Trouessart, Cat. Mamm. Viv. Foss., pt. 2, p. 258, 1897. South 

Africa, etc. 

ICTONTX STRIATUS ALBESCENS Heller 

Icionyx capensis albescens Heller, Smithsonian Misc. Coll., 61: no. 13, p. 13, 16 
Sept. 1913. Mt. Lololokwi, northern Guaso Nyiro district, Kenya Colony. 

IcTONYX STRIATUS ELGONis Granvik 

Ictonyx striaius elgonis Granvik, Acta Univ. Lundensis, (2) 21 : no. 3, p. 13, 
1924. Mt. Elgon region, Kenya Colony. 

Ictonyx striatus erythre.e De Winton 

Ictonyx erythrsea (sic) De Winton, Ann. Mag. Nat. Hist. (7) 1 : 248, Mar. 1898. 

Suakin, Red Sea coast, Anglo-Egyptian Sudan. 
Ictonyx erythrese Anderson and De Winton, Zool. of Egypt, Mamm., p. 240, 

pi. 37, 1902 (correction of misprint). 
Range : Eastern Sudan to southeastern Somaliland. 

Ictonyx striatus intermedius Anderson and De Winton 
Ictonyx senegalensis intermedia Anderson and De Winton, Zool. of Egypt, 
Mamm., p. 242, 1902. "Nile basin." 

Ictonyx striatus lancasteri Roberts 

Ictonyx striatus lancasteri Roberts, Ann. Transvaal Mus., 15: 8, 1 Oct. 1932. 
Choma, Northern Rhodesia. 

Ictonyx striatus limpopoensis Roberts 

Ictonyx capensis limpopoensis Roberts, Ann. Transvaal Mus., 5: 265, 16 May 
1917. Mooivlei, Rustenburg district, Transvaal. 

IcTONiTC STRIATUS MAXiMUS Roberts 

Ictonyx striatus maximus Roberts, Ann. Transvaal Mus., 10: 65, 31 Jan. 1924. 
Wakkerstroom, South Africa. 

Ictonyx striatus obscuratus de Beaux 

Ictonyx capensis obscurata de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. 
Nat. Milano, 63: 1, July 1924. Lugh, Italian Somaliland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 181 

ICTONYX STRIATtJS PONDOENSIS RobcrtS 

Ictonyx striatus pondoensis Roberts, Ann. Transvaal Mus., 10: 66, 31 Jan. 1924. 
Port St. John's, South Africa. 

Ictonyx striatus senegalensis (J. B. Fischer) 

Mustela zorilla p. senegalensis J. B. Fischer, Synopsis Mamm., p. 219, 1829. 

Senegal. 
Zorilla striata var. senegalensis Gray, Proc. Zool. Soc. London, p. 151, June 

1865. Senegal. See De Winton, W. E., Ann. Mag. Nat. Hist. (7) 1 : 248, 

Mar. 1898. 

Ictonyx striatus sho^ Thomas 

Ictonyx capensis shose Thomas, Ann. Mag. Nat. Hist. (7) 18: 301, Oct. 1906j 
Addis Ababa, Ethiopia, 8500 ft. 

Ictonyx striatus shortridgei Roberts 

Ictonyx striatus shortridgei Roberts, Ann. Transvaal Mus., 15: 8, 1 Oct. 1932. 
Maschi River, Caprivi border. Southwest Africa. 

Ictonyx striatus sudanicus Thomas and Hinton 

Ictonyx striatus sudanicus Thomas and Hinton, Proc. Zool. Soc. London, p. 

254, 6 July 1923. Foothills of Jebel Marra, Darfur, Anglo-Egyptian 

Sudan, 4000 ft. , 

Mellivora Storr. Ratels or Honey Badgers 

Mellivora Storr, Prodromus Methodi Mamm., p. 34, tab. A, 1780. Genotype 

Viverra ratel Sparrman = Viverra capensis Schreber. 
Ratelus Bennett, Gardens and Menagerie Zool. Soc, 1: Quad., 13, text-f., 

1830. Genotype Ratelus mellivorus Bennett. 
Ursitaxus Hodgson, Journ. Asiatic Soc. Bengal, (2) 4: 522, Sep. 1835. Geno- 

tj^e Ursitaxus inauritus Hodgson. 
Melitoryx Gloger, Gemeinniitz. Hand- u. Hilfsbuch Naturg., 1: pp. xxix, 57, 

1841. Genotype honey badger of South Africa and India. 
Lipotus Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 1842, 

p. 199, 1843. Genotype Ursus mellivorus Cuvier. 

Mellivora capensis capensis (Schreber) 

Viverra capensis Schreber, Saugthiere, 3: 451, 588, pi. 125, 1776. Cape of Good 

Hope. See HoUister, Proc. Biol. Soc. Washington, 25: 93, 1912. 
Viverra ratel Sparrman, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 38: 

147, 1777. Cape of Good Hope. 
C/rsMs mellivorus G. Cuvier, Tabl. Element. Hist. Nat. des Anim., p. 112, 

1798. Cape of Good Hope. 
Ratellus typicus A. Smith, So. African Quart. Journ., 2: 83, 1833. "South 

Africa." 



182 bulletin: museum of comparative zoology 

Mellivora capensis abyssinica Hollister 

Mellivora abyssinica Hollister, Smithsonian Misc. Coll., 56: no. 13, p. 1, 10 
Oct. 1910. Near Addis Ababa, Ethiopia; in later report corrected to 
Suksukki River, which connects Lake Zwai with Lake Horo Schala, 4500- 
5000 feet. 

Mellivora capensis brockmani Wroughton and Cheesman 
Mellivora brockmani Wroughton and Cheesman, Ann. Mag. Nat. Hist. (9) 5: 
197, Feb. 1920. Upper Sheikh, northern Somaliland. 

Mellivora capensis buchanani Thomas 

Mellivora buchanani Thomas, Ann. Mag. Nat. Hist. (9) 16: 190, July 1925. 
Elmiki River, Asben (Air), Sahara, French West Africa. 

Mellivora capensis concisa Thomas and Wroughton 

Mellivora concisa Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 376, 
May 1907. Yo, Lake Chad. 

Mellivora capensis cottoni Lydekker 

Mellivora cottoni Lydekker, Froc. Zool. Soc. London, 1: 112, pi. 7, 7 June 1906. 

Eastern fringe of the Ituri forest, near Mawambi, Congo Beige. Probably 

a melano. 

Mellivora capensis leuconota P. L. Sclater 

Mellivora leuconota P. L. Sclater, Proc. Zool. Soc. London, p. 98, pi. 8, May 
1867. "West Africa." 
Range: Southern Morocco south to French Congo. 

Mellivora capensis maxwelli Thomas 

Mellivora maxwelli Thomas, Ann. Mag. Nat. Hist. (9) 12: 340, Sep. 1923. 
Lorian Swamp, Kenya Colony. 

Mellivora capensis sagulata Hollister 

Mellivora sagulata Hollister, Smithsonian Misc. Coll., 56: no. 13, p. 2, 10 Oct. 
1910. Mt. Kilimanjaro, Tanganyika Territory, 5000 feet. 
For commentary on supposed size characters see Lonnberg, E., Der Honig- 
dachs vom Kilimandscharo, Zool. Anzeiger, 37: 74-75, 31 Jan. 1911. 

Mellivora capensis signata Pocock 

Mellivora signata Pocock, Proc. Zool. Soc. London, p. 394, pi. 61, 23 Aug. 
1909. About 100 miles inland from Sierra Leone. 

Mellivora capensis vernayi Roberts 

Mellivora capensis vernayi Roberts, Ann. Transvaal Mus., 15: 7, 1 Oct. 1932. 
Kwai, Mababe Flats, northern Bechuanaland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 183 



MusTELA Linnaeus. Weasels and Stoats 

Mustela Linnaeus, Syst. Nat., ed. 10, 1: 45, 1758. Genotype (nine species 

originally included) Mustela erminea Linnaeus. 
Ardogale Kaup, Entw.-Gesch. u. Naturl. Syst. Europ. Thierwelt, 1: 30, 1829. 

Genotype Mustela erminea Linnaeus. 
Gale Wagner, Schreber's Saugthiere, Suppl., 2: 234, 1841. As a subgenus of 

Mustela, to include M. frenata Lichtenstein, M. erminea Linnaeus, M. 

boccamela Bechstein, M. vulgaris Erxleben. 
Neogale Gray, Proc. Zool. Soc. London, p. 114, June 1865. As a subgenus, 

type not specified; included Mustela brasiliensis Sevastianoff, M. aureo- 

ventris Gray, M. xanthogenys Gray. 
Gymnopus Gray, Proc. Zool. Soc. London, p. 120, June 1865. Genotype 

Mustela africana Desmarest. 
Mustelina Bogdanov, Proc. Imp. Univ. Kazan, 1: 167, 1871. Genotype ? 
Eumustela Acloque, Faune de France, Mamm., p. 62, 1900. As a section of 

Mustela, to include M. vulgaris Linnaeus and M. erminea Linnaeus; no 

type designated. 
See Barrett-Hamilton, G. E. H. Note on the weasel, Putorius (Idis) 
nivalis, Linn., and some of its subspecies. Ann. Mag. Nat. Hist., (7) 5: 41-50, 
Jan. 1900. 

Mustela nivalis subpalmata (Hemprich and Ehrenberg) 
Mustela subpalmata Hemprich and Ehrenberg, Symbolae Physicae, Mamm., 
dec. 2, folio k, p. 2, Sep. 1832. In houses, Cairo and Alexandria, Egypt. 
Status: see Thomas, O., Proc. Zool. Soc. London, p. 128-131, June 1895. 

Mustela numidica (Pucheran) 

Putorius numidicus Pucheran, Rev. et Mag. de Zool., 1855, p. 393. Morocco 

and Algeria; (fixed as Tangiers by Cabrera). 
Putorius nivalis atlas Barrett-Hamilton, Ann. Mag. Nat. Hist. (7) 13: 323, 

Apl. 1904. Atlas Mountains of Morocco. 
Putorius nivalis siculus Barrett-Hamilton, Ann. Mag. Nat. Hist. (7) 5: 46, 

Jan. 1900 (in part). Marsala, Sicily. 
Putorius ermineus algiricus Thomas, Ann. Mag. Nat. Hist. (6) 15: 451, May 

1895. Near Algiers, Algeria. 
Mustela erminea, var. 1, africana^ Gray, Proc. Zool. Soc. London, p. Ill, 

June 1865 (not of Desmarest). Algiers. 
Range: Sicily and Algeria (to Oran Province). 

Note: Mustela africana Desmarest (Nouv. Dictionnaire d'Hist. Nat., 19: 
376, 1818) said to be "d'Afrique," is shown to have been based on a species 
from South America by Cabrera (Bol. Soc. Real Espanola. Hist. Nat., Mad- 
rid, 13: 428, 1913; 14: 176, pi. 1, 1914), 



184 bulletin: museum of comparative zoology 



Subgenus Putorius Cuvier. Polecats and Ferrets 

Putorius Cuvier, Regne Animale, ed. 1, 1: 147, 1817. As a subgenus, to in- 
clude eight species; type Mustela putorius Linnseus. 

FcElorius Keyserling and Blasius, Wirbelthiere Europa's, p. 68, 1840. Geno- 
type not indicated; seven species included. 

Cynomyonax Coues, U. S. Geol. Surv. Terr., Misc. Publ. no. 8, "Fur-bearing 
Animals," p. 99, 1877. As a subgenus of Putorius; type Putorius nigripes 
Audubon and Bachman. 

Mustela putorius furo Linnseus. The Ferret 

Mustela furo Linnseus, Syst. Nat., ed. 10, 1: 46, 1758. "Africa." 

Putorius foetidus var. subfuro Gray, Proc. Zool. Soc. London, p. 110, June 1865. 

Bred in captivity. 

Putorius putorius furo Pocock, Proc. Zool. Soc. London, p. 692, 25 Sept. 1936. 

Range: Mountains of the extreme west of the.Rif, from Nekor River to 

Gomara in east of Yebala. Now reared and introduced to Europe and 

elsewhere (see Cabrera, A., in Los mamiferos de Marruecos, p. 136, 1932.) 



P(ecilictis Thomas and Hinton 

PoEcilictis Thomas and Hinton, Ann. Mag. Nat. Hist. (9) 5: 367, Apl. 1920. 
Genotype Mustela libyca Hemprich and Ehrenberg. 

Review: Thomas, O., and Hinton, M.A.C. On the group of African zorils 
represented by Ictonyx libyca. Ann. Mag. Nat. Hist. (9) 5: 367-369, Apl. 1920. 

PcECiLiCTis LIBYCA LIBYCA (Hcmprich and Ehrenberg) 

Mustela libyca Hemprich and Ehrenberg, Symbolse Physicse, Mamm., dec. 2, 
foho k, p. 2, Sep. 1832. Libya, northern Africa. 
Range: Lower Egypt. 

PCECILICTIS LIBYCA MULTIVITTATA (Wagner) 

Rhabdogale multivittata Wagner, Schreber's Saugthiere, Suppl., 2: 221, foot- 
note, pi. 133B, 1841. Upper Nile (Pruner, coll.). 

Ictonyx frenata Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 
for 1842, p. 212, pi. 4, f. 1, 1843. Sennaar. 

PcECiLiCTis LIBYCA ORALIS Thomas and Hinton 

Pcecilictis oralis Thomas and Hinton, Ann. Mag. Nat. Hist. (9) 5: 368, Apl. 

1920. Suakin, Egypt. 
Mephitis africana, var. /?. Lichtenstein, Abh. K. Akad. Wiss., Berlin, p. 284, 

pi. 1, f. 3, 1836. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 185 

PcECiLiCTis LiBYCA ROTHscHiLDi Thomas and Hinton 

Pcecilictis rothschildi Thomas and Hinton, Novitates Zool., 27: 316, 15 June 
1920. Famiso, near Kano, northern Nigeria. 

PCECILICTIS LIBYCA VAILLANTII (Loche) 

Zorilla vaillantii Loche, Rev. et Mag. de Zool., (2) 8: 497, pi. 22, Nov. 1856. 
Algeria. 
Range: Algeria and Tunis. 

PcECiLOGALE Thomas 

Pcecilogale Thomas, Ann. Mag. Nat. Hist. (5) 11: 370, text-fig., 1 May 1883. 
Genotype Zorilla albinucha Gray. 

PcECILOGALE ALBINUCHA ALBINUCHA (Gray) 

Zorilla albinucha Gray, Proc. Zool. Soc. London, p. 69, pi. 10, July 1864. 
Locality unknown, but Cape of Good Hope may be assumed. 

PCECILOGALE ALBINUCHA BECHUAN^ Roberts 

PcEcilogale albinucha bechuanse Roberts, Ann. Transvaal Mus., 14: 226, 28 July 
1931. Vryburg, South Africa. 

PcECiLOGALE ALBINUCHA DOGGETTi Thomas and Schwann 

Pcecilogale doggeiti Thomas and Schwann, Abstr. Proc. Zool. Soc. London, 

no. 6, p. 22, 26 Apl. 1904; Proc. Zool. Soc. London, 1: 460, 2 Aug. 1904. 

Burumba, Ankole, southwestern Uganda. 

Pcecilogale albinucha flavistriata (Bocage) 

Zorilla flavistriata Bocage, Proc. Zool. Soc. London, p. 402, Oct. 1865. Duque 
de Braganca district, Angola. (Proposed on the basis of Angola specimens, 
as a substitute for albinucha, but may represent a distinct race.) 

Zorilla africana Peters, Proc. Zool. Soc. London, p. 400, Oct. 1865 (nomen 
nudum). Golungo Alto, Angola. 

Pcecilogale albinucha lebombo Roberts 

Pcecilogale albinucha lebombo Roberts, Ann. Transvaal Mus., 14: 226, 28 July 
1931. Ubombo, northern Zululand, South Africa. 

Pcecilogale albinucha transvaalensis Roberts 

Pcecilogale albinucha transvaalensis Roberts, Ann. Transvaal Mus., 11: 247, 14 
Sep. 1926. Tzaneen, northeastern Transvaal. 

Subfamily Luteins. Otters 

Review: Pohle, H. Die Unterfamilie der Lutrinte. (Eine systematisch- 
tiergeographische Studie an dem Material der Berliner Museen.) Arch. f. 
Naturgesch., 85: sect. A, no. 9, p. 1-246, 19 text-figs., 10 pis., Dec. 1920. 



186 bulletin: museum of comparative zoology 



AoNYX Lesson. Clawless Otters 

Aonyx Lesson, Manuel de Mammalogie, p. 157, 1827. Genotype Aonyx 

delalandi hesson = Lutra capensis Schinz. 
Anahyster Murray, Proc. Roy. Phys. Soc. Edinburgh, 2: 157, 1860. Genotype 

Anahyster calabaricus Murrsiy = Aonyx capensis (Schinz). 

AoNirx CAPENSIS CAPENSIS (Schinz) 

Lutra capensis Schinz, Cuvier's Thierreich, 1: 214, 1821. Cape Colony. 

Lutra inunguis G. Cuvier, Dictionnaire des Sci. Nat., 27: 247, 1823. Cape of 

Good Hope. 
Aonyx delalandi Lesson, Manuel de Mammalogie, p. 157, 1827. Cape of Good 

Hope. 
Lutra poensis Waterhouse, Proc. Zool. Soc. London, p. 60, July 1838. Fernando 

Po. 
Anahyster calabaricus Murray, Proc. Roy. Phys. Soc. Edinburgh, 2: 158, 1860. 

Old Calabar, West Africa. 
hutra lenoiri Rochebrune, Vert. Nouv. Afr. Occid., ser. 3, p. 9, 1888 (fide 

Trouessart, Cat. Mamm. Viv. Foss., ed. 1, pt. 2, p. 285, 1897). Senegal. 

Aonyx capensis angol.e Thomas 

Aonyx capensis angolse Thomas, Ann. Mag. Nat. Hist. (8) 1 : 388, May 1908. 
Coporole River, Angola, lat. 13° S. 

Aonyx capensis coombsi Roberts 

Aonyx capensis coombsi Roberts, Ann. Transvaal Mus., 11: 246, 14 Sep. 1926. 
Hennops River, Pretoria, Transvaal. 
For Cecil Coombs, coll. 

Aonyx capensis helios Heller 

Aonyx capensis helios Heller, Smithsonian Misc. Coll., 61: no. 19, p. 1, 8 Nov. 
1913. Sotik district, 40 miles southwest of Kericho, Kenya Colony. 

Aonyx capensis hindei (Thomas) 

Lutra capensis hindei Thomas, Ann. Mag. Nat. Hist. (7) 15: 78, Jan. 1905. 
Fort Hall, Kenya Colony. 

Aonyx capensis meneleki (Thomas) 

Lutra capensis meneleki Thomas, Proc. Zool. Soc. London, for 1902, 2: 309, 
1 Apl. 1903. Zegi, Lake Tsana, Abyssinia, 4000 ft. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 187 



LuTRA Brisson. River Otters 

Jjutra Brisson, Regnum Animale in classes IX distrib., ed. 2, 13: 201, 1762. 

Genotype Lutra lutra Brisson = MitsteZa Intra Linnseus. 
Hydrogale Gray, Proc. Zool. Soc. London, p. 131, fig. of skull, June 1865 (not 

Hydrogale Kaup, 1829). Genotype Lutra maculicollis Lichtenstein. 

Subgenus Lutra Brisson 

Lutra lutra angustifrons Lataste 

Lutra angustifrons Lataste, Actes Soc. Linn, de Bordeaux, (4) 9: 168, 237, 1885. 

Algeria. 
Lutra lutra s-plendida Cabrera, Bol. Real Soc. Espan. Hist. Nat., Madrid, 6: 

360, July 1906. Mogador, Morocco. 

Subgenus Hydrictis Pocock 

Hydrictis Pocock, Proc. Zool. Soc. London, p. 543, 9 Sep. 1921. Genotype 
Lutra maculicollis Lichtenstein. Although Pocock regards this as a dis- 
tinct genus, it may better be retained as a subgenus. 

Lutra maculicollis maculicollis Lichtenstein 

Lutra maculicollis Lichtenstein, Arch. f. Naturgesch., for 1835, 1: 89, pi. 2, f. 1, 

1835. "Aus dem Kafferlande," i.e.. South Africa. 
Lutra grayii Gerrard, Cat. Bones of Mammalia in Brit. Mus., p. 101, 1862 

(nomen nudum). Port Natal, South Africa. 

Lutra maculicollis chobiensis Roberts 

Lutra malculicollis (sic) chobiensis Roberts, Ann. Transvaal Mus., 15: 7, 1 Oct. 
1932. Kabulabula, Chobe River, Bechuanaland, South Africa. 

Lutra maculicollis kivuana Pohle 

Lutra maculicollis kivuana Pohle, Arch. f. Naturgesch., for 1919, 85: sect. A, 
no. 9, p. 46, 50, f. 5, Dec. 1920. Kissenji, Lake Kivu, Tanganyika Territory, 

Lutra maculicollis matschiei Cabrera 

Lutra matschiei Cabrera, Bol. Soc. Espafiola Hist. Nat., Madrid, 3: 182, Apl. 

1903. Muni River, northern Gaboon. 
Lutra maculicollis matschiei Thomas", Ann. Mag. Nat. Hist. (8) 8: 726, Dec. 

1911; "unrecognizable from the description" (J.A.Allen) probably =L. 

nmculicollis. 

Lutra maculicollis nilotica Thomas 

Lutra concolor Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 55, 1902 (pre- 
occupied by Lutra concolor Rafinesque, 1832, from India). Addis Ababa, 
Ethiopia. 



188 bulletin: museum of comparative zoology 

Lutra macuUcollis nilotica Thomas, Ann. Mag. Nat. Hist. (8) 8: 726, Dec. 1911 
Upper Nile, Malek, south of Bor, Anglo-Egyptian Sudan. 

Lutra maculicollis tenuis Pohle 

Lutra tenuis Pohle, Arch. f. Naturgesch., for 1919, 85: sect. A, no. 9, p. 53, 

Dec. 1920. Lake Mohasi, Ruanda district, eastern Congo Beige. 
Lutra maculicollis viutandse Hinton, Ann. Mag. Nat. Hist. (9) 7: 368, Apl. 1921. 

Lake Mutanda, British Ruanda. 
Range : British Ruanda into Uganda. 



Paraonyx Hinton 

Paraonyx Hinton, Ann. Mag. Nat. Hist. (9) 7: 195, Feb. 1921. Genotype Para- 
onyx philippsi Hinton. 

Paraonyx congica (Lonnberg) 

Aonyx capensis congica Lonnberg, Arkiv f. Zool., 7: no. 9, p. 1, text-f. la, 2a, 

15 Dec. 1910. "Lower Congo." 
Paraonyx congica Hinton, Ann. Mag. Nat. Hist. (9) 7: 198, Feb. 1921. 

Paraonyx microdon (Pohle) 

Aonyx microdon Pohle, Arch. f. Naturgesch., 85: sect. A, pt. 9, p. 145, Dec. 
1920. Bomse, Nana River, Cameroons. 

Paraonyx philippsi Hinton 

Paraonyx philippsi Hinton, Ann. Mag. Nat. Hist. (9) 7: 196, Feb. 1921. South- 
western Lake Bunyonyi, British Ruanda, Central Africa, lat. 1° 20' S., 
long. 29° 50' E. 



Family CANID.E. Dogs, Jackals, Foxes 

See De Winton, W. E. On the species of Canidae found on the continent of 
Africa. Proc. Zool. Soc. London, p. 533-552, 13 text-figs., Oct. 1899. 



Fennecus Desmarest. Fennec 

Fennecus Desmarest, Dictionnaire d'Hist. Nat., ed. 1, 24: tabl. methodique d. 

mamm., p. 18, 1804; Nouv. Dictionnaire d'Hist. Nat., 11: 341, 1817. 

Genotype Fennecus arabicus Desmarest = Canis zerda Zimmermann. 
Megalotis IWiger, Prodromus Syst. Mamm. et Avium, p. 131, 1811. Genotype 

Canis cerdo Gmelin = C. zerda Zimmermann. 
Canis, Megalotis, Viverra, Vulpes, in part, of older authors. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 189 

Fennecus zerda (Zimmermann) 

Vulpes miimnus saarensis Skjoldebrand, Kongl. Svenska Vet.-Akad. Handl., 

Stockholm, 38: 267, pi. 6, 1777. Algerian Sahara, on the farther side of 

the Atlas Mts. 
This name if considered valid, would supersede Cards zerda, but although the 
author states that he wishes to include the animal in the Linntean system, he 
gives a trinomial name. 
Canis zerda Zimmermann, Geograph. Geschichte, 2: 247, 1780. Sahara and 

other regions back of the Atlas Mts., and in TripoH. 
Canis cerdo Gmelin, Linnseus's Syst. Nat., ed. 13, 1 : 75, 1788. Desert of Sahara. 
Viverra aurita F. A. A. Meyer, Syst.-summarisch. Uebersicht d. Entdeckungen 

in Neu-HoUand u. Africa, p. 91, 1793. Biskra, Beni Mezzab and Weryleh, 

Algeria. 
Fennecus arabicus Desmarest, Dictionnaire d' Hist. Nat., ed. 1, 24: tabl. 

methodique d. mamm., p. 18, 1804; Nouv. Dictionnaire d'Hist. Nat., 11: 

342, 1817. Barbary, Nubia, Abyssinia. 
Fennecus briicei Desmarest, Encyclop. Methodique, Mammalogie, p. 235, 1820. 

Libya, Tunis, Algeria and Sennaar. 
Canis fennecus Lesson, Manuel de Mammalogie, p. 168, 1827. "I'Afrique cen- 

trale." 
Vulpes denhand Boitard, Le Jardin des Plantes, p. 163, 1842. "L'interieur de 

I'Afrique." 
Vulpes zaarensis Gray, List. Spec. Mamm. Brit. Mus., p. 62, 13 May 1843. 

Egypt. 
Megalotis fennecus Knight, Pict. Mus. Anim. Nature, p. 207, circa 1860. 

Lycaon Brookes. Hunting Dog 

Lycaon Brookes, in Griffith's Cuvier's Animal Kingdom, 5: 151, 1827. Geno- 
type Lycaon tricolor Brookes = Hysena picta Temminck. 

Cynhysena F. Cuvier, Dictionnaire des Sci. Nat., 59: 454, 1829. Genotype 
Hysena picta Temminck. 

Hyenoides Boitard, Le Jardin des Plantes, p. 163, 1842. Genotype Hyaena picta 
Temminck. 

Hysenoides Gervais, Hist. Nat. des Mammiferes, 2: 53, 1855 (variant of Hyen- 
oides) . 

Lycaon pictus pictus (Temminck) ^ 

Hysena picta Temminck, Ann. Gen. Sci. Phys., 3: 54, pi. 35, 1820. Coast of 

Mozambique. 
Canis (Lycaon) tricolor Brookes, in Griffith's Cuvier's Animal Kingdom, 5: 151, 

1827. Cape of Good Hope. 
Lycaon typicus A. Smith, South African Quart. Journ., 2: 91, 1833. 
Lycaon windhorni Matschie, Sitzb. Ges. Naturf . Freunde, Berlin p. 378, Oct. 

1915. Rustenburg, Transvaal. 



190 bulletin: museum of comparative zoology 

Lycaon pictus ltjpinus Thomas 

Lycaon pictus Iwpinus Thomas, Ann. Mag. Nat. Hist. (7) 9: 439, June 1902. 

Nyuki River Swamp, Rift Valley, Kenya Colony, 2000 m. 
Lycaon pictus prageri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 250, 

Apl. 1912; p. 311, May 1912. Nzoia River, Uasin Gishu plateau south 

of Mt. Elgon, Kenya Colony. 
Lycaon lademanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 315, Oct. 

1915. Mbarangandu, near Ssongea-Liwale Road, Tanganyika Territory. 
Lycaon ssongese Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 317, Oct. 

1915. Ussangire, 14 km. west-southwest of Ssongea, headwaters of the 

Rovuma, Tanganyika Territory. 
Lycaon hennigi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 320, Oct. 

1915. Makangaga, 40 km. west of Kilwa, Tanganyika Territory. 
Lycaon stierlingi Matschie, Sitzb. Ges. Naturf. Freunde, BerHn, p. 323, Oct. 

1915. Upper Rijuni, Ssongea district, Tanganyika Territory (about 30 

km. from source of the Mbarangandu-Luwegu) . 
Lycaon huebneri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 326, Oct. 

1915. Kibwesi, Kenya Colony. 
Lycaon ruwanx Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 328, Oct. 

1915. Ruwana Plains, northeast of Speke Gulf, Tanganyika Territory. 
Lycaon kondose Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 331, Oct. 1915- 

Bubu, near Kondoa-Irangi, Tanganyika Territory. 
Lycaon langheldi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 333, Oct. 

1915. Njawa, Ussongo, northeast of Tabora, in the western Wembsere 

Plains, Tanganyika Territory. 
Lycaon dieseneri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 336, Oct. 

1915. Myombo forest, between Usinsa and East Ussuwi, south of Vic- 
toria Nyanza, Tanganyika Territory. 
Lycaon gansseri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 340, Oct. 

1915. Tabora district, Tanganyika Territory. 
Lycaon taborse Matschie, Sitzb. Ges. Naturf. Freunde, BerUn, p. 344, Oct. 1915. 

Tabora district, Tanganyika Territory. 
Lycaon wintgensi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 347, Oct. 

1915. Luegere River, Kungue Bay, east shore of Lake Tanganyika, Tan- 
ganyika Territory. 
Lycaon richteri Matschie, Sitzb. Ges. Naturf. Freunde, Berhn, p. 350, Oct. 1915. 

Dar-es-Salaam, Tanganyika Territory. 
Lycaon styxi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 352, Oct. 1915. 

Mpwapwa, Tanganyika Territory. 

Lycaon picttjs manguensis Matschie 

Lycaon manguensis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 364, 

Oct. 1915. Djannaga, north of Sansanne Mangu, near the Oti, (Togo) 

northern Dahomey. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 191 

Lycaon mischlichi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 366, Oct. 
1915. Northern part of Kete-Kratschi district, probably Bimbila, be- 
tween the Oti and Daka, (Togo) northern Dahomey. 

Lycaon pictus sharicus Thomas and Wroughton 

Lycaon pictus sharicus Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 

375, May 1907. Mani, lower Shari River, east of Lake Chad. 
Lycaon ebermaieri Matschie, Sitzb. Ges. Naturf. Freunde, BerUn, p. 369, Oct. 

1915. Lake Chad region (probably from near Dikoa). 
According to Heim de Balsac (Suppl. au Bull. Biol, de France etde Belgique, 
Paris, 21 : 392-393, 1936) specimens from Tanezrouft, Algeria, have larger buUse 
and may indicate that the form of the southern and central Sahara is a distinct 
race, to which possibly the above name is applicable. 

Lycaon pictus somalicus Thomas 

Lycaon pictus somalicus Thomas, Ann. Mag. Nat. Hist. (7) 14: 98, Aug. 1904, 

Gorahai, Somaliland. 
Lycaon riippelli Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 315, Oct. 

1915. Bajuda Steppe region, Anglo-Egj^ptian Sudan (based on Riippell's 

plate 12). 
Lycaon luchsingeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 355, Oct. 

1915. Edaballa, Mulu, near Hawash River, Ethiopia. 
Lycaon zedlitzi Matschie, Sitzb. Ges. Naturf. Freunde, BerUn, p. 358, Oct. 1915. 

Scetel River, on plains southeast of Agordat, Eritrea. 
hycaon takanus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 360, Oct. 

1915. Near Khor el Gasch, Taka district, Anglo-Egyptian Sudan. 

Lycaon pictus venaticus (Burchell) 

Hyaena venatica Burchell, Travels in Interior So. Africa, 1: 456, 1822; 2: 229, 

1824. Northeast of Asbestos Range, upper Orange River. ^ 
Lycaon pictus venaticus Thomas, Ann. Mag. Nat. Hist. (7) 9: 438, May 1902. 
Lycaon lalandei Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 310, Oct. 

1915. Between Cape Town and Keiskama River, South Africa. 
Lycaon fuchsi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 371, Oct. 1915. 

Rio Cubal, Benguella, Angola. 
Lycaon gobabis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 373, Oct. 1915. 

Gobabis, Southwest Africa. 
Lycaon krebsi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 376, Oct. 1915. 

Near Baviaans Kloof, Cape Colony. 

Lycaon pictus zuluensis Thomas 

Lycaon pictus zuluensis Thomas, Ann. Mag. Nat. Hist. (7) 14: 98, footnote, 
Aug. 1904. Pongola River, Zululand, Natal. 



192 bulletin: museum of comparative zoology 



Otocyon Miiller. Big-eared Foxes 

Otocyon Miiller, Jahresber. iiber die Fortschritte d. anat.-phya. Wiss., 1835, 
p. 1 (Roman numeral), with Arch. f. Anat. u. Phys., 1836. Genotype 
Otocyon coffer Lichtenstein = Cams megalotis Desmarest. 

Agriodus Hamilton Smith, Jardine's Naturalist's Library, Mamm., 10: 258, 
1840. As a subgenus of Canis; type Agriodus auritus H. Smith = Cams 
megalotis Desmarest. 

Otocyon megalotis megalotis (Desmarest) 

Canis megalotis Desmarest, Encyclop. M^thodique, Mammalogie, suppl., 

p. 538, 1821. Cape of Good Hope. 
Canis lalandi Desmoulins, Dictionnaire Class. d'Hist. Nat., 4: 18, pi. 15, f. 2, 

1823. "Cafrerie,"= South Africa. 
Otocyon caffer Wiegmann, Arch. f. Naturgesch., 4: sect. 1, p. 290, 1838. No 

locality. 
Agriodus auritus Hamilton Smith, Jardine's Naturalist's Library, Mamm., 10: 

260, pi. 23, 1840. Cape of Good Hope. 

Otocyon megalotis canescens Cabrera 

Otocyon canescens Cabrera, Ann. Mag. Nat. Hist. (8) 6: 462, Nov. 1910. Ten 
miles south of Burao, Somaliland. 
Range: Interior plateau of Somaliland. 

Otocyon megalotis steinhardti Zukowsky 

Otocyon steinhardti Zukowsky, Arch. f. Naturgesch., 90: sect. A, pt. l,p. 52, 3 

text-ff., July 1924. Goreis, Ugab Mts., 45 km. west of Outjo, Southwest 

Africa. 

Otocyon megalotis virgatus Miller 

Otocyon virgatus Miller, Smithsonian Misc. Coll., 52: 485, pi. 60-62, 18 Dec. 
1909. Naivasha Station, Kenya Colony. 



SiMENiA Gray. Simenian Fox 

Simenia Gray, Proc. Zool. Soc. London, for 1868, pp. 494, 506, Mar. 1869. 
Genotype Canis simensis Riippell. 

Simenia simensis simensis (Riippell) 

Cams simensis Riippell, NeueWirbelth. zu Fauna Abyssinien gehorig, Saugeth., 

1 : 39, pi. 14, 1835. Mountains of Simen, Ethiopia. 
Canis sinus Gervais, Hist. Nat. des Mammiferes, 2: 58, 1855. 
Canis walgie Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Dresden, 30: no. 2, 

suppl., p. 3, 1863. Simeen, Ethiopia, above 10,000 feet. 
Canis semiensis Heuglin, Reise in Nordost-Afrika, 2: 48, 1877. Semi6n, 

Vogara, south to Wolo-Gala, Ethiopia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 193 

SiMENiA siMENsis ciTERNii (de Beaux) 

Canis (Simenia) simensis citernii de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. 
Civ. Stor. Nat., Milano, 61 : 25, Feb. 1922. Barofa, Arussi, Ethiopia. 

Thos Oken. Jackals 

On the characters of Thos, see Heller, E., Smithsonian Misc. Coll., 63: no. 7, 
p. 1-3, 24 June 1914. 

Thos Oken, Lehrbuch d. Naturgesch., zool., sect. 3, pt. 2, p. 1037, 1816. Type 

by subsequent designation, Thos vulgaris Oken = Canis aureus Linnaeus. 
Vidpicanis Blainville, Ann. des Sci. Nat., Zool., (2) 8: 279, 1837. Genotype 

Canis aureus Linnseus. 
According to Cabrera (Trabajos Mus. Nac. de Ciencias Nat., Madrid, 
zool. ser., no. 57, p. 106, 30 Dec. 1932) this name should replace Thos, which 
he regards as invalid on the ground that Oken is not binomial. 
Sacalius Hamilton Smith, Jardine's Naturalist's Libr., Mamm., 9: 206, 1839. 

As a subgenus, type Canis aureus Linnseus. 
Oxygous Hodgson, Calcutta Journ. Nat. Hist., 2: 213, July 1841. Genotype 

Oxygoils indicus Hodgson = Canis pallipes Sykes. 
Dieba Gray, Cat. Carniv., Pachyderm, and Edentate Mamm. Brit. Mus., p. 

180, 189, 1869. Genotype Dieba anthus = Canis anthus F. Cuvier. 
Lupulella Hilzheimer, Zool. Beobachter, 47: 363, Dec. 1906. Genotype Thos 

mesomelas (Schreber). 
Schseffia Hilzheimer, Zool. Beobachter, 47: 364, Dec. 1906. Subgenus of Thos 

to include Canis adustus, C. lateralis, C. kaffensis. 
Alopedon Hilzheimer, Zool. Beobachter, 47: 365, Dec. 1906. As a subgenus, 

type Canis thooides Hilzheimer = Cam's anthus Cretzschmar, not of 

Cuvier. 

Thos adustus adustus (Sundevall). Side-striped Jackal 

Canis adustus Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. Forhandl., 

Stockholm, 3: no. 4, p. 121, 1846. In Caffraria interiore. 
Canis holubi Lorenz, Verh. K. K. Zool.-Bot. Ges. Wien, 45: 111, 1895. Leshumo 

Valley, near Victoria Falls, western Matabeleland, Rhodesia. 
Canis wunderlichi Noack, Zool. Anzeiger, 20: 519, 27 Dec. 1897. Probably 

southeastern Africa. 

Thos adustus bweha Heller 

Thos adustus bweha Heller, Smithsonian Misc. Coll., 63: no. 7, p. 2, 3, 24 June 
1914. Kisumu, Kenya Colony. 

Thos adustus centralis (Schwarz) 

Canis adustus centralis Schwarz, Jahrb. d. Nassauischen Ver. f. Naturk., 
Wiesbaden, 68: 60, pi. 12, f. 1; pi. 13, 1915. Bate, Cameroons. 



194 bulletin: museum of comparative zoology 

Thos adustus grayi (HilzheimeiO 

Canis lupaster grayi Hilzheimer, Zool. Beobachter, 47: 367, Dec. 1906. Mo- 
rocco and Tunis. 
Canis studeri Hilzheimer, Zool. Beobachter, 47: 368, Dec. 1906. Tunis. 

Thos adustus kaffensis (Neumann) 

Canis kaffensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 53, 1902. 
Anderatscha, Kaffa, southwestern Ethiopia. 

Thos adustus lateralis (P. L. Sclater) 

Canis lateralis P. L. Sclater, Proc. Zool. Soc. London, p. 279, pi. 23, Nov. 1870. 
Rio Fernand Vaz, south of Gaboon. 

Thos adustus notatus Heller 

Thos adustus notatus Heller, Smithsonian Misc. Coll., 63: no. 7, p. 2, 4, 24 
June 1914. Loita Plains, Kenya Colony. 

Thos aureus algirensis (Wagner). Algerian Jackal 

Canis aureus algirensis Wagner, Schreber's Saugth., Suppl., 2: 384, 1841. 

Algeria (based on I. Geoflfroy, Exped. Sci. de Moree, p. 22, 1833). 
Canis aureus tripolitanus Wagner, Schreber's Saugth., Suppl., 2: 384, 1841. 

No locality; Tripoli, Tunis, implied. 
A valid race according to Cabrera (Trabajos Mus. Nac. de Ciencias, Madrid, 
zool. ser., no. 57, p. 107, 30 Dec. 1932), but the status of all the named African 
races of Thos aureus is still in much need of revision; see especially, Schwarz, 
E., Ueber Typenexemplare von Schakalen, Senckenbergiana (Frankfurt), 8: 
34-47, 10 Mar. 1926. 

Thos aureus anthus (F. Cuvier). Senegalese Jackal 

[Canis] anthus F. Cuvier, Hist. Nat. des Mammiferes, pt. 17, pi. 173 and text, 

June 1820. Senegal. Status uncertain. 
Thos senegalensis Hamilton Smith, Jardine's Naturalist's Library, Mamm., 

9: 201, pi. 13, 1839. Senegal. 

Thos aureus bea Heller. East African Jackal 

Thos aureus bea Heller, Smithsonian Misc. Coll., 63: no. 7, p. 2,5,24 .June 1914. 
Loita Plains, Kenya Colony. 

Thos aureus lupaster (Hemprich and Ehrenberg). Egyptian Jackal 

Canis lupaster Hemprich and Ehrenberg, Symbolae Physicse, Mamm., dec. 2, 

folio ff, Nov. 1832. Fayum, Egypt. 
Canis sacer Hemprich and Ehrenberg, Symbolae Physicae, Mamm., dec. 2, 

folio ff, Nov. 1832. Fayum, Egypt. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 195 

Thos aureus maroccanus Cabrera. Moroccan Jackal 

Thos lupaster maroccanus Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 
21: 263, 14 Oct. 1921. Mogador, Morocco. 

Thos aureus nubianus Cabrera. Nubian Jackal 

Canis variegatus Cretzschmar, Riippell's Atlas zu d. Reise im Nordlich. Afrika, 

Saugeth., p. 31, pi. 10, 1826. Nubia and upper Egypt. 
Thos aureus nubianus Cabrera, Bol. Real Soc. Espafiola Hist. Nat., Madrid, 

21 : 264, 14 Oct. 1921. New name in place of Canis variegatus Cretzschmar, 

not C. familiaris variegatus Gmelin, 1788. 

Thos aureus riparius (Hemprich and Ehrenberg). Red Sea Jackal 

Canis riparius Hemprich and Ehrenberg, Symbolae Physicse, Mamm., dec. 
2, fol. ff, Nov. 1832. Coast of Abyssinia, near Arkiko. 

Canis hagenbecki Noack, Zool. Garten, 27: 234, Aug. 1886; Zool. Anzeiger, 
20: 518, 27 Dec. 1897. Somaliland. 

Canis mengesi Noack, Zool. Anzeiger, 20: 518, 27 Dec. 1897. Coast of Somali- 
land. 

Canis somalicus Lorenz, Anzeiger K. K. Akad. Wiss. Wien, math.-nat. Klasse, 
43: 306, 1906. Ireso, near Agada, Somaliland. 

Thos aureus soudanicus (Thomas). Sudanese Jackal 

Canis anthus soudanicus Thomas, Proc. Zool. Soc. London, 1 : 295, Aug. 1903. 

El Obeid, Kordofan. 
According to Schwarz, this is probably, with the following synonyms, the 
same as Thos aureus nubianus Cabrera. 
Canis doederleini Hilzheimer, Zool. Anzeiger, 30: 116, 17 Apl. 1906. Upper 

Egypt. 
Canis thooides Hilzheimer, Zool. Beobachter, 47: 364, Dec. 1906. Sennaar. 

Thos gallaensis (Lorenz) 

Canis gallaensis Lorenz, Anzeiger K. K. Akad. Wiss. Wien, math.-nat. Klasse, 
43: 307, 1906. Ginea, Arussi, Ethiopia. Status uncertain. 

Thos lamperti (Hilzheimer) 

Canis mengesi lamperti Hilzheimer, Zool. Beobachter, 47: 371, Dec. 1906. 
?Somaliland. Status uncertain. 

Thos mesomelas mesomelas (Schreber). Black-backed Jackal 

Canis mesomelas Schreber, Saugthiere, 3: 370, 586, pi. 95, 1778. Cape of Good 

Hope. 
Canis variegatoides A. Smith, So. African Quart. Journ., 2: 87, 1834. 

Thos mesomelas achrotes Thomas 

Thos mesomelas achrotes Thomas, Proc. Zool. Soc. London, p. 295, 29 Apl. 
1926. Rooibank, Kuisib River, Southwest Africa. 



196 bulletin: museum of comparative zoology 

Thos mesomelas arenarum Thomas 

Thos mesomelas arenarum Thomas, Proc. Zooh Sec. London, p. 295, 29 Apl. 
1926. Berseba, Southwest Africa. 

Thos mesomelas elgon^ Heller 

Thos mesomelas elgonse Heller, Smithsonian Misc. Coll., 63: no. 7, p. 3, 6, 
24 June 1914. Guas Ngishu plateau, Kenya Colony. 

Thos mesomelas mcmillani Heller 

Thos 7nesomelas mcmillani Heller, Smithsonian Misc. Coll., 63: no. 7, p. 3, 6, 
24 June 1914. Mtoto Andei, Kenya Colony. 

Thos mesomelas schmidti (Noack) 

Canis mesomelas var. schmidti Noack, Zool. Anzeiger, 20: 519, 27 Dec. 1897. 
Interior of Somaliland. 



VuLPES Oken. Red Foxes 

Vulpes Oken, Lehrbuch d. Naturgesch., sect. 3, pt. 2, p. 1033, 1266, 1816. 
Genotype Vulpes vulgaris Oken = Canis vulpes Linnaeus. 

Vulpes chama (A. Smith). Chama Fox 

Canis chama A. Smith, South African Quart. Journ., 2: 89, 1833. "Namaqua- 

land and the country on both sides of the Orange River," South Africa. 
Megalotis caania Hamilton Smith, Jardine's Natm-alist's Library, Mamm., 9: 

236, pi. 19, 1839. South Africa. 
Megalotis hodsoni Noack, Zool. Anzeiger, 35: 461, 15 Feb. 1910. Transvaal. 

Vulpes pallida pallida (Cretzschmar). Sand Fox 

Canis pallidus Cretzschmar, in Riippell, Atlas zu d. Reise im Nordlich. Afrika, 

Saugeth., p. 33, pi. 11, 1826. Kordofan. 
? Canis sabbar Hemprich and Ehrenberg, Symbolse Physicse, Mamm., dec. 2, 

folio ff, Nov. 1832. Dongola. 
See Thomas, O. Some notes on the small sand-foxes of North Africa. Ann. 
Mag. Nat. Hist. (9) 1: 242-245, Mar. 1918. 

Vulpes pallida cyrenaica Festa 

Vulpes cyrenaica Festa, Boll. Mus. Zool. ed Anat. Comp. Univ. Torino, 36: no. 
740, p. 3, 15 Dec. 1921. Near Benghazi, Barka Cyrenaica, Libya. 

Vulpes pallida edwardsi Rochebrune 

Vulpes edwardsi Rochebrune, Bull. Soc. Philom., Paris, (7) 7: 8, 1883. Cayor, 
Oualo, Gandiole, and forests of Gommiers, Senegambia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 197 

VuLPES PALLIDA HARTERTi Thomas and Hinton 

Vulpes pallida harterti Thomas and Hinton, Novitates Zool., 28:4, May 1921. 

Takoukout, Damergou, northern Nigeria, 1550 ft. alt. 
Canis pallidus haterti Zimara, Sitzb. K. K. Akad. Wiss., Wien, math.-nat. Kl., 

144: sect. 1, no. 1-2, p. 9, 1935. Gourma-Bahrus, French West Africa. 

Vulpes pallida oertzeni (Matschie) 

Canis (Cynalopex) pallidus oertzeni Matschie, Sitzb. Ges. Naturf. Freunde, 

Berhn, p. 370, Oct. 1910. Dikoa, northeastern Cameroons (=northern 

Nigeria) . 

Vulpes RtjppELLii RtJpPELLii (Schinz) 

Canis riippelii (sic) Schinz, Cuvier's Thierreich, 4: 508, 1825. Dongola. 
Canis famelicus Cretzschmar, in Riippell, Atlas zu d. Reise im NordUch. Afrika, 
Saugeth., p. 15, pi. 5, 1826. Nubian desert and Kordofan. 

Vulpes rxjppellii c^sia Thomas and Hinton 

Vulpes ruppelli csesia Thomas and Hinton, Novitates Zool., 28: 5, May 1921. 
South side of Mt. Baguezan, Asben, northern Nigeria, 3300 ft. 

Vulpes RtJppELLii somali^e Thomas 

Vulpes ruppelli somalise Thomas, Ann. Mag. Nat. Hist. (9) 1: 244, Mar. 1918. 
Berbera, Somaliland. 

Vulpes vulpes ^gyptiaca (Sonnini). Egyptian Red Fox 

Canis segyptiacus Sonnini, Nouv. Dictionnaire des Sci. Nat., 6: 524, 1816. 

Egypt. 
Canis niloticus Desmarest, Encyclop. Methodique, Mammalogie, p. 204, 1820. 

Egypt. 
Canis anubis Hemprich and Ehrenberg, Symbolae Physicae, Mamm., dec. 2, 

folio ff, Nov. 1832. Fayum, Egypt. 
Canis vulpecula Hemprich and Ehrenberg, Symbolae Physicae, Mamm., dec. 2, 

folio ff, Nov. 1832. Fayum, Egypt. 

Vulpes vulpes atlantica (A. Wagner) 

Canis vulpes, var. atlantica A. Wagner, in M. Wagner, Reisen in d. Regenschaft 

Algier, 3: 31, 62, pi. 3, 1841. Atlas Mts., Mitiyd, Algeria. 
Vulpes algeriensis Loche, Cat. Mamm. et Oiseaux observes en Algerie, p. 4, 

1858; Explor. Sci. de I'Algerie, Zool., Mamm., p. 21, 1867. Wooded parts 

of Algeria. 

Vulpes vulpes barbarus (Shaw). Barbary Fox 

Canis barbarus Shaw, General Zool., 1: Mamm., pt. 2, p. 311, 1800. Barbary, 
i.e. coast of northwestern Africa (based on specimen described by Bufifon, 
Hist. Nat., Suppl., 4: 50, pi. 14, 1778, and Pennant, Hist. Quadrupeds, 1: 
260, 1781). 
This name, if identifiable, should supersede V. v. acaab Cabrera. 



198 bulletin: museum of compakative zoology 

Vulpes vulpes acaab Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 16: 
884, Oct. 1916. Marraquex, western Morocco. 
Range: Western Morocco. 

Vulpes vxjlpes dorsalis (Gray) 

Canis vulpes dorsalis Gray, Proc. Zool Soc. London, for 1837, p. 132, 14 June 
1838. Senegal. 

Family VIVERRID.^. Civets, Mungooses, etc. 

Subfamily Viverrin^. Civets, Genets 

CiVETTiCTis Pocock. African Civets 

Civettictis Pocock, Proc. Zool. Soc. London, 1: 134, 26 March 1915. Geno- 
type Viverra civetta Schreber. 
Viverra in part, of authors. 

Civettictis civetta civetta (Schreber). West African Civet 

Viverra civetta Schreber, Saugthiere, 3: 418, 587, pi. Ill, 1778. "Guinea, 

Kongo, das Vorgebirge der guten Hofnung und Aethiopien," type locality 

taken as French Guinea. 
Viverra poortmanni Pucheran, Rev. et Mag. de Zool. (2) 7: 304, June 1855. 

Gaboon (see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 117, 11 Apl. 

1924). 

Civettictis civetta congica Cabrera 

Civettictis civetta congica Cabrera, Mem. Real Soc. Espanola Hist. Nat., 
Madrid, 16: 36, footnote, 10 July 1929. Niapu, Congo Beige. Based on 
the account of C. c. orientalis J. A. Allen (not Matschie), Bull. Amer. Mus. 
Nat. Hist., 47: 109, text-fig. 12-15, pis. 12, 13, 11 Apl. 1924; type, the 
specimen from Niapu figured in PI. 12, no. 51811 A.M.N.H. 

Civettictis civetta schwarzi Cabrera. East African Civet 

Viverra civetta orientalis Matschie, Arch. f. Naturgesch., 57: sect. A, pt. 1, 

p. 352, Oct. 1891 (not Viverra orientalis Hodgson, 1842, = F. zibetha 

Linnaeus). Zanzibar, Bagamoyo, Tanganyika Territory, of which the 

former is selected as type locality by Schwarz. 
Viverra megaspila Noack, Jahrb. Hamburg. Wiss. Anstalt., 9: 124, pi. 1, 

f.5; pl.2, f.5,6, 1891 (not of Blyth). 
Civettictis civetta schwarzi Cabrera, Mem. Real Soc. Espanola Hist. Nat., 

Madrid, 16: 36, footnote, 10 July 1929 (new name to replace orientalis 

Matschie, preoccupied). Type locality selected as Zanzibar (see Schwarz, 

Ann. Mag. Nat. Hist. (10) 14: 261, Aug. 1934). 
[Viverra civetta] matschiei Pocock, Journ. Bombay Nat. Hist. Soc, 36: 429, 

footnote, 15 Apl. 1933, to replace orientalis, preoccupied. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 199 

Fossa Gray. The Foussa 

Fossa Gray, Proc. Zool. Soc. London, for 1864, p. 518, May 1865. Genotype 
Fossa daubentonii Gray = Viverra fossa Schreber. 

Fossa fossa (Schreber). The Foussa 

Viverrafossa Schreber, Siiugthiere, 3 : 424, 587, 1777; pi. 114, 1776 (plate name). 

Madagascar. 
Viverra fossana P. L. S. Miiller, von Linne, Natur-systems, Suppl., p. 32, 1776, 

based on Buffon and Daubenton, Hist. Nat., ed. 2, 13: 88, pi. 21, 1770. 

No locality. 
Fossa daubentonii Gray, Proc. Zool. Soc. London, for 1864, p. 518, May 1865. 

Madagascar. 

Genetta Oken. The Genets 

Genetta Oken, Lehrbuch d. Naturgesch., sect. 3, zool., pt. 2, p. 1010, 1816. 

As a subgenus of Viverra; type Viverra genetta Linnaeus. 
Viverra Linnseus et auct. vet. (in part). 

Review: Schwarz, E. Die Sammlung afrikanischer Saugethiere im Congo- 
Museum. Ginsterkatzen (Gattung Genetta Oken). Rev. Zool. et Bot. Afri- 
caines, 19: 275-286, 30 July 1930. 

Genetta abyssinica (Rtippell) 

Viverra abyssinica Riippell, Neue Wirbelth. zu Fauna Abyssinien gehorig, 

Saugeth., 1: 33, pi. 11, 1836. Abyssinia. 
Genetta abyssinica Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 30 July 

1930. 
Range: Abyssinia (= Ethiopia). 

Genetta genetta afra F. Cuvier 

Genetta afra F. Cuvier, in F. Cuvier and Geoffroy, Hist. Nat. des Mammif^res, 

pt. 52, pi. 195 and pt. 51, text, Sep. 1825. Barbary. 
Genetta genetta afra Schwarz, Rev. de Zool. et Bot. Africaines, 19: 279, 30 July 

1930. 
Genetta barbara Hamilton Smith, Jardine's Naturalist's Library, Mamm., 13: 

171, 1842. Barbary. 
Genetta bonaparti Loche, Rev. et Mag. de Zool. (2) 9: 385, pi. 13, Sep. 1857. 

Algeria. 
Range: Western Morocco, Algeria and Tunis. 

Genetta genetta felina (Thunberg) 

Viverra felina Thunberg, Kongl. Svenska Vet.-Acad. Nya Handl., Stockholm, 
32: 165, 1811. Cape of Good Hope. 



200 bulletin: museum of comparative zoology 

Geneita genetta felina Schwarz, Rev. de Zool. et Bot. Africaines, 19: 279, 30 

July 1930. 
Viverra niacrura Jentink, ex Temminck MS., Mus. d'Hist. Nat. Pays-Bas, 

Leiden, 11: 112, 1892. Nomen nudum. 

Genetta genetta hararensis Neumann 

Genetta hararensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 183, 1902. 

Harar, Abyssinia (Ethiopia). 
Genetta guardafuensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 183, 

1902. Cape Guardafui, eastern SomalUand. 
Genetta albipes Trouessart, Cat. Mamm. Viv. Foss., pt. 1, p. 250, 1904. Harar, 

Abyssinia (based on Matschie's "weissfiissige Genette, G. spec", in 

Verb. d. V. Internat. Zool. Congress. Berlin, 1901, p. 1139, 1902). 
Genetta dongolana tedescoi de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. 

Nat., Milano, 62: 267, Feb. 1924. Tigiglo, a third the way from Webi 

Shebeli to Juba River, Italian SomalUand. 

Genetta genetta hintoni Schwarz 

Genetta genetta hintoni Schwarz, Ann. Mag. Nat. Hist. (10) 3: 47, Jan. 1929. 
N'dola, northeastern Rhodesia, 4220 ft. 

Genetta genetta neumanni Matschie 

Genetta neumanni Matschie, Verb. d. V. Internat. Zool. Congress. Berlin, 1901, 

p. 1140, 1902. Irangi, northern Ugogo, and Kitui in Ukamba, Kenya 

Colony. 
Genetta genetta neumanni Schwarz, Rev. de Zool. et Bot. Africaines, 19: 279, 

30 July 1930. 
Range: Mt. Elgon, Kenya Colony, to Mt. Kilimanjaro region. 

Genetta genetta pulchra Matschie 

Genetta pulchra Matschie, Verb. d. V. Internat. Zool. Congress. Berlin, 

1901, p. 1139, 1902. Okondoto, Okawango, Otawi, Windhoek, Southwest 

Africa. 
Genetta geneita pulchra Schwarz, Rev. de Zool. et Bot. Africaines, 19: 279, 

30 July 1930. 
Genetta bella Matschie, Verb. d. V. Internat. Zool. Congress. Berlin, 1901, p. 

1140, 1902. Loanda, Angola. 
Genetta India Thomas and Schwann, Proc. Zool. Soc. London, p. 579, 10 Oct. 

1906. Klein Letaba, northeastern Transvaal. 

Genetta genetta senegalensis (J. B. Fischer) 

Viverra senegalensis J. B. Fischer, Synopsis Mamm., p. 170, 1829. Senegal. 
Genetta genetta senegalensis Schwarz, Rev. de Zool. et Bot. Africaines, 19: 279, 
30 July 1930. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 201 

Viverra dongolana Hemprich and Ehrenberg, Symbolse Physicse, Zool., Mamm., 

dec. 2, folio k, p. 2, Sep. 1832. Dongola, Anglo-Egyptian Sudan. 
Viverra leptura Reichenbach, Regnum Animale, 1 : 23, f. 270, 1836. 

Genetta maculata (Gray) 

Viverra maculata Gray, Spicilegia Zool., 2: 9, pi. 9, 1 Aug. 1830. North Africa, 

but doubtless in reality from Senegal (see Schwarz, Rev. Zool. et Bet. 

Africaines, 19: 277, footnote, 30 July 1930). 
Genetta pardina I. Geoffroy, Mag. de Zool., 2: cl. 1, pi. 8 and text, 1832. In- 
terior of Senegal. 
Genetta poensis Waterhouse, Proc. Zool. Soc. London, p. 59, July 1838. Island 

of Fernando Po? (see Pocock, Proc. Zool. Soc. London, 1907, 2: 1039- 

1041, 4 June 1908). 
Viverra genettoides Temminck, Esquisses Zool. sur la Cote de Guin6, p. 89, 

1853. Rio Boutry and La Mina, Gold Coast. 
Genetta dubia Matschie, Verh. des V. Internat. Zool. Congress. Berlin, 1901, 

p. 1141, 1902. West Africa. 
Genetta johnstoni Pocock, Proc. Zool. Soc. London, for 1907, p. 1041, pi. 54, 

f. 1, 2, 4 June 1908. West of Putu Mts., west of Cavally River, Liberia. 
Range: Forests of Upper Guinea from the Niger to Gambia (Schwarz). 

Genetta servalina servalina Plicheran 

Genetta servalina Pucheran, Rev. et Mag. de Zool. (2) 7: 154, Mar. 1855. 

Gaboon. 
Genetta aubryana Pucheran, Rev. et Mag. de Zool. (2) 7: 154, Mar. 1855. 

Gaboon. 
Range: Forest region from Niger to the Rift Valley, west on both banks of 

the Congo to the West Coast. 

Genetta servalina bettoni Thomas 

Genetta bettoni Thomas, Ann. Mag. Nat. Hist. (7) 9: 365, May 1902. Lagari, 
Mau district, south of El Burgon Range, Kenya Colony. 
Range: Uele region to Lake Ivivu and Mt. Elgon. 

Genetta servalina intensa Lonnberg 

Genetta servalina intensa Lonnberg, Kungl. Svenska Vet.-Akad. Handl., Stock- 
holm, (2) 58: no. 2, p. 59, 1 Sep. 1917. Beni and Masisi, Congo Beige. 

Genetta tigrina tigrina (Schreber) 

Viverra tigrina Schreber, Saugthiere, 3: 425, pi. 115, 1778. Cape of Good Hope. 

Genetta rubiginosa Pucheran, Rev. et Mag. de Zool. (2) 7: 154, Mar. 1855. Cape 

of Good Hope. 
Range: As a species, from Abyssinia and the Sudan, west to Senegal, south 

of the Sahara, to the edge of the Congo rainforest and on the east coast to 

Cape of Good Hope. 



202 bulletin: museum of compakative zoology 

Genetta tigrina .equatorialis Heuglin 

Genetta sequatorialis Heuglin, Sitzb. K. Akad. Wiss., Wien, math.-nat. KL, 54: 
sect. 1, p. 559, 1866. West bank of Bahr-el-Abiad, 7°-8° N., Anglo- 
Egyptian Sudan. 

Genetta tigrina sequatorialis Schwarz, Rev. de Zool. et Bot. Africaines, 19: 277, 
30 July 1930. 
Range: Uele to Ituri forest, south to Lake Tanganyika. 

Genetta tigrina albiventris Roberts 

Genetta rubiginosa albiventris Roberts, Ann. Transvaal Mus., 15: 1, 1 Oct. 
1932. Maun, Ngamiland, Bechuanaland. 

Genetta tigrina amer Gray 

Genetta amer Gray, List Spec. Mamm. Brit. Mus., p. 49, 13 May 1843. 

Genetta tigrina amer Gray, Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278 

and footnote, 30 July 1930. ?Nomen nudum, cites Rueppell specimen in 

BM. 
Genetta matschiei Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 183, 1902. 

Harar, Abyssinia (Ethiopia). 
Genetta schraderi Matschie, Verb. d. V. Internat. Zool. Congress. Berlin, 1901, 

p. 1140, 1902. Massawa, Eritrea. 

Genetta tigrina angolensis Bocage 

Genetta angolensis Bocage, Jorn. Sci. Math.,Phys. e Nat., Lisboa, (1) 9: 29, July 

1882. Caconda, Angola. 
Genetta tigrina angolensis Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 

30 July 1930. 
Genetta gleimi Matschie, Verb. d. V. Internat. Zool. Congress. Berlin, 1901, 

p. 1142, 1902. Loanda, Angola. 
Genetta loandse Matschie, Schwarz, Rev. Zool. et Bot. Africaines, 19: 278, 30 

July 1930 (lapsus cal. for G. gleimi Matschie?). 
Range: Lower Congo to Angola. 

Genetta tigrina erlangeri Matschie 

Genetta erlangeri Matschie, Verb. d. V. Internat. Zool. Congress. Berlin, 1901, 

p. 1143, 1902. Kitui, Kenya Colony. 
Genetta tigrina erlangeri Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 30 

July 1930. 
Genetta pumila HoUister, Smithsonian Misc. Coll., 66: no. 1, p. 4, 10 Feb. 1916. 

Mt. Gargues, Kenya Colony, 6000 feet. 

Genetta tigrina fieldiana DuChaillu 

Genetta fieldiana DuChaillu, Proc. Boston Soc. Nat. Hist., 7: 302, July 1860. 

Interior of Gaboon. See J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 119, 

11 Apl. 1924. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 203 

Genetta tigrina insularis Cabrera 

Genetta insularis Cabrera, Bol. Real Soc. Espafiola Hist. Nat., Madrid, 21: 261, 

14 Oct. 1921. Rebola, Island of Fernando Po. 
Genetta tigrina insularis Schwarz, Rev. Zool. et Bot. Africaines, 19: 277, 30 July 

1930 with ?. 

Genetta tigrina letab^ Thomas and Schwann 

Genetta letabse Thomas and Schwann, Proc. Zool. Soc. London, p. 578, 10 Oct. 

1906. Klein Letaba, northeastern Transvaal. 
Genetta tigrina letabse Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 30 July 

1930. 
Genetta rubiginosa ziduensis Roberts, Ann. Transvaal Mus., 10: 67, 31 Jan. 

1924. Banks of the White Umfolosi, Umfolosi Game Reserve, Zululand, 

Natal. 

Genetta tigrina mossambica Matschie 

Genetta mossambica Matschie, Verh. d. V. Intemat. Zool. Congress. Berhn, 

1901, p. 1138, 1902. Mossimboa, Querimba Id., and southern Tanganyika 

Territory. 
Genetta tigrina mossambica Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 

30 July 1930. 
Genetta zambesiana Matschie, Verh. d. V. Internat. Zool. Congress. Berlin, 

1901, p. 1141, 1902. Boror, Nyasaland. 

Genetta tigrina soror Schwarz 

Genetta tigrina soror Schwarz, Ann. Mag. Nat. Hist. (10) 3: 48, Jan. 1929. 
Kimdilungo Mts., southwest of Lake Mweru, Congo Beige. 

Genetta tigrina stuhlmanni Matschie 

Genetta stuhlmanni Matschie, Verh. d. V. Internat. Zool. Congress. Berlin, 

1901, p. 1142, 1902. Bukoba, Tanganyika Territory. 
Genetta tigrina stuhlmanni Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 

30 July 1930. 

Genetta tigrina suahelica Matschie 

Genetta suahelica Matschie, Verh. d. V. Internat. Zool. Congress. Berlin, 1901, 

p. 1143, 1902. East coast of Tanganyika Territory to Mombasa, Kenya 

Colony. 
Genetta tigrina suahelica Schwarz, Rev. de Zool. et Bot. Africaines, 19: 278, 30 

July 1930. 
Range: Coast of Tanganyika Territory and Kenya Colony to ? Mt. Elgon. 

Genetta tigrina thierryi Matschie 

(jene/to i/iierrt/i Matschie, Verh. d. V. Internat. Zool. Congress. Berlin, 1901, 
p. 1142, 1902. Interior of Togo, Dahomy, 9° N. 



204 bulletin: museum of compakative zoology 

Genetta tigrina thierryi Schwarz, Rev. de Zool. et Bot. Africaines, 19: 277, 30 
July 1930. 

Genetta victorijE Thomas 

Genetta victorise Thomas, Proc. Zool. Soc. London, 2: 87, pi. 5, 1 Oct. 1901. 
Entebbe, Uganda. 
Range: Local in forests between Ubangi, Congo, and Lake Victoria. 



Nandinia Gray. Tree Civet 

Nandinia Gray, Cat. Spec. Mamm. Brit. Mus., p. 54, 13 May 1843, nomen 
nudum; Proc. Zool. Soc. London, for 1864, p. 529, May 1865. Genotype 
Viverra binotata Reinwardt. 

Nandinia binotata binotata (Reinwardt) 

Viverra binotata Reinwardt, in Gray, Spicilegia Zool., 2: 9, 1830. Fernando Po. 

Paradoxurus hamiltonii Gray, Proc. Zool. Soc. London, p. 67, 5 June 1832; 

lUustr. Indian Zool., pi. 10, 1833-34. No locality; Ashanti according to 

Temminck. 

Nandinia binotata arborea Heller 

Nandinia binotata arborea Heller, Smithsonian Misc. Coll., 61: no. 13, p. 9, 16 
Sep. 1913. Lukosa River, northeast of Kisumu, Kenya Colony. 

Nandinia binotata gerrardi Thomas 

Nandinia gerrardi Thomas, Ann. Mag. Nat. Hist. (6) 12: 205, Sep. 1893. 
Lower Shire River, Nyasaland. 

Nandinia binotata intensa Cabrera and Ruxton 

Nandinia binotata intensa Cabrera and Ruxton, Ann. Mag. Nat. Hist. (9) 17: 
596, May 1926. St. Joseph de Luluabourg, southern Congo Beige. 



OsBOKNiCTis J. A. Allen. Aquatic Civet 

Osbornictis J. A. Allen, Journ. Mammalogy, 1 : 25, 28 Nov. 1919; Bull. Amer. 
Mus. Nat. Hist., 47: 135, 11 Apl. 1924. Genotype Obsornidis piscivora 
J. A. Allen. 

Osbornictis piscivora J. A. Allen 

Osbornictis piscivora J. A. Allen, Journ. Mammalogy, 1: 25, 28 Nov. 1919; 
Bull. Amer. Mus. Nat. Hist., 47: 140, text-f. 21, 23, A, B; 24, A; 25, A; 26, 
A, B; 28, A, C; 29, A, C; pis. 18, 19, 11 Apl. 1924. Niapu, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 205 



PoiANA Gray. African Linsang 

Linsang Gerrard, Cat. Bones Mamm. Brit. Mus., p. 71, 1862 (in part). 
Poiana Gray, Proc. Zool. Soc. London, for 1864, p. 520, May 1865. Genotype 

Linsang richardsoni Gerrard = Genetta poensis Waterhouse, Proc. Zool. Soc. 

London, p. 59, 1838 (juv.), not of Waterhouse, ibid, (adult). 

Poiana richardsonii richardsonii (Thomson) 

Genetta richardsonii Thomson, Ann. Mag. Nat. Hist. (1) 10: 204, Nov. 1842. 

Fernando Po. 
Genetta poensis Waterhouse, Proc. Zool. Soc. London, p. 59, July 1838 (the 

young individual but not the adult on which the name was based), in part. 

Poiana richardsonii liberiensis Pocock 

Poiana richardsoni liberiensis Pocock, Proc. Zool. Soc. London, for 1907, p. 

1043, pi. 54, f. 3, 4 June 1908. West of Putu Mts., west of Cavally River, 

Liberia. 
Poiana richardsoni leightoni Pocock, Proc. Zool. Soc. London, for 1907, p. 1045, 

4 June 1908. West of Putu Mts., Liberia (lapsus cal. for liberiensis). 

Poiana richardsonii ochracea Thomas and Wroughton 
Poiana richardsoni ochracea Thomas and Wroughton, Ann. Mag. Nat. Histi 
(7) 19: 372, May 1907. Near Yambuya, Aruwimi River, Congo, about 1° 

N., 25° E. 

ViVERRicuLA Hodgson 

Viverricula Hodgson, Ann. Mag. Nat. Hist. (1) 1: 152, Apl. 1838. Genotype 
not specified but included Viverra indica Geoffroy ( = V. malaccensis 
Gmelin) and V. rape ( = F. rasse Horsfield). 

Viverricula rasse (Horsfield). The Rasse. 

Viverra rasse Horsfield, Zool. Researches in Java, no. 6, pi. and text (un- 
numbered), 1821. Java. 

Viverra schlegelii Pollen, in Schlegel, Nederlandsch Tijdschr. v. Dierkunde, 3: 
78, 1866. Islands of Mayotte and Nossi-Faly, northwest of Madagascar. 
(Introduced). 
Range: Java; introduced in Socotra, Madagascar and Comoro Islands. 



Subfamily Herpestin.^. Mungooses 
(See J. A. Allen, BuU. Amer. Mus. Nat. Hist., 47: 153, 11 Apl. 1924.) 



206 bulletin: museum of comparative zoology 

Atilax F. Cuvier. Marsh Mungooses 

Atilax F. Cuvier, Hist. Nat. des Mammiferes, pt. 54, pi. and text on the "Van- 
sire" (not of Buffon and Daubenton), June 1826. Genotype Herpestes 
paludinosus G. Cuvier (see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 
171, 11 Apl. 1924). 

Athylax I. Geoffroy, Mag. de Zool. (2) 1 : mammif., art. 4, p. 25, 1839; Gray, 
Proc. Zool. Soc. London, for 1864, p. 556, 1865. Genotype Herpestes palu- 
dinosus G. Cuvier. 

Atilax paludinosus paludinosus (G. Cuvier) 

Mustela galera Schreber, Saugthiere, 3: pi. 135, 1777. Madagascar. This is the 
Vansire of Buffon and Daubenton, but the name is regarded as unidenti- 
fiable (see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 163, 11 Apl. 1924). 

Mustela voang-shire Zimmermann, Specimen Zoologiae Geographicae, p. 487, 
1777. See Allen, J. A., Bull. Amer. Mus. Nat. Hist., 16: 16, 1 Feb. 1902. 

Herpestes paludinosus G. Cuvier, Regne Anim., ed. 2, 1: 158, (Apl.) 1829. 
Cape of Good Hope. 

Mangusta urinatrix A. Smith, Zool. Journ., 4: 437, May, 1829. South Africa. 

Herpestes atilax Wagner, Schreber's Saugthiere, Suppl., 2: 305, 1841. South 
Africa. 

Athylax paludosus Gray, Proc. Zool. Soc. London, for 1864, p. 557, May 1865; 
Cape of Good Hope. 

Atilax paludinosus macrodon J. A. Allen 

Atilax macrodon J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 205, text-ff. 61, 

C-D; 62-64, pi. 27, f. 2; 29, 11 Apl. 1924. Niapu, northeastern Congo 

Beige. 

Atilax paludinosus mitis (Thomas and Wroughton) 

Herpestes galera mitis Thomas, Proc. Zool. Soc. London, for 1902, 2: 309, Apl. 
1903. Zegi, Lake Tana, Ethiopia. 

Atilax paludinosus mordax (Thomas) 

Mungos paludinosus mordax Thomas, Ann. Mag. Nat. Hist. (8) 10: 588, Dec. 
1912. Rombashi River, northwest of north end of Lake Nyasa, Tangan- 
yika Territory. 

Atilax paludinosus naso (De Winton) 

Herpestes naso De Winton, Bull. Liverpool Museums, 3: 35, pi. 1, 1 text-f., 

May 1901. Cameroon River, Cameroons. 
Mungos paludinosus naso Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 58: no. 2, p. 64, 1 Sep. 1917. 
Herpestes almodovari Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 2: 

139, Mar. 1902. Cape San Juan, Biafra. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 207 

Atilax paludinosus nigerianus (Thomas) 

Mungos naso nigerianus Thomas, Ann. Mag. Nat. Hist. (8) 10: 588, Dec. 1912. 
Niaji, 20 miles northeast of Oban, Nigeria. 

Atilax paludinosus pluto (Temminck) 

Herpesies pluto Temminck, Esquisses Zool. sur la Cote de Guin6, p. 95, 1853. 
Dabocrom, Gold Coast. 
Range: Gold Coast to Sierra Leone. 

Atilax paludinosus robustus Gray 

Athylax robusius Gray, Proc. Zool. Soc. London, for 1864, p. 558, text-f. (skull). 
May 1865. White Nile. 

Atilax paludinosus rubellus (Thomas and Wroughton) 
Mungos paludinosus rubellus Thomas and Wroughton, Proc. Zool. Soc. London, 
p. 166, 17 Sep. 1908. Tambarara, Gorongoza district, Mozambique. 

Atilax paludinosus rubescens (HoHister) 

Mungos paludinosus rubescens HolUster, Proc. Biol. Soc. Washington, 25: 1, 
23 Jan. 1912. Mt. Kilimanjaro, Tanganyika Territorj^ 4000 ft. 

Atilax paludinosus spadiceus Cabrera 

Atilax paludinosus spadiceus Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., 
Madrid, 21 : 262, 14 Oct. 1921. Cape San Juan, Spanish Guinea. 

Atilax paludinosus transvaalensis Roberts 

Atilax paludinosus transvaalensis Roberts, Ann. Transvaal Mus., 15: 266, 1 
Aug. 1933. Mokeetsi, northern Transvaal. 



Bdeogale Peters 

Bdeogale Peters, Mitth. Ges. Naturf. Freunde, Berlin, 19 Nov. 1850; Monatsb. 
K. Preuss. Akad. Wiss., Berlin, p. 81, Feb. 1852. Genotype Bdeogale 
crassicauda Peters. 

Bdeogale crassicauda crassicauda Peters 

Bdeogale crassicauda Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 81, 

Feb. 1852; Reise nach Mossambique, Saugeth., p. 119, 120, pi. 27, 1852. 

Tette, Boror, Mozambique, 17°-18° S. 

Bdeogale crassicauda omnivora Heller 

Bdeogale crassicauda omnivora Heller, Smithsonian Misc. Coll., 61: no. 13, 
p. 12, 16 Sep. 1913. Mazeras, Kenya Colony. 



208 bulletin: museum of comparative zoology 

Bdeogale puisa Peters 

Bdeogale puisa Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 82, Feb. 

1852; Reise nach Mossambique, Saugeth., p. 124, pi. 28, 1852. Mossim- 

boa, Mozambique, 11° S. 

Bdeogale tenuis Thomas and Wroughton 

Bdeogale tenuis Thomas and Wroughton, Proc. Zool. Soc. London, p. 168, 17 
Sep. 1908. Zanzibar Island. 

Crossarchus F. Cuvier 

Crossarchus F. Cuvier, Hist. Nat. des Mammiferes, pt. 47, plate and text, 
p. 3, Feb. 1825. Genotype Crossarchus obscurus F. Cuvier. 

Crossarchus alexandri Thomas and Wroughton 

Crossarchus alexandri Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 
373j May 1907. Banzyville, Ubangi, Congo Beige. 

Crossarchus ansorgei Thomas 

Crossarchus ansorgei Thomas, Ann. Mag. Nat. Hist. (8) 5: 195, Feb. 1910. 
Dalla Tando, northern Angola. 

Crossarchus gambianus (Ogilby) 

Herpestes gambianus Ogilby, Proc. Zool. Soc. London, p. 102, 9 Oct. 1835. 
Gambia. 

Crossarchus obscurus F. Cuvier 

Crossarchus obscurus F. Cuvier, Hist. Nat. des Mammiferes, pt. 47, pi. and 

p. 3, text, Feb. 1825. West Africa, probably southern Gambia. 
Herpestes punctatissimus Temminck, Esquisses Zool. sur la Cote de Guin6, 

p. 108, 1853. Gaboon River. 

Cynictis Ogilby. Mierkat 

Cynictis Ogilby, Proc. Zool. Soc. London, p. 48, 24 May 1833. Genotype 
Cynictis steedmanni Ogilhy = Herpestes penicillatus G. Cuvier. 

Cynictis penicillata penicillata (G. Cuvier) 

Herpestes penicillatus G. Cuvier, Regne Anim., ed. 2, 1 : 158, (Apl.) 1829. Cape 

of Good Hope. 
Mangusta levaillantii A. Smith, Zool. Journ., 4: 437, May 1829. South Africa. 
Cynictis steedmanni Ogilby, Proc. Zool. Soc. London, p. 49, 24 May 1833. 

"Uitenhage, on the borders of Caffraria," Cape of Good Hope. 
Cynictis typicus A. Smith, South African Quart. Journ., 2: 116, 1834. South 

Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 209 

Cynictis penicillata brachyura Roberts 

Cynictis 'penicillata brachyura Roberts, Ann. Transvaal Mus., 10: 68, 31 Jan. 
1924. Boschkop, near Johannesburg, Transvaal. 

Cynictis penicillata bradfieldi Roberts 

Cynictis penicillata bradfieldi Roberts, Ann. Transvaal Mus., 10: 69, 31 Jan. 
1924. "Quickborn," Okahandja district. Southwest Africa. 

Cynictis penicillata bechuan^ Roberts 

Cynictis penicillata bechuanse. Roberts, Ann. Transvaal Mus., 15: 4, 1 Oct. 1932. 
Gaberones, Bechuanaland, South Africa. 

Cynictis penicillata Cinderella Thomas 

Cynictis bradfieldi Cinderella Thomas, Proc. Zool. Soc. London, p. 375, 12 July 
1927. Ondongwa, central Ovamboland, Southwest Africa. 
Range: Between Windhoek and Bechuanaland border. 

Cynictis penicillata coombsi Roberts 

Cynictis penicillata coombsi Roberts, Ann. Transvaal Mus., 13: 90, 6 July 1929. 
Swarthoek, Zoutpansberg, Transvaal. 

Cynictis penicillata intensa Schwann 

Cynictis penicillata intensa Schwann, Proc. Zool. Soc. London, p. 104, 7 June 
1906. Deelfontein, Cape Colony. 
Range: Central Cape Colony. 

Cynictis penicillata kalaharica Roberts 

Cynictis penicillata kalaharica Roberts, Ann. Transvaal Mus., 15: 4, 1 Oct. 
1932. Kaotwe Pan, central Kalahari, Bechuanaland, South Africa. 

Cynictis penicillata leptura (A. Smith) 

Cynictis lepturus A. Smith, Illustr. Zool. South Africa, Mamm., pi. 17 and 
text, Sep. 1839. South Africa, "arid plains towards the Tropic of Capri- 
corn." 
Range: Desert from Kuruman to northern Bechuanaland. 

Cynictis penicillata ogilbyi (A. Smith) 

Mangusta ogilbyi A. Smith, South African Quart. Journ., 2: 117, 1834. West- 
ern Orange Free State. 

Cynictis penicillata ogilbyi Shortridge, Mammals South West Africa, 1: 141, 
1934. 
Range: Great Karoo. 



210 bulletin: museum of comparative zoology 

Cynictis penicillata pallidior Thomas and Schwann 

Cynictis -penicillata ■pallidior Thomas and Schwann, Abstr. Proc. Zool. Sec. 
London, no. 2, p. 5, 9 Feb. 1904; Proc. Zool. Soc. London, 1: 175, 9 June 
1904. Klipfontein, Little Namaqualand, Southwest Africa. 

Dologale Thomas 

Dologale Thomas, Ann. Mag. Nat. Hist. (9) 17: 183, Jan. 1926. Genotype 
Crossarchus dybowskii Pousargues. 

Dologale dybowskii (Pousargues) 

Crossarchus dybowskii Pousargues, Bull. Soc. Zool. de France, 18: 51, 1893; 

Nouv. Arch. Mus. d' Hist. Nat., Paris, (3) 6: 121, pi. 1, 1894. Ubangi, 

Congo Beige. 
For characters and synonymy, see Hayman, R. W., Ann. Mag. Nat. Hist. 
(10) 18: 626-630, Dec. 1936. 
Herpestes (Calogale) nigripes Kershaw,, Ann. Mag. Nat. Hist. (9) 13: 22, Jan. 

1924. Moru district, Mongalla, Anglo-Egyptian Sudan. See Hayman, 

R. W., Ann. Mag. Nat. Hist. (10) 18: 629, Dec. 1936, as to status. 
Helogale hirtula robusta J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 183, 

text-f. 44D, 47, 48, pi. 24, f. 1, 11 Apl. 1924. Faradje, northeastern Congo 

Beige. See Hayman, R. W., Ann. Mag. Nat. Hist. (10) 18: 629, Dec. 

1936, as to status. 

EuPLERES Doyere 

Eupleres Doyere, Bull. Soc. des Sci. Nat., Paris, no. 3, p. 45, Mar. 1835; Ann. 
des Sci. Nat., Zool., (2) 4: 280, pi. 8, Nov. 1835. Genotype Eupleres gou- 
dotii Doyere. 

Eupleres goudotii Doyere 

Eupleres goudotii Doyere, Bull. Soc. des Sci. Nat., Paris, no. 3, p. 45, Mar. 

1835; Ann. des Sci. Nat., Zool., (2) 4: 281, pi. 8, Nov. 1835. Tamatave, 

eastern Madagascar. 

Eupleres major Lavauden 

Eupleres major Lavauden, Compt. Rend. Acad. Sci. Paris, 189: 198, 1929. 
Forests near Beangona, Valley of Sombirano, northern Madagascar. 

Galerella Gray 

Galerella Gray, Proc. Zool. Soc. London, for 1864, p. 564, May 1865. Genotype 
Herpestes ochraceus Gray. 
See J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 175-179, 11 Apl. 1924, for 
history of name. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 211 

Galerella ochracea ochracea (Gray) 

Herpestes ochraceus Gray, Proc. Zool. Soc. London, for 1848, p. 138, 1 June 
1849. Abyssinia (Ethiopia). 

Galerella ochracea bocagei (Thomas and Wroughton) ? generic reference 

uncertain. 
Herpestes bocagei Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 16: 170, 

Aug. 1905. Caconda, Benguella, Angola. 
Herpestes gracilis var. punctulatus Bocage, Jorn. Sci. Math., Phys. e Nat., 

Lisboa, (2) 1: 179, Dec. 1889 (not of Gray, 1849). Caconda, also Humbe 

and interior, Angola. 

Galerella ochracea fulvidior (Thomas) 

Herpestes ochraceus fulvidior Thomas, Ann. Mag. Nat. Hist. (7) 14: 97, Aug. 
1904. Mandeira, south of Berbera, Somaliland, 3500 ft. 

Galerella ochracea perfulvida (Thomas) 

Herpestes ochraceus perfulvidus Thomas, Ann. Mag. Nat. Hist. (7) 14: 96, Aug. 
1904. Wardair, northeast of Gerlogobi, central Somaliland. 



Galeriscus Thomas 

Galeriscus Thomas, Ann. Mag. Nat. Hist. (6) 13: 522, June 1894. Genotype 
Galeriscus jacksoni Thomas. 
See Matschie, P., Die Saugethiere Deutsch-Ost-Afrikas, p. 147, 1895 (a 
Bdeogale, not a mustelid as originally described); Pocock, R. I., Ann. Mag. Nat. 
Hist. (8) 17: 179, Feb. 1916; Thomas, Ann. Mag. Nat. Hist. (10) 1: 344, Mar. 
1928 (valid genus for jacksoni and Bdeogale nigripes). 

Galeriscus jacksoni Thomas 

Galeriscus jacksoni Thomas, Ann. Mag. Nat. Hist. (6) 13: 522, June 1894. 
Mianzini, near Lake Naivasha, Kenya Colony. 

Galeriscus nigripes (Pucheran) 

Bdeogale nigripes Pucheran, Rev. et Mag. de Zool., (2) 7: 111, Feb. 1855. 

Gaboon. 
Galeriscus nigripes Thomas, Ann. Mag. Nat. Hist. (10) 1 : 344, Mar. 1928. 



Galidia I. Geoffrey 

Galidia I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 5: 580, 1837; Ann. des 
Sci. Nat., zool., (2) 8: 251, Oct. 1837. Genotype by elimination, Galidia 
elegans I. Geoffroy. 



212 bulletin: museum of comparative zoology 

Galidia elegans I. Geoffrey 

Mustela afra Kerr, in Linnseus's Animal Kingdom, p. 175, 1792. Based on the 

"Vansire" of Madagascar, =? Galidia elegans (see Thomas and Wrough- 

ton, Proc. Zool. Soc. London, p. 167, 17 Sep. 1908). 
Galidia elegans I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 5: 581, 1837. 

Madagascar. 

Galidictis I. Geoffroy 

Galictis I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 5: 581, 1837 (not of Bell, 

1826). Genotype Mustela striata E. Geoffroy. 
Galidictis I. Geoffroy, Mag. de Zool., (2) 1: Mamm., art. 5, p. 33, footnote, 

37, footnote, pis. 18, 19, 1839. Genotype Mustela striata E. Geoffroy. 

Galidictis fasciata (Gmelin) 

Viverra fasciata Gmelin, Linne's Syst. Nat., ed. 13, 1: pt. 1, p. 92, 1788. 

Based on "Le Chat Sauvage a bandes noires des Indes" of Sonnerat, but 

clearly from Madagascar (see Schwarz, E., Ann. Mag. Nat. Hist. (8) 7: 

638, June 1911). 
Viverra striata Desmarest, Encyclop. Methodique, Mammalogie, p. 210, 

1820. "La cote de Coromandel," but actually Madagascar. (Not 

Mustela striata E. Geoffroy.) 

Galidictis ornata Pocock 

Galidictis ornatus Pocock, Ann. Mag. Nat. Hist. (8) 16: 118, pi. 7, f. 2, Aug. 
1915. Andevoranto, east coast of Madagascar. 

Galidictis striata (E. Geoffroy) 

Mustela striata E. Geoffroy, Dictionnaire Class. d'Hist. Nat., 10: 214, June 

1826 (not Viverra striata Desmarest which is a synonym of Galidictis 

fasciata (Gmelin)). Madagascar. 
Pocock (Ann. Mag. Nat. Hist. (8) 16: 115, Aug. 1915) shows that the new 
form he describes as Galidictis exiniius "certainly agrees" essentially with 
E. Geoffroy's description of Mustela striata, of which it must be regarded as a 
synonym. 
Galidictis vittata Schinz, Syst. Verzeichn. Saugeth., 1: 360, 1844. Madagascar. 

The name is based primarily on the G. vittata of I. Geoffroy (Mag. de 

Zool., (2) 1: 32, pi. 18, 1839), which is probably the same as G. striata 

(E. Geoffroy), not G. vittata Gray, 1848. 
Galidia decernlineata A. Grandidier, Revue et Mag. de Zool., (2) 19: 85, Mar. 

1867. West coast of Madagascar. 
Galidictis vittata, var. rufa A. Grandidier, Revue et Mag. de Zool., (2) 21: 338, 

Sep. 1869. Madagascar. 
Galidictis exiniius Pocock, Ann. Mag. Nat. Hist. (8) 16: 116, pi. 7, f. 1, Aug. 

1915. Ambinanindrano, eastern Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 213 



Helogale Gray. Dwarf Mungooses 

Helogale Gray, Proc. Zool. Soc. London, for 1861, p. 308, text-f. (skull), Apl. 
1862. Genotype Herpestes -parvulus Sundevall. 

Helogale brunnula brtjnnula Thomas and Schwann 

Helogale brunnula Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 

33, p. 10, 5 June 1906; Proc. Zool. Soc. London, p. 581, 10 Oct. 1906. 

Klein Letaba, Zoutpansberg district, northeastern Transvaal, 1050 ft. 

Helogale brunnula brunetta Thomas 

Helogale brunetta Thomas, Proc. Zool. Soc. London, 1 : 293, 29 Apl. 1926. Rua 
Cana Falls, Cunene River, northwestern Ovamboland, Southwest Africa. 

Helogale brunnula ruficeps Kershaw 

Helogale brumiula ruficeps Kershaw, Ann. Mag. Nat. Hist. (9) 10: 103, July 
1922. Monze, 200 miles south of N'dola, northwestern Rhodesia. 

Helogale hirtula hirtula Thomas 

Helogale hirtula Thomas, Ann. Mag. Nat. Hist. (7) 14: 97, Aug. 1904. Gabride- 
hari, 60 miles west of Gerlogobi, Somaliland. 

Helogale hirtula ahlselli Lonnberg 

Helogale hirtula ahlselli Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 64, Jan. 1912; 
Kungl. Svenska Vet.-Akad. Handl., Stockholm, 48: 70, pi. 2, f. 4, 5; pi. 5 
(col., type), 26 June 1912. Thornbush north of Northern Guaso Nyiro, 
Kenya Colony. 

Helogale hirtula annulata Drake-Brockman 

Helogale hirtula annulata Drake-Brockman, Ann. Mag. Nat. Hist. (8) 10: 
612, Dec. 1912. Afgo, Webi Shebeh, near Mogadishu, Italian Somaliland. 

Helogale hirtula lutescens Thomas 

Helogale hirtula lutescens Thomas, Ann. Mag. Nat. Hist. (8) 8: 725, Dec. 
1911. Plains near north end of Lake Rudolf, Abyssinia (Ethiopia). 

Helogale hirtula powelli Drake-Brockman 

Helogale hirtula powelli Drake-Brockman, Ann. Mag. Nat. Hist. (8) 10: 612, 
Dec. 1912. Eil Hur, near Obbia, ItaHan Somaliland. 
For H. T. Powell, Treasurer of Somaliland Protectorate. 

Helogale ivori Thomas 

Helogale ivori Thomas, Ann. Mag. Nat. Hist. (9) 4: 31, July 1919. Lumbo, 
Mozambique. 
For Hon. Ivor Montague. 



214 bulletin: museum of comparative zoology 

Helogale macmillani Thomas 

Helogale jrmcmillani Thomas, Ann. Mag. Nat. Hist. (7) 18: 300, Oct. 1906. 
Delbena River, Konso, Abyssinia. 

Helogale mimetra Thomas 

Helogale mimetra Thomas, Ann. Mag. Nat. Hist. (9) 17: 183, Jan. 1926. 
Ganguela, Angola. 

Helogale parvtjla parvula (Sundevall) 

Herpestes -parvulus Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. For- 

handl., Stockholm, 3: no. 4, 121, 1846. "E Caffraria superiore, juxta 

tropicum." 

Helogale parvula nero Thomas 

Helogale parvula nero Thomas, Ann. Mag. Nat. Hist. (10) 2: 408, Nov. 1928, 

Okorosave, Kaokoveld, Southwest Africa. 
Helogale bradfieldi Roberts, Ann. Transvaal Mus., 12: 323, 29 Dec. 1928. 

Waterberg, Southwest Africa. 

Helogale percivali percivali Thomas 

Helogale percivali Thomas, Ann. Mag. Nat. Hist. (8) 8: 724, Dec. 1911. Orr 
Valley, near Mt. Nyiro, northern Kenya Colony, 4500 ft. 

Helogale percivali tenebrosa Lonnberg 

Helogale percivali tenebrosa Lonnberg, Revue Zool. Airicaine, 5: 175, 30 Apl. 
1918. Kerio River, near Mt. Elgon, Kenya Colony. 

Helogale tjndulata undulata (Peters) 

Herpestes undulatus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 81, 

Feb. 1852; Reise nach Mossambique, Saugeth., p. 114, pi. 25, 1852. 

Mossimboa, Mozambique. 

Helogale undulata affinis HoUister 

Helogale undulata affinis Hollister, Smithsonian Misc. Coll., 66: no. 1, p. 7, 10 
Feb, 1916. Summit of Mt. Lololokwi, Kenya Colony, 6000 ft. 

Helogale undulata atkinsoni Thomas 

Helogale atkinsoni Thomas, Ann. Mag. Nat. Hist, (6) 20: 378, Oct, 1897. Har- 
gaisa, Somaliland, 1500 m. 

Helogale undulata rufula Thomas 

Helogale undulata rufula Thomas, Ann. Mag. Nat. Hist. (8) 5: 194, Feb. 1910. 
Rogoro, Kikuyu, Kenya Colony, 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 215 

Helogale varia Thomas 

Helogale varia Thomas, Proc. Zool. Soc. London, 1: 119, 1 June 1902. North- 
ern Nyasaland. 

Helogale vetula Thomas 

Helogale vetula Thomas, Ann. Mag. Nat. Hist. (8) 8: 724, Dec. 1911. Lamu, 
coast of Kenya Colony. 

Helogale victorina victorina Thomas 

Helogale victorina Thomas, Proc. Zool. Soc. London, 1: 120, 1 June 1902. 
Nassa, Speke Gulf, Lake Victoria, Kenya Colony. 

Helogale victorina ochracea Thomas 

Helogale victorina ochracea Thomas, Ann. Mag. Nat. Hist. (8) 5: 194, Feb. 1910. 
Ketoma, Ankole, southern Uganda. 



Herpestes Illiger. Gray Mungooses 

Ichneumon Lac^pede, Tabl. des Div., Ord. Gen. Mamm., p. 7, 1799. Genotype 

Viverra ichneumon Gmelin; preoccupied by Ichneumon Linnaeus, 1758, in 

Hymenoptera. 
Mungos Geoffroy and Cuvier, Mag. Encycloped., 2: 184, 1795 (in part). 
Herpestes lUiger, Prodromus Syst. Mamm. et Avium, p. 135, 1811 {Herpertes 

corrected to Herpestes in Errata, p. 302). Genotype Viverra ichneumon 

Gmelin. 
Mangusta Horsfield, Zool. Researches in Java, pt. v, pi. and text, 1824 (in part). 

Genotype not specified, but four species are included; description based 

chiefly on Mangusta ( = Herpestes) javanica. 

Herpestes ichneumon ichneumon (Linnaeus) 

Viverra ichneumon Linnaeus, Syst. Nat., ed. 10, 1: 43, 1758. Egypt, "ad ripas 

Nili." 
Ichneumon pharaon Lacepede, Mem. de ITnst. Nat. des Sci. et Arts, sci. math. 

et phys., Paris, 3: 492, 1801. 
Viverra egypti Tiedemann, Zool., 1 : 364, 1808. 
Ichneumon major E. Geoffroy, Description de I'Egypte, 2: 139, footnote, 1818. 

Egypt. See Thomas and Wroughton, Proc. Zool. Soc. London, p. 167, 17 

Sep. 1908. 

Herpestes ichneumon angolensis Bocage 

H[erpestes] angolensis Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 2: 
32, 1890. Quissange, Angola. 



216 bulletin: museum of comparative zoology 

Herpestes ichneumon cafer (Gmelin) 

Viverracafra Gmelin, Linnseus's Syst. Nat.,ed. 13, 1 :pt. 1,85, 1788. Capeof Good 
Hope (based on description of a specimen from the Cape, in Schreber's 
Saugthiere, 3: 434, 1778). It is doubtful whether Schreber's description 
does not refer rather to a Myonax. 

Viverra nems Kerr, Linnseus's Animal Kingdom, p. 160, 1792. "East coast of 
Africa," but supposed by Thomas and Wroughton (Proc. Zool. Soc. 
London, p. 167, 17 Sep. 1908) to be probably an Indian species; the des- 
cription seems to be based on Schreber's account of a mungoose which 
Gmelin independently named Viverra cafra in 1788. 

Herpestes madagascarensis A. Smith, South African Quart. Journ., 2: 114, 1834. 
Madagascar (introduced). 

Herpestes Bennettii Gray, Charlesworth's Mag. Nat. Hist., 1: 578, Nov. 
1837. Madagascar (introduced). 

Herpestes dorsalis Gray, Proc. Zool. Soc. London, for 1864, p. 549, May 1865. 
South Africa; based on Ichneumon pharaonis, var. A. Smith. 

Herpestes ichneumon centralis (Lonnberg) 

Mungos ichneumon centralis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 
Stockholm, (2) 58: no. 2, p. 61, 1 Sep. 1917. Beni, Congo Beige. 

Herpestes ichneumon funestus (Osgood) 

Mungos ichneumon funestus Osgood, Publ. Field Mus. Nat. Hist., zool. ser 
10: no. 3, p. 17, 7 Apl. 1910. Naivasha, Kenya Colony. 

Herpestes ichneumon mababiensis Roberts 

Herpestes cafer mababiensis Roberts, Ann. Transvaal Mus., 15: 4, 1 Oct. 1932; 
Tsotsoroga Pan, Mababe Flats, Bechuanaland, South Africa. 

Herpestes ichneumon numidicus (F. Cuvier) 

Ichneumon numidicus F. Cuvier, Hist. Nat. des Mammiferes, pt. 68, pi. 191, 

and text, 1834. Algeria. 
Herpestes ichneumon numidicus Cabrera, Trab. Mus. Nac. de Cienc. Nat., 

Madrid, zool. ser., no. 57, p. 149, 30 Dec. 1932. 
Herpestes numidianus Gray, Proc. Zool. Soc. London, for 1864, p. 550, May 

1865. Numidia. 
Range: Northern Morocco and Algeria. 

Herpestes ichneumon parvidens (Lonnberg) 

Mungos ichneumon parvidens Lonnberg, Arkiv f. Zool., 4: no. 16, p. 3, 29 Apl. 
1908. Mukimbungu, below Manyanga, Lower Congo, Congo Beige. 

Herpestes ichneumon sabiensis Roberts 

Herpestes caffer sabiensis Roberts, Ann. Transvaal Mus., 11 : 249, 14 Sep. 1926. 
Sabi Game Reserve, eastern Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 217 

Herpestes ichneumon sangronizi Cabrera 

Herpestes ichneumon sangronizi Cabrera, Bol. Real Soc. Espafiola Hist. Nat., 
Madrid, 24: 217, 28 Apl. 1924; Trab. Mus. Nac. de Cienc. Nat., Madrid, 
zool. ser., no. 57, p. 151, pi. 6, 30 Dec. 1932. Mogador, Morocco. 
Range: Southwestern Morocco. 

IcHNEUMiA I. Geoffroy. White-tailed Mungooses 

Lasiopus I. Geoffroy, in Gervais's Resume des Legons de Mammalogie, pro- 
fessees au Mus. Paris, 1: 37, 1835. Genotype Herpestes albicaudus G. 
Cuvier (not Lasiopus Dejean, 1833, in Coleoptera). 

Ichneumia I. Geoffroy, Ann. des Sci. Nat., Zool. (2) 8: 251, 1837. New name to 
replace Lasiopus, same genotype. 

Ichneumia albicauda albicauda (G. Cuvier) 

Herpestes albicaudus G. Cuvier, Regne Anim., ed. 2, 1: 158, 1829. Senegal. 

Herpestes leucurus Hemprich and Ehrenberg, Symbolse Physicse, Mamm., dec. 

2, fol. h, i, k, pi. 12, Sep. 1832. Dongola, Anglo-Egyptian Sudan. 
Ichneumia albescens I. Geoffroy, Mag. de Zool., mamm., p. 16 of description 

accompanying pi. 12, 1839. Sennaar; ? Dongola. 
Ichneumia abu-wudan Fitzinger and Heuglin, Sitzb. K. Akad. Wiss. Wien, 

math.-nat. CI., 54: sect. 1, p. 561, 1866. Berbera, Somaliland. 

Ichneumia albicauda dialeucos (Hollister) 

Mungos albicaudus dialeucos Hollister, Smithsonian Misc. Coll., 66: no. 1, 
p. 6, 10 Feb. 1916. Mt. Lololokwi, Kenya Colony. 

Ichneumia albicauda grandis (Thomas) 

Herpestes grandis Thomas, Proc. Zool. Soc. London, for 1889, p. 622, pi. 62 
(skull and teeth), Apl. 1890. ?Limpopo or Zululand, South Africa. 

Ichneumia albicauda haagneri Roberts 

Ichneumia grandis haagneri Roberts, Ann. Transvaal Mus., 10: 68, 31 Jan. 
1924. Bridgewater, Rustenburg district, Transvaal. 

Ichneumia albicauda ibeana (Thomas) 

Herpestes albicaudus ibeanus Thomas,^ Ann. Mag. Nat. Hist. (7) 13: 409, June 

1904. Athi-ya-Maui (Stony Athi), Kenya Colony. 
Mungos albicaudus ferox Heller, Smithsonian Misc. Coll., 61: no. 13, p. 11, 16 

Sep. 1913. Changamwe, Kenya Colony. 

Ichneumia albicauda loand^ (Thomas) 

Herpestes albicaudus loandx Thomas, Ann. Mag. Nat. Hist. (7) 13: 408, 
text-f. b, June 1904. Pungo Andongo, northern Angola. 



218 bulletin: museum of comparative zoology 

IcHNEUMiA ALBiCAUDA LOEMPO (Temminck) 

Herpesies loempo Temminck, Esquisses Zool. sur la Cote de Guine, p. 93, 1853 

(not of Gray, 1865). Guinea Coast. 
Ichneumia nigricauda Pucheran, Rev. et Mag. de Zool. (2) 7: 394, Aug. 1855. 

Senegal. 

MuNGOS E. Geoffrey and G. Cuvier 

Mungos E. Geoffroy and G. Cuvier, Magasin Encyclopedique, 2: 184, 187, 

1795. Genotype Viverra mungo Gmelin. 
Ariela Gray, Proc. Zool. Soc. London, for 1864, p. 565, May 1865. Genotype 

Herpesies tsenionotus (A. Smith). 
Mungos Gray, Proc. Zool. Soc. London, for 1864, p. 575, May 1865. Genotype 

Mungos fasciatus Gray = Herpestes mungo Desmarest. 
Crossarchus auct. recent., not F. Cuvier, (in part). 

Mungos mungo mungo (Gmelin). Banded Mungoose 

Viverra mungo Gmelin, Linnaeus's Syst. Nat., ed. 13, 1: pt. 1, p. 84, 1788. 

Supposed to be Asia, but in reality Africa, taken as Cape of Good Hope 

(see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 155-156, 11 Apl. 1924). 
Herpestes fascdatus Desmarest, Dictionnaire des Sci. Nat., 29: 58, 1823. 

"L'Inde." 
Ichneumon tsenionotus A. Smith, South African Quart. Journ., 2: 114, 1834. 

Natal. 

Mungos mungo bororensis Roberts 

Mungos mungo bororensis Roberts, Ann. Transvaal Mus., 13: 88, 6 July 1929. 
Buruma's Village, Boror, Mozambique. 

Mungos mungo caurinus Thomas 

Mungos caurinus Thomas, Ann. Mag. Nat. Hist. (9) 17: 182, Jan. 1926. 
Gunnal, Portuguese Guinea. 

Mungos mungo colonus (Heller) 

Crossarchus fasciatus colonus Heller, Smithsonian Misc. Coll., 56: no. 17, p. 16, 
28 Feb. 1911. Southern Guaso Nyiro, Sotik district, Kenya Colony. 

Mungos mungo gothneh (Heuglin and Fitzinger) 

Herpestes gothneh Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.- 

nat. CI., 54: sect. 1, p. 560, 1866. Kordofan. 
Herpestes adailensis Heuglin, Nova Acta Acad. Ca^s. Leop.-Carol., Halle, 28: 

no. 10, p. 5, pi. 2, f. 4 (skull), 1861; Petermann's Geogr. Mittheil., p. 17, 

1861. Adail coast. Red Sea. 
Herpestes leucostethicus Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien, 

math.-nat. CI., 54: sect. 1, p. 561, 1866. Junction of the Sobat and Bahr-el- 

Abiad, Anglo-Egyptian Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 219 

MuNGOS MUNGo GRisoNAX Thomas 

Mungos mungo grisonax Thomas, Proc. Zool. Soc. London, p. 294, 29 Apl. 
1926. Otjumbumbi, 15 miles above Cunene Falls, Southwest Africa. 

Mungos mungo macrurus (Thomas) 

Crossarchus fasciatus macrurus Thomas, Ann. Mag. Nat. Hist. (7) 19: 120, 

Jan. 1907. Ruwenzori Southeast, Uganda. 
Crossarchus fasciatus macrosus Lydekker, Zool. Record for 1907, Mammalia, 

p. 61, 1908 (lapsus for macrurus). 

Mungos mungo mandjarum (Schwarz) 

Crossarchus fasciatus mandjarum Schwarz, Jahrb. d. Nassauischen Ver. f. 
Naturk. in Wiesbaden, 68: 63, 1915. Bate, Cameroons. 

Mungos mungo ngamiensis Roberts 

Mungos mungo ngamiensis Roberts, Ann. Transvaal Mus., 15: 5, 1 Oct. 1932. 
Maun, Ngamiland, Bechuanaland, South Africa. 

Mungos mungo pallidipes Roberts 

Mungos mungo pallidipes Roberts, Ann. Transvaal Mus., 13: 69, 6 July 1929. 
Mooivlei, Rustenburg district, Transvaal. 

Mungos mungo rossi Roberts 

Mungos mungo rossi Roberts, Ann. Transvaal Mus., 13: 89, 6 July 1929. Lake 
Bangweolo, Rhodesia. 

Mungos mungo senescens (Thomas and Wroughton) 

Crossarchus fasciatus senescens Thomas and Wroughton, Proc. Zool. Soc. 
London, p. 291, 1 Aug. 1907. Coguno, Inhambane, Mozambique. 

Mungos mungo somalicus (Thomas) 

Crossarchus somalicus Thomas, Ann. Mag. Nat. Hist. (6) 15: 531, June 1895. 
Sunerdorler, Webi Shebeli, SomalUand, 1500 feet. 

Mungos mungo talboti (Thomas and Wroughton) 

Crossarchus talboti Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 
374, May 1908. Bornu, northern Nigeria. 

Mungos mungo zebra (Riippell) 

Herpestes zebra Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 

Saugeth., 1: 30, pi. 9, f. 2; pi. 10, f. 1, 1835. Lowlands of Kulla and at 

Massawa, Eritrea. 

Mungos mxjngo zebroides (Lonnberg) 

Crossarchus zebroides Lonnberg, Arkiv f. Zool., 4: no. 16, p. 7, 29 Apl. 1908. 
Mukimbungu, below Manyanga, Lower Congo. 



220 bulletin: museum of comparative zoology 



MUNGOTICTIS PoCOck 

Mungotictis Pocock, Ann. Mag. Nat. Hist. (8) 16: 120, Aug. 1915. Genotype 
Mungotictis lineatus Focock = Galidictis vittatus Gray, 1848, not Schinz, 
1844. 

Mungotictis lineata Pocock 

Galidictis vittaia Gray, Proc. Zool. Soc. London, p. 22, pi. 1, Mamm., 1 Aug. 
1848. Tulyah Bay, Madagascar (not Galidictis vittatus Schinz, 1844). 

Mungotictis lineatus Pocock, Ann. Mag. Nat. Hist. (8) 16: 506, Dec. 1915 
(new name for Galidictis vittata Gray, 1848, not of Schinz, 1844). Mada- 
gascar. 

Mungotictis substriata Pocock 

Mtmgotictis substriatus Pocock, Ann. Mag. Nat. Hist. (8) 16: 121, pi. 7, f. 4, 
Aug. 1915. Madagascar. 



Myonax Thomas. Black-tipped Mungooses 

Galerella Gray, Proc. Zool. Soc. London, for 1864, p. 509, May 1865 (in part); 

J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 175, 11 Apl. 1924. Genotype 

Herpestes ochraceus Gray. 
Calogale Gray, Proc. Zool. Soc. London, for 1864, p. 509, 560, May 1865 (in 

part); Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 438, 1914 (in part). 

Genotype Herpestes nepalensis Gray. 
Myonax Thomas, Ann. Mag. Nat. Hist. (10) 2: 408, Nov. 1928. Genotype 

Herpestes gracilis Riippell. 

Review: Wroughton, R. C. On the African mungooses usually referred to 
the Herpestes gracilis group. Ann. Mag. Nat. Hist. (7) 20: 110-121, Aug. 1907. 

Schwarz (Ann. Mag. Nat. Hist. (10) 15: 300, Feb. 1935) regards Myonax as 
inseparable from Galerella. 

Myonax cauui cauui (A. Smith) 

Ichneumon cauui A. Smith, Rept. Exped. for Exploring Central Africa, app., 

p. 42, 1836. Kurrichane, northwestern Transvaal. 
Herpestes punctulatus Gray, Proc. Zool. Soc. London, p. 11, 11 Dec. 1849. 

Port Natal, South Africa = Durban, Natal. 
Herpestes ornalus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 81, Feb. 

1852; Reise nach Mossambique, Saugeth., p. 117, pi. 26, 1852. Tette, 

Mozambique. 
Calogale venatica Gray, Proc. Zool. Soc. London, for 1864, p. 563, May 1865. 

"East Africa." 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 221 

Myonax cauui bradfieldi Roberts 

Myonax cauui bradfieldi Roberts, Ann. Transvaal Mus., 15: 2, 1 Oct. 1932. 
Quickborn Farm, 60 miles north of Okahandja, Southwest Africa. 

Myonax cauui caldatus (Thomas) 

Herpestes cauui caldatus Thomas, Proc. Zool. Soc. London, p. 374, 12 July 
1927. Sandfontein, Gobabis-Bechuanaland border, Southwest Africa. 

Myonax cauui kalaharicus Roberts 

Myonax cauui kalaharicus Roberts, Ann. Transvaal Mus., 15: 2, 1 Oct. 1932. 
Gemsbok Pan, Bechuanaland, South Africa. 

Myonax cauui kaokoensis Roberts 

Myonax cauui kaokoensis Roberts, Ann. Transvaal Mus., 15: 2, 1 Oct. 1932. 
Okorosave, Kaokoveld, Southwest Africa. 

Myonax cauui khanensis Roberts 

Myonax cauui khanensis Roberts, Ann. Transvaal Mus., 15: 3, 1 Oct. 1932. 
Khan Mountains, east of Swakopmund, Southwest Africa. 

Myonax cauui lancasteri Roberts 

Myonax cauui lancasteri Roberts, Ann. Transvaal Mus., 15: 4, 1 Oct. 1932. 
Kafue River, Northern Rhodesia. 

Myonax cauui ngamiensis Roberts 

Myonax cauui ngamiensis Roberts, Ann. Transvaal Mus., 15: 3, 1 Oct. 19.32. 
Maun, Ngamiland, Bechuanaland, South Africa. 

Myonax cauui okavangensis Roberts 

Myonax cauui okavangensis Roberts, Ann. Transvaal Mus., 15: 3, 1 Oct. 1932. 
Karakuwise, Grootfontein district, Southwest Africa. 

Myonax cauui sw alius (Thomas) 

Herpestes cauui swalius Thomas, Proc. Zool. Soc. London, p. 292, 29 Apl. 

1926. Great Brukaros Mountain, Great Namaqualand, Southwest 

Africa, 3500 feet. 

Myonax cauui swinnyi (Roberts) 

Mungos caaui (sic) swinnyi Roberts, Ann. Transvaal Mus., 4: 75, 20 Oct. 1913. 
Ngqeleni district, Pondoland, Cape of Good Hope. 

Myonax dentifer (Heller) 

Mungos dentifer Heller, Smithsonian Misc. Coll., 61: no. 13, p. 10, 16 Sep. 
1913. Maji-ya-Chumvi, Kenya Colony. 



222 bulletin: museum of comparative zoology 

Myonax grantii (Gray) 

Calogale grantii Gray, Proc. Zool. Soc. London, for 1864, p. 561, May 1865- 

Mgunda Mkali, Tanganyika Territory. 
Herpestes neumanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 121,^ 

1894. Tisso, northern Ugogo, Tanganyika Territory. 

Myonax ignitus (Roberts) 

Mungos ignitus Roberts, Ann. Transvaal Mus., 4: 76, 20 Oct. 1913. Malava, 
Boror, Mozambique. 

Myonax nigratus nigratus Thomas 

Myonax nigratus Thomas, Ann. Mag. Nat. Hist. (10) 2: 408, Nov. 1928. 
Okorosave, Kaokoland, Southwest Africa. 

Myonax nigratus shortridgei Roberts 

Myonax shortridgei Roberts, Ann. Transvaal Mus., 15: 1, 1 Oct. 1932. Cunene 
Falls, South Africa. 

Myonax pulverulentus pulverulentus (Wagner) 

Herpestes pulverulentus Wagner, Gelehrte Anzeigen d. K. Bayer. Akad. Wiss., 

Miinchen, 9: 426, 12 Sep. 1839. Cape of Good Hope. 
Herpestes apiculatus Gray, Proc. Zool. Soc. London, for 1864, p. 551, May 1865. 

Cape of Good Hope. 

Myonax pulverulentus basuticus Roberts 

Myonax pulverulentus basuticus Roberts, Ann. Transvaal Mus., 18: 253, 21 Feb. 
1936. Senqunyane Valley, Maluti Mountains, Basutoland, 

Myonax pulverulentus maritimus (Roberts) 

Mungos pulverulentus maritimus Roberts, Ann. Transvaal Mus., 6: 114, ApL 
1919. Lambert's Bay, Cape of Good Hope. 

Myonax ratlamuchi ratlamuchi (A. Smith). Ruddy Mungoose 
Ichneumon ratlamuchi A. Smith, Rept. Exped. for Exploring Central Africa, 

app., p. 42, 1836. "Between Latakoo and the Tropic," South Africa. 
Herpestes badius A. Smith, Illustr. Zool. South Africa, Mamm., pi. 4, and text, 
1838. Between Old Latakoo and Kurrichane, 120 miles eastward, Trans- 
vaal, South Africa. 

Myonax ratlamuchi auratus (Thomas and Wroughton) 
Mungos auratus Thomas and Wroughton, Proc. Zool. Soc. London, p. 543, 
30 Oct. 1908. Tette, Mozambique. 

Myonax ratlamuchi ignitoides Roberts 

Myonax auratus ignitoides Roberts, Ann. Transvaal Mus., 15: 2, 1 Oct. 1932. 
Macequece, Mozambique. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 223 

Myonax ratlamuchi rufescens (Lorenz) 

Herpestes ornatus rufescens Lorenz, Abh. Senckenburg. Naturf. Ges., Frankfurt 

a. M., 21: 462, 1898. Island of Zanzibar. 
Mungos melanurus lQ,sti Wroughton, Ann. Mag. Nat. Hist. (7) 20: 114, Aug. 

1907. Zanzibar Island. 

Myonax ratlamuchi upinqtoni Shortridge 

Myonax ratlamuchi upingtoni Shortridge, Mammals of South West Africa, 1 : 
124, footnote, 1934. Between Upington and Louisvale, Orange River. 

Myonax ruddi (Thomas) 

Herpestes ruddi Thomas, Ann. Mag. Nat. Hist. (7) 12: 465, Oct. 1903. Klip- 
fontein. Little Namaqualand, Southwest Africa. 

Myonax sanguineus sanguineus (Riippell) 

Herpestes sanguineus Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 
Saugeth., 1: 27, pi. 8, f. 1; pi. 10, f. 3, 1835. Kordofan. 

Myonax sanguineus canus (Wroughton) 

Mungos melanurus canus Wroughton, Ann. Mag. Nat. Hist. (7) 20: 114, Aug. 
1907. Cape Verde. 

Myonax sanguineus flavescens (Bocage) 

Herpestes gracilis var. flavescens Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, 
(2) 1: 179, Dec. 1889. Benguella, Angola. (Specific reference uncertain.) 

Myonax sanguineus galbus (Wroughton) 

Mungos sanguineus galbus Wroughton, Ann. Mag. Nat. Hist. (8) 3: 514, June 
1909. Mubende, Uganda. 

Myonax sanguineus gracilis (Riippell) 

Herpestes gracilis Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 1 : 

29, pi. 8, f. 2; pi. 10, f. 2, 1835. Near Massawa, Eritrea. 
Ichneumia nigricaudatus Geoffrey, Mag. de Zool., (2) 1: 18, 1839. Abyssinia; 

renaming of Ichneumia gracilis Riippell. 
Herpestes galinieri Guerin, in Ferret and Galinier's Voyage en Abyssinie, 3: 

169, atlas, zool., pi. 1, fig. 1, 1847. Ethiopia. (On the plate the name is 

credited to Pucheran.) 
Herpestes lefebvrii Prevost and Desmurs, in Lefebvre's Voyage en Abyssinie, 

atlas, zool., pi. 1, fig. 1, index, 1850. Abyssinia. 
Herpestes ochromelas Pucheran, Rev. et Mag. de Zool., (2) 7: 393, Aug. 1855. 

Renaming of H. gracilis Riippell, and of Prevost and Desmurs. 
Herpestes jodoprymnus Heuglin, Nova Acta Acad. Caes. Leop.-Carol., Halle, 

29: no. 8, p. 23, 1861. Eastern Abyssinia (Ethiopia). 
Herpestes ruficauda Heuglin, Reise in Nordost-Afrika, 2: 43, 1877. Highlands 

of Wolo-Gala, Abyssinia, 12,000 ft. 



224 bulletin: museum of comparative zoology 

Myonax sanguineus ibe^ (Wroughton) 

Mungos sanguineus ibex Wroughton, Ann. Mag. Nat. Hist. (7) 20: 118, Aug. 

1907. Kikuyu, Kenya Colony. 
Calogale elegans Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 456, Dec. 

1914. Kikuyu (Fort Smith), Kenya Colony. 
Calogale marse Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 453, Dec. 1914. 

On the Ngare Mbusse, a southern tributary of the Ngare Dobasch (Mara), 

between Lake Natron and Lake Victoria, Tanganyika Territory. 

Myonax sanguineus melanurus (Martin) 

Cynictis melanurus Martin, Proc. Zool. Soc. London, p. 56, 13 Sep. 1836. 

Sierra Leone. 
Galerella sanguinea melanura Schwarz, Ann. Mag. Nat. Hist. (10) 15: 300, 

Feb. 1935. 

Myonax sanguineus mustela (Schwarz) 

Galerella sanguinea mustela Schwarz, Ann. Mag. Nat. Hist. (10) 15: 300, Feb. 
1935. Efulen, Ebolowa district, southern Cameroons. 

Myonax sanguineus mutgigella (Riippell) 

Herpestes mutgigella Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 
Saugeth., 1: 29, pi. 9, f. 1, 1835. Dembea and Simen districts, Abyssinia 
(Ethiopia). 

Herpestes fuscus Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 
Saugeth., 1: 31, footnote, pi. 10, f. 4 (skull), 1835. Name given inad- 
vertently to dark phase of H. inutgigella. 

Herpestes mutscheltschela Heuglin, Reise in Nordost-Afrika, 2: 41, 1877 (emen- 
dation). 

Myonax sanguineus orestes (Heller) 

Mungos sanguineus orestes Heller, Smithsonian Misc. Coll., 56: no. 17, p. 15, 
28 Feb. 1911. West slope of Mt. Kenya, Kenya Colony, 8500 ft. 

Myonax sanguineus parvipes (Hollister) 

Mungos sanguineus parvipes Hollister, Smithsonian Misc. Coll., 66: no. 1, p. 5, 
10 Feb. 1916. Kaimosi, Kenya Colony. 

Myonax sanguineus phcenicurus (Thomas) 

Mungos phcenicurus Thomas, Ann. Mag. Nat. Hist. (8) 10: 280, Sep. 1912. 
Panyam, Bauchi Province, northern Nigeria. 

Myonax sanguineus proteus (Thomas) 

Mungos gracilis proteus Thomas, Ann. Mag. Nat. Hist. (7) 19: 119, Jan. 1907. 
Ruwenzori East, Uganda, 7000 ft. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 225 

Myonax sanguineus rendilis (Lonnberg) 

Mungos sanguineus rendilis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 
Stockholm, 48: no. 5, p. 66, pi. 2, f. 3 (skull), 26 June 1912. Thornbush 
country, north of Northern Guaso Nyiro, Kenya Colony. 

Myonax sanguineus SAHARiE (Thomas) 

Herpestes phoenicurus saharx Thomas, Ann. Mag. Nat. Hist. (9) 16: 189, July 
1925. Aouderas, Asben, French Equatorial Africa. 

Myonax sanguineus ugand^ (Wroughton) 

Mungos sanguineus ugandse Wroughton, Ann. Mag. Nat. Hist. (8) 3: 514, June 
1909. Entebbe, Uganda. 

Myonax sanguineus zomb^ (Wroughton) 

Mungos melanurus zomhse Wroughton, Ann. Mag. Nat. Hist. (7) 20: 115, Aug. 
1907. Zomba, Nyasaland. 

The status of the following is uncertain : — 

Calogale bruneo-ochracea Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 451 1 

Dec. 1914. Mengo, Uganda. 
Calogale conradsi Matschie, Sitzb. Ges. Naturf. Freunde, Berlm, p. 454, Dec. 

1914. Ukerewe Id., Lake Victoria, Tanganyika Territory. 
Calogale emini Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 452, Dec. 1914. 

Bukoba, Lake Victoria, Tanganyika Territory. 
Calogale ererensis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 449, Dec. 

1914. Erer River, east of Harar, northern GaUaland, Abyssinia (Ethiopia). 
Calogale erlangeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 445, Dec. 

1914. Near Dadab, hinterland of Zeila, on road to Harar, Abyssinia 

(Ethiopia). 
Calogale flaviventris Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 454, 

Dec. 1914. Tchamliho Mts., north of middle Ruwana River, east side of 

Lake Victoria. 
Calogale lademanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 441, Dec. 

1914. Bubu, near Kondoa Irangi, Tanganyika Territory. 
Calogale mossambica Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 438, 

Dec. 1914. On the Cabaceira, Mozambique. 
Calogale ruasse Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 449, Dec. 1914. 

Mission station of Ruasa near Lake Luhondo, northeast of Lake Elivu, 

Tanganyika Territory. 
Calogale schimperi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 444, Dec. 

1914. Hamedo Plains, Tigre, Abyssinia (Ethiopia). 
Calogale tiirstigi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 446, Dec. 

1914. Uplands of Erkauit, southwest of Suakin, Anglo-Egyptian Sudan. 



226 bulletin: museum of comparative zoology 



Paracynictis Pocock 

Paracynictis Pocock, Ann. Mag. Nat. Hist. (8) 17: 177, Feb. 1916. Genotype 
Cynidis selousi de Winton. 

Paracynictis selousi selousi (De Winton) 

Cynictis selousi De Winton, Ann. Mag. Nat. Hist. (6) 18: 469, Dec. 1896 
(skull); Proc. Zool. Soc. London, 1: 2-3, pi. 1 (colored; exterior), 1 June 
1901. Essex Vale, near Bulawayo, Matabeleland, Southern Rhodesia. 

Paracynictis selousi bechuan^e Roberts 

Paracynictis selousi hechuans Roberts, Ann. Transvaal Mus., 15:5, 1 Oct. 1932. 
Tsessebe Siding, Tati district, Bechuanaland, South Africa. 

Paracynictis selousi ngamiensis Roberts 

Paracynictis selousi ngamiensis Roberts, Ann. Transvaal Mus., 15: 5, 1 Oct. 
1932. 30 miles northwest of Maun, Lake Ngami, Bechuanaland. 

Paracynictis sengaani Roberts 

Paracynictis sengaani Roberts, Ann. Transvaal Mus., 14: 227, pis. 6, 7, 8, ff. a, 
28 July 1931. Maputa, northern Zululand, Natal. 

Rhynchogale Thomas 

Rhynchogale Thomas, Proc. Zool. Soc. London, p. 139, June 1894. Genotype 

Rhinogale melleri Gray. 
Rhinogale Gray, Proc. Zool. Soc. London, for 1864, p. 573, text-f. (cranium), 

May 1865; not of Gloger, 1842. ,Genotype Rhinogale melleri Gray. 

Rhynchogale caniceps Kershaw 

Rhynchogale caniceps Kershaw, Ann. Mag. Nat. Hist. (9) 13: 79, Jan. 1924. 
Near Mbweni, 2.5 miles west of Kilosa, Tanganyika Territory. 

Rhynchogale melleri (Gray) 

Rhinogale melleri Gray, Proc. Zool. Soc. London, for 1864, p. 575, text-f. 
(cranium). May 1865. "East Africa" = Zomba, Nyasaland. 

Salanoia Gray 

Galidia I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 5: 580, 1837; Ann. des Sci. 
Nat., Zool., (2) 8: 251, Oct. 1837 (in part). Genotype Galidia elegans: 
originally included also G. unicolor and G. olivacea, now placed in Salanoia. 

Salanoia Gray, Proc. Zool. Soc. London, for 1864, p. 523, May 1865. As a sub- 
genus of Galidia to include Galidia concolor I. Geoffroy andG. olivacea I. 
Geoffroy. Genotype here designated as Galidia olivacea I. Geoffroy. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 227 

Hemigalidia Mivart, Proc. Zool. Soc. London, p. 143, June 1882. Genotype 
Galidia olivacea I. Geoffroy. 

Salanoia OLIVACEA (I. Geoffroy) 

Galidia olivacea I. Geoffroy, Mag. de Zool. (2) 1: mamm., art. 5, p. 38, pi. 16, 
1839. Madagascar. 

Salanoia unicolor (I. Geoffroy) 

Galidia unicolor I. Geoffroy, Compt. Rend. Acad. Sci. Paris, 5: 581, 1837. 

Madagascar. 
Galidia concolor I. Geoffroy, Mag. de Zool. (2) 1: mamm., art. 5, p. 30, pi. 15, 

1839. Madagascar. 

SuRiCATA Desmarest 

Suricata Desmarest, Dictionnaire d'Hist. Nat., ed. 1,24: Tabl. Methodique d. 

Mamm., p. 15, 1804. Genotype Suricata ca-pensis Desmarest = FiiJerra 

suricaUa Erxleben. 
Ryzsena Illiger, Prodromus Syst. Mamm. et Avium, p. 134, 1811. Includes 

Viverra tetradactyla Schreber and V. zenik Gmelin. 
Rizxna Blainville, Nouv. Dictionnaire d'Hist. Nat., 9: 339, 1817 (misprint). 

No species given. 
Ryssena Lesson, Manuel de Mammalogie, p. 178, 1827 (lapsus). 
Rhyzsena Wagner, Schreber's Saugthiere, Suppl., 2: 330, 1841 (emendation). 

Genotj^pe Rhyzsena ( = Viverra) tetradactyla Schreber. 

Suricata suricatta suricatta (Erxleben). Slender-tailed Mierkat 

Viverra suricatta Er.xleben, Syst. Regni Animalis, p. 488, 1777. "ad caput 

Bona) Spei?" 
Viverra tetradactyla Schreber, Saugthiere, 3: 434, pi. 117, 1778. Cape of Good 

Hope. 
Mus zenik Scopoli, Delicise Faunae et Flor. Insubricae, 2: 84, 1786. No locality; 

based on Sonnerat, Voy. a la Chine, 2: 145, pi. 92, 1782. 
Suricata capensis Desmarest, Dictionnaire d'Hist. Nat., ed. 1, 24: Tabl. 

Methodique d. Mamm., p. 15, 1804. Near Cape of Good Hope. 
Surikata viverrina Desmarest, Nouv. Dictionnaire d'Hist. Nat., ed. 2, 32: 

297, pi. 27, f. 2, 1819. Cape of Good Hope. 
Ryzsena surakatta A. Smith, Descrip. Cat. So. African Mus., p. 32, 1826. 
Rhyzsena typicus A. Smith, South African Quart. Journ., 2: 117, 1834. South 

Africa. 

Suricata suricatta hahni Thomas 

Suricata suricatta hahni Thomas, Proc. Zool. Soc. London, p. 376, 12 July 1927. 
Gobabis, Southwest Africa. 



228 bulletin: museum of comparative zoology 

SuRiCATA SURICATTA HAMiLTONi Thomas and Schwann 

Suricata suricatta hamiltoni Thomas and Schwann, Proc. Zool. Soc. London, 1 : 
133, 1 June 1905. Wakkerstroom, Transvaal. 

Suricata suricatta lophurus Thomas and Schwann 

Suricata suricatta lophurus Thomas and Schwann, Proc. Zool. Soc. London, 1 : 
133, 1 June 1905. Near Grahamstown, Cape Colony. 
Range: Southeastern Cape Colony. 

Suricata suricatta marjori^ Bradfield 

Suricata marjorise Bradfield, Description of New Races of Kalahari Birds and 

Mammals, (privately printed), Benoni, South Africa, p. 1, 26 Sep. 1935; 

The Auk, 53: 131, Jan. 1936 (reprinted). Saltpan, 10 miles north of 

Swakopmund, Southwest Africa. 

Suricata suricatta namaquensis Thomas and Schwann 
Suricata suricatta namaquensis Thomas and Schwann, Proc. Zool. Soc. London, 
1 : 134, 1 June 1905. Klipfontein, Little Namaqualand, Cape Colony. 

Xenogale J. A. Allen 

Xenogale J. A. Allen, Journ. Mammalogy, 1 : 26, 28 Nov. 1919. Genotype 
Xenogale microdon J. A. Allen. 

Xenogale microdon J. A. Allen 

Xenogale microdon J. A. Allen, Journ. Mammalogy, 1 : 27, 28 Nov. 1919; Bull. 

Amer. Mus. Nat. Hist., 47: 199, text-f. 58-60, 61, A, B, pi. 27, 11 Apl. 

1924. Akenge, Niapu, northeastern Congo Beige. 



Family PROTELID^. Aard-wolves 
Proteles I. Geoffrey. Aard-wolf 

Proteles I. Geoffroy, Mem. Mus. d'Hist. Nat., Paris, 11: 355, pi. 20, 1824. 

Genotype Proteles lalandii I. Geoffroy = Fiyerra cristala Sparrman. 
Geocyon Wagler, Naturl. Syst. d. Amphibien, p. 30, 1830. Genotype Proteles 

lalandii I. Geoffroy. 

Proteles cristatus cristatus (Sparrman) 

Viverra cristata Sparrman, Resa Goda-Hopps-Udden, 1 : 581, 1783. Near 

Little Fish River, Somerset East, Cape Colony. 
Viverra hyenoides Desmarest, Encyclop. Method., Mammalogie, Suppl., p. 538, 

1821. Cape of Good Hope. 
Proteles lalandii I. Geoffroy, Mem. Mus. d'Hist. Nat., Paris, 11: 371, pi. 20, 

1824. Near Algoa Bay, Cape of Good Hope. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 229 

Proteles typicus A. Smith, South African Quart. Journ., 2: 96, 1834. South 
Africa. 

Proteles cristatus harrisoni W. Rothschild 

Proteles cristatus harrisoni W. Rothschild, Novitates Zool., 9: 443, July 1902. 
Umpata, Mossamedes district, southern Angola. 

Proteles cristatus pallidior Cabrera 

Proteles cristatus pallidior Cabrera, Ann. Mag. Nat. Hist. (8) 6: 464, Nov, 
1910. Suakin, Nubia. 

Proteles cristatus septentrionalis W. Rothschild 

Proteles cristatus septentrionalis W. Rothschild, Novitates Zool., 9: 443, July 
1902. Somaliland. 

Proteles cristatus termes Heller 

Proteles cristatus termes Heller, Smithsonian Misc. Coll., 61 : no. 13, p. 9, 16 

Sep. 1913. Kabalolot Hill, headwaters of Amala River, west of Loita 

Plains, Kenya Colony. 

Proteles cristatus transvaalensis Roberts 

Proteles cristata transvaalensis Roberts, Ann. Transvaal Mus., 15: 6, 1 Oct. 
1932. Roodekuil, Pretoria, Transvaal. 



Family HYiENIDiE. Hyenas 

Crocuta Kaup. Spotted Hyenas 

Crocuta Kaup, Oken's Isis, 21: column 1145, 1828. Genotype Catiis crocuta 

Erxleben. 
Crocotta Kaup, Skizzirte Entw.-Geschichtl. u. Natlirl. Syst. Europ. Thierwelt, 

1 : 78, 1829. Genotype Canis crocuta Erxleben. 

Crocuta crocuta crocuta (Erxleben) 

Canis crocuta Erxleben, Systema Regni Animalis, p. 578, 1777. ". . . ad caput 

Bonse Spei," = Senegambia (see Cabrera, A., Proc. Zool. Soc. London, p. 95, 

1911). 
Hysena maculata Thunberg, Mem. Acad. Imp. Sci. St.Petersbourg, 3: 302, 1811. 

South Africa. 
Hysena rufa Desmarest, Nouveau Diet. d'Hist. Nat., 15: 499, 1817. Cape of 

Good Hope. Based on G. Cuvier's account in Recherches sur Ossemens 

Foss. 
Hysena capensis Desmarest, Nouveau Diet. d'Hist. Nat., 15: 499, 1817. Cape 

of Good Hope. 



230 bulletin: museum of comparative zoology 

Hysena maculata Odmann, Kongl. Svenska Vet.-Akad. Handl., Stockholm, p. 

70, 1820. Cape of Good Hope (renaming of Erxleben's H. crocuta). 
Hysena encrita A. Smith, Trans. Linn. Soc. London, 15: 461, 1827. Misprint for 

crocuta. 
Hysena cuvieri Boitard, Le Jardin des Plantes, p. 223, 1842. Cape of Good 

Hope. 
Hysena {Crocotta) wissmanni Matschie, Sitzb. Ges. Naturf. Freunde, BerHn, 

p. 22, 1900. Epukiro, Southwest Africa. 
Hysena {Crocotta) gariepensis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 25, 1900. Bamboo Mts., between the Orange, Count Reynett and 

Cradock, north of the Liqua River, South Africa. 
Hysena weissmanni Trouessart, Cat. Mamm. Viv. Foss., part 1, p. 243, 1904 

(errorim). 

Crocuta crocuta fisi Heller 

Crocuta crocuta fisi Heller, Smithsonian Misc. Coll., 61: no. 22, p. 5, 26 Jan. 
1914. Merelle waterholes, Marsabit Road, northern Kenya Colony. 

Crocuta crocuta fortis J. A. Allen 

Crocuta crocuta fortis J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 214, 11 Apl. 
1924. Faradje, northeastern Congo Beige. 

Crocuta crocuta germinans (Matschie) 

Hysena {Crocotta) germinans Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 26, 1900. Lake Rukwa, Tanganyika Territory. 
Crocotta kibonotensis Lonnberg, Wiss. Ergebn. Schwed. Zool. Exp. Kilimandjaro 

1905-06, u. Leitung Sjostedt, Mammals, p. 16, pi. 5, f. 1; pi. 7, f. I, 1908. 

Kibonoto steppe, Kilimanjaro, Tanganyika Territory. 
Crocotta panganensis Lonnberg, Wiss. Ergebn. Schwed. Zool. Exp. Kilimand- 
jaro,. 1905-06, u. Leitung Sjostedt, Mammals, p. 18, pi. 5, f. 2; pi. 7, f. 2, 

1908. Kibonoto, Kilimanjaro, Tanganyika Territory. 
Crocuta nzoyse Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 11: 200, 

Apl. 1911. Nzoia River, Guas Ngishu plateau, Kenya Colony. 
Crocuta nyasse Cabrera, Proc. Zool. Soc. London, p. 99, 22 Mar. 1911. Mt. 

Milanji, southern Nyasaland. 

Crocuta crocuta habessynica (Blainville) 

Hysena crocuta habessynica Blainville, Osteographie, 2: les hyenes, p. 82; atlas, 

pi. 6, fig. of teeth, 1844. Abyssinia (implied). 
Hysena {Crocuta) leontieun Satunin, Zool. Anzeiger, 29: 556, 28 Nov. 1905. 

Abyssinia (=Ethiopia), exact locality unknown. 
Crocuta rufopicta Cabrera, Proc. Zool. Soc. London, p. 97, 22 March 1911. 

Odweina, 100 miles south of Berbera, Somaliland. 
Crocuta thomasi Cabrera, Proc. Zool. Soc. London, p. 98, 22 Mar. 1911. An- 

kole, Uganda. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 231 

Crocuta crocuta thierryi (Matschie) 

Hysena (CrocoUa) thierryi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 30, 

1900. Sansanne Mangu, northern Togo, (now French West Africa). 
Hyaena (CrocoUa) togoensis Matschie, Sitzb. Ges. Naturf. Freunde, BerHn, p. 

31, 1900. Kratyi, or Krachi, Togo (now French West Africa). 
Hyiena (CrocoUa) noUei Matschie, Sitzb. Ges. Naturf. Freunde, BerHn, p. 211, 

2 text-figs., 1900. Upper Sanaga, South Cameroons. 



Hyaena Brisson. Striped Hyenas 

Hyaena Brisson, Regnum Animale in Classes IX Distrib., ed. 2, pp. 13, 169, 
1762. Genotype Hysena hyaena Brisson = Canzs hyaena Linnseus, of India. 

Euhysena Falconer, Palaeontol. Memoirs, 2: 464, 1868. As a subgenus of 
Hyaena; type Canis hyaena Linnseus. 

Review: Pocock, R. I. The races of the Striped and Brown Hyenas. Proc. 
Zool. Soc. London, for 1934, p. 799-825, 14 Jan. 1935. 

HyjEna brunnea brunnea Thunberg. Brown Hyena 

Hyaena brunnea Thunberg, Kongl. Svenska Vet.-Akad. Handl., Stockholm, p. 

59, pi. 2, 1820. Cape of Good Hope. 
Hyaena fusca E. Geoffroy, Dictionnaire Class, d' Hist. Nat., 8: 444, pi. 14, f. 2, 

Sep. 1825. No locality. 
Hyaena villosa A. Smith, Trans. Linn. Soc. London, 15: 461, pi. 19, 1827. South 

Africa. 

Hy^na brunnea melampus Pocock 

Hyaena brunnea melampus Pocock, Proc. Zool. Soc. London, for 1934, p. 824, 
14 Jan. 1935. Otjitundua, central Kaokoveld, Southwest Africa, 4500 ft. 
Range: Kaokoveld, Damaraland, and Upington district, about 500 miles 
inland from coast. Southwest Africa. 

Hy^na HYiENA BARBARA Blainville 

Hyaena vulgaris barbara Blainville, Osteographie, Mamm., 2: les Hyenes, pp. 6, 

81; atlas, pis. 2, 6, 1844. Oran, western Algeria. 
Hyaena suilla FiHppi, Mem. Real Acqad. Sci. Torino, (2) 13: 131, 1853. Locality 

imknown. 

HYiENA HY^NA DUBBAH Meyer 

Hyaena dubbah F. A. A. Meyer, Syst.-summarisch. Uebersicht d. Entdeckun- 

gen in Neu-Holland u. Africa, p. 94, 1793. Atbara, Anglo-Egyptian Sudan; 

based on Bruce's Travels to discover Source of Nile, 5: 115, pi. 26, 1791, 

German ed. 



232 bulletin: museum of comparative zoology 

Cams hysenomelas Desmarest, Encyclop. M^thodique, Mammalogie, p. 215, 

footnote, 1820. Atbara, Anglo-Egyptian Sudan (based on Latreille's 

reference to Bruce in Sonnini's ed. of Buff on, Hist. Nat., 27: 25, 1800). 
Hysma dubia Schinz, Das Thierreich von Cuvier, 4: 509, 1825. Dongola. 
Hyaena hienomelas Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 53, 1900. 

Atbara, Anglo-Egyptian Sudan (based on Bruce's description). 
Hysena (Hyaena) schillingsi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 55, 

1900. West Njiri Swamp, Massi plains, near Kilimanjaro, Tanganyika 

Territory. 
Hyaena (Hyaena) hienomelas bergeri Matschie, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 365, Oct. 1910. Elgeyo Escarpment, east of Sirgoi, Kenya 

Colony. 
Hyaena schillingsi rendilis Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 64, Jan. 1912. 

North of Northern Guaso Nyiro, Kenya Colony. 

Hy^NA hyjEna vulgaris Desmarest 

Hyaena vulgaris Desmarest, Encyclop. Methodique, Mammalogie, p. 215, 1820. 

No exact locality, but assigned by Geoffroy and Cuvier (Hist. Nat. des 

Mamm., 1: no. 146 and text, Sep. 1819) to Egypt. 



Family FELID.^. Cats 

Subfamily CRYPTOPROCTIN.E 

Cryptoprocta Bennett 

Cryptoprocta Bennett, Proc. Zool. Soc. London, p. 46, 24 May 1833. Genotype 
Cryptoprocta ferox Bennett. 

Cryptoprocta ferox Bennett. Le Fosa 

Cryptoprocta ferox Bennett, Proc. Zool. Soc. London, p. 46, 24 May 1833. Mad- 
agascar. 

Cryptoprocta typicus A. Smith, South African Quart. Journ., 2: 134, 1834. Mad- 
agascar. 

Subfamily Feline. Typical Cats 
AciNONYX Brookes. Cheetahs 

Acinonyx Brookes, Cat. Anat. and Zool. Mus. of Joshua Brookes, p. 16, 1828. 

Genotype Acinonyx venator Brookes = /^eZis venatica H. Smith. 
Cynailurus Wagler, Natiirl. Syst. Amphib., p. 30, 1830. Genotype Felis 

jubata Linnseus. 
Guepardus Duvernoy, L'Institut, Paris, 2: 145, 5 May 1834. Genotype not 

specified; includes Guepardus flavus Schreber (pi. 105) and Felis guttata 

Hermann. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 233 

Guepar Boitard, Le Jardin des Plantes, p. 174, 1842. No type specified; in- 
cluded Guepar jubatus (Schreber) and Felis guttata Hermann. 

Cynofelis Lesson, Nouv. Table Regne Anim., Mamm., p. 48, 1842. Included 
Felis jubata Schreber and F. guttata Hermann, of which the latter is 
excluded, leaving the former as genotype. 

Gueparda Gray, List Spec. Mamm. Brit. Mus., p. 46, 13 May 1843. Genotype 
Felis jubata Schreber. 

AciNONYX JUBATUS JUBATUS (Schreber). South African Cheetah 

Felis jubata Schreber, Saugthiere, 3: 392, 586, pi. 105, 1776. Cape of Good 

Hope (see Holhster, Proc. Biol. Soc. Washington, 24: 225, 31 Oct. 1911; 

also Hilzheimer, M., Sitzb. Ges. Naturf. Freunde, Berlin, p. 283-292, 

4 text-f., 1913, who believes Schreber's figure is that of an Indian cheetah). 
Felis fearonii A. Smith, South African Quart. Journ., 2: 245, 1834. Northeast of 

Natal. 
Felis fear onis Fitzinger, Sitzb. K. Akad. Wiss., Wien, math.-nat. CI., 59: sect. 

1, p. 664, 1869. Cape of Good Hope. 
Felis lanea P. L. Sclater, Proc. Zool. Soc. London, p. 532, Oct. 1877. Beaufort 

West, Cape of Good Hope. 

AciNONYX JUBATUS HECKi Hilzheimer 

Acinonyx hecki Hilzheimer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 288, text- 

f. 1, 1913. Senegal, West Africa. 
Felis jubata senegalensis Blainville, Ost^ographie, Felis, atlas, pi. 10, 1843. 

Senegal. Not Felis leo senegalensis von Meyer, 1826. 

AciNONYX JUBATUS NGORONGORENSis Hilzheimer 

Acinonyx guttatus ngorongorensis Hilzheimer, Sitzb. Ges. Naturf. Freunde, 
Berlin, p. 290, text-f. 3, 1913. NgorongorO) south of Lake Natron, Tan- 
ganyika Territory. 

AciNONYX JUBATUS OBERGi Hilzheimer 

'Acinonyx guttatus obergi Hilzheimer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 
289, text-f. 2, 1913. Ketmannshoop, Southwest Africa. 

AciNONYX JUBATUS RAiNEYi Heller 

Acinonyx jubatus raineyi Heller, Smithsonian Misc. Coll., 61: no. 19, p. 9, 8 
Nov. 1913. Ulu Station, Kapiti Plains, Kenya Colony. 

AciNONYX JUBATUS SOEMMERINGII (FitziugCr) 

Cynailurus soemmeringii Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. 

CL, 17: pt. 2, p. 245, 1855. Steppes of Kababish, south of Bajuda Desert, 

Kordofan, Anglo-Egyptian Sudan. 
Felis megabalica Heuglin, Leopoldina, Amtliche Organ K.Leop.-Carol. Deutsch. 

Akad. d. Naturf., 4: no. 3, p. 23, May 1863. West bank of Bahr-el-Abiad, 

Anglo-Egyptian Sudan. 



234 bulletin: museum of comparative zoology 

Acinonyx wagneri Hilzheimer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 285, 
1913. Kordofan (based on description of specimen collected by Riippell, 
in Wagner's Schreber's Saugthiere,. Suppl., 2: 503, 1841). 

AciNONYT JUBATUS VELOx Heller 

Acinonyx jubat us velox Heller, Smithsonian Misc. Coll., 61 : no. 19, p. 7, 8 Nov. 
1913. Agate's, Loita Plains, Kenya Colony. 

Acinonyx rex Pocock 

Acinonyx rex Pocock, Abstr. Proc. Zool. Soc. London, no. 283, p. 18, 1 Mar. 

1927; Proc. Zool. Soc. London, p. 250, pi. 1 (col.), text-f. 6, 8, 6 Apl. 1927. 

Umvukwe Range, northwest of Salisbury, Rhodesia. 
Cabrera (Los mamiferos de Marruecos, p. 191, 1932) regards this as an indi- 
vidual variation of the usual type of Cheetah found in Rhodesia. 

Caracal Gray. Caracals 

Caracal Gray, List Spec. Mamm. Brit. Mus., p. 46 (as Caracala on p. xx), 13 
May 1843 (nomina nuda); Proc. Zool. Soc. London, p. 277, Oct. 1867. 
Genotype Caracal melanotis Gray = Felis caracal Schreber. 

Lynx G. Fischer, Zoognosia, 3: 229, 1814, in part. Genotype Felis lynx Lin- 
naeus. 

Urolynchus Severtzow, Rev. et Mag. de Zool., (2) 10: 389, .Sep. 1858. Geno- 
type Felis caracal Schreber. 

Caracal caracal caracal (Schreber). The Cape Caracal. 

Felis caracal Schreber, Saugthiere, pi. 110, before July, 1776; te.xt, 3: 413, 587, 

1777. Table Mountain, Cape of Good Hope. See Allen, J. A., Bull. Amer. 

Mus. Nat. Hist., 47: 279, 11 Apl. 1924, as to type locality and authority. 
Caracal melanotis Gray, List Spec. Mamm. Brit. Mus., p. 46, 13 May 1843; 

Cat. Carnivorous, Pachyd. and Edentate Mamm. Brit. Mus., p. 38, 1869. 

Substitute for Felis caracal Schreber (the specific name is a barbaric word 

said to mean black-eared). Cape of Good Hope. 

Caracal caracal algirus (A. Wagner) 

Felis caracal P. L. S. Miiller, Linne's Natursystems, Suppl., p. 30, 1776, not 
F. caracal Schreber. "Arabia," but probably near Constantine, Algeria, 
based on "le Caracal" of Buffon (see Allen, J. A., Bull. Amer. Mus. Nat. 
Hist., 47:280, 11 Apl. 1924). 

Felis caracal, var. algira A. Wagner, in M. Wagner, Reisen in d. Regenschaft 
Algier, 3: 76, pi. 4, l841. Algeria. 

Caracal berberorum Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 114, 
1892. Constantine, Algeria. 

Felis (Caracal) berberorum spatzi Matschie, Sitzb. Ges. Naturf. Freunde, Ber- 
lin, p. 61, 1912. Between Feriana and Tebessa, Tunis. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 235 

C[aracal] berberorum medjerdae Matschie, Sitzb. Ges. Naturf . Freunde, Berlin, 
p. 62, 1912. Tunis. 

Felis (Caracal) nubicus corylinus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 63, 1912. TjTJe purchased in Hamburg from a ship sailing from a Mor- 
occo harbor, and supposed to be from Tangiers. 

Caracal caracal colonize Thomas 

Caracal caracal colonise Thomas, Ann. Mag. Nat. Hist. (9) 17: 181, Jan. 1926. 
Deelfontein, central Cape Colony, South Africa. 

Caracal caracal damarensis (Roberts) 

Lynx caracal damarensis Roberts, Ann. Transvaal Mas., 11 : 248, 14 Sep. 1926. 
Quickborn, Okahandja, Damaraland, Southwest Africa. 

Caracal caracal limpopoensis (Roberts) 

Lynx caracal limpopoensis Roberts, Ann. Transvaal Mus., 11: 248, 14 Sep. 
1926. Njellele River, north of Zoutpansberg, near Limpopo River, north- 
■ ern Transvaal. 

Caracal caracal lucani (Rochebrune) 

Lynx lucani Rochebrune, Bull. Soc. Philomathique, Paris, (7) 9: 87, 1885. 
Landana, northern Angola, north of the Congo mouth. 
Possibly valid race. 

Caracal caracal nubicus (J. B. Fischer) 

Felis caracal y nubicus J. B. Fischer, Synopsis Mamm., p. 210, 1829. Nubia 
(implied). 

Caracal caracal pcecilotis Thomas and Hinton 

Caracal caracal pcecilotis Thomas and Hinton, Novitates Zool., 28: 3, May 1921. 
Mt. Baguezan, Asben, northern Nigeria. 

Caracal caracal roothi (Roberts) 

Lynx caracal roothi Roberts, Ann. Transvaal Mus., 11: 247, 14 Sep. 1926. 
Elandshoek, Drakensberg, Barberton district, Transvaal. 



Felis Linnseus 

Note : The old genus Felis Linnaeus was used in a widely inclusive sense, and 
has in later years been variously divided into groups by different authors, some 
of whom give generic, others subgeneric rank to all or certain ones of these 
divisions. At present, a conservative treatment suggests retaining most of the 
groups in a subgeneric sense as members of Felis. See Allen, J. A., Severt- 



236 bulletin: museum of comparative zoology 

zow's classification of the Felidse, Bull. Amer. Mus. Nat. Hist., 41: 335-340,. 
22 Sep. 1919; Pocock, R. I., The classification of existing Felidge, Ann. Mag. 
Nat. Hist. (8) 20: 328-350, Nov. 1917. 



Subgenus Felis Linnaeus 

Felis Linnaeus, Syst. Nat., ed. 10, 1 : 41, 1758. Genotype Felis catus Linnaeus. 
Calus Fitzinger, Wiss.-pop. Nat. Saugethiere, 1 : 265, 1855. Genotype Felis 

catus Linnseus. 
Catolynx Severtzow, Rev. et Mag. de Zool., (2) 10: 389, Sep. 1858. Genotype 

Felis caligata Temminck. 

Felis catus Linnaeus. House Cat 

Felis catus Linnseus, Syst. Nat.,ed. 10, 1 :42, 1758. Upsala, Sweden (see Thomas, 
O., Proc. Zool. Soc. London, p. 136, Mar. 1911). 
Domesticated and now occasionally feral in parts of Africa. 

Felis obscura Desmarest, Encyclop. Methodique, Mammalogie, p. 230, 1820. 
Cape of Good Hope. Status indeterminate; "probably based upon a me- 
lanistic specimen of the domestic cat described as F. torquata, or upon a 
mongrel between that breed and F. ocreatd" (Pocock, R. I.j Proc. Zool. Soc. 
London, p. 669, 9 Oct. 1907). 

Felis caligata, /? hybrida J. B. Fischer, Synopsis Mamm., p. 209, 1829. The 
F. obscura of Desmarest and Chat noir du Cap of F. Cuvier. In Africa 
from Egypt to Cape of Good Hope. 

Felis bouvieri Rochebrune, ex A. Milne-Edwards MS., Actes Soc. Linn. Bor- 
deaux, (4) 7: 126, 1883. Cape Verde Islands; also found in Senegambia. 
Probably a breed of F. catus. 

Felis chaus nilotica De Winton. Swamp Cat 

Felis nippelii (sic) Brandt, Bull. Soc. Imp. des Nat., Moscou, 4: 209, 1832. 

Egypt. (Not Felis riippellii Schinz, 1825, which is a synonym of F. lybica 

ocreata Gmelin.) Based on Felis chaus Riippell, Atlas zu d. Reise im nordl. 

Afrika, pt. 1, Zool., pi. 4, 1826. 
Felis chaus nilotica De Winton, Ann. Mag. Nat. Hist. (7) 2: 292, Oct. 1898. 

Near Cairo, Egypt. 
Range: Nile Delta, south to Qena Province, and west to about 155 miles west 

of Alexandria (Flower, S.S., Proc. Zool. Soc. London, p. 390, 8 July 1932). 

Felis lybica lybica Forster 

Felis lybica Forster, in Buffon's Naturgesch. d. Vierfuss. Thiere, 6: 313, 1780. 

Gafsa, Tunis (name based on Bruce's description; see Cabrera, A., Los 

Mamiferos de Marruecos, p. 167, 1932). 
Felix (sic) lynx lybiensis Kerr, Linnaeus's Animal Kingdom, p. 156, 1792. 

Capsa, Libya. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 237 

Felis cristata Lataste, Actes Soc. Linn. Bordeaux, (4) 9: 229, 1885 (not of 

Falconer and Cautley). Haidra, Tunis. 
Felis lybica cyrenarum Ghigi, Mem. Real. Accad. Sci. 1st. Bologna, cl. di sci. 

fis., (7) 7: 79, Dec. 1920. Cirene, Cyrenaica, Libya. 
Range: Extreme southeastern Morocco, along the northern border of the 

Sahara to southern Tunis (Cabrera, 1932). 

Felis lybica cafra Desmarest 

Felis cafra Desmarest, Encyclop. Method., Mammalogie, Suppl., p. 540, 1822. 
South Africa, the Cape of Good Hope. 

Felis lybica constantina Forster 

Felis constantina Forster, in Buffon's Naturgesch. d. Vierfiiss. Thiere, 6: 313, 

1780. Vicinity of Constantine, Algeria (name based on Buffon's account, 

derived from James Bruce; see Cabrera, A., Los Mamiferosde Marruecos, 

p. 169, pi. 7 (col.), 1932). 
Felis constantinensis Link, Beytr. Naturgesch., 1: pt. 2, p. 91, 1795 (nomen 

nudum). 
Felis caracal, /? algiricus J. B. Fischer, Synopsis Mamm., p. 210, 1829. Algeria 

(implied). 
Felis libyca I. Geoffroy, Descr. d. Collections de V. Jaquemont, Mamm. et 

Oiseaux, p. 56, 1843. Tangiers. 
Felis ocreata mauritana Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., Madrid, 

6: 362, July 1906. Mogador, Morocco. 
Range : Morocco, from at least the Grand Atlas and Atlantic Ocean on the 

west, to northern Algiers and Tunis (Cabrera). 

Felis lybica griselda Thomas 

Felis ocreata griselda Thomas, Ann. Mag. Nat. Hist. (9) 17: 180, Jan. 1926. 
Fifty miles south of Dombe Grande, Benguella, Angola. 

Felis lybica haussa Thomas and Hinton 

Felis haussa Thomas and Hinton, Novitates ZooL, 28: 2, May 1921. Zinder, 
northern Nigeria. 

Felis lybica mellandi Schwann 

Felis ocreata mellandi Schwann, Anti. Mag. Nat. Hist. (7) 13: 423, June 1904. 
Mpika, northeastern Rhodesia. 

Felis lybica namaquana Thomas 

Felis ocreata namaquana Thomas, Ann. Mag. Nat. Hist. (9) 17: 180, Jan. 1926. 
KUpfontein, Little Namaqualand, Southwest Africa. 
Range: High desert portion of northwestern South Africa. 



238 bulletin: museum of comparative zoology 

Felis lybk'a nand.e Heller 

Felis ocreata nandse Heller, Smithsonian Misc. Coll., 61: no. 13, p. 14, 16 Sep. 

1913. Headwaters of Lukosa River, Nandi Escarpment, Kenya Colony, 

7000 feet. 

Felis lybica ocreata Gmelin 

Felis ocreata Gmelin, Anhang zu Bruce, J., Reisen in Africa, 2: 27, 79, 1791. 
Ras-el-Fil, north of Lake Tana, Abyssinia (Ethiopia). 

Felis guttata Hermann, Observationes Zool., p. 38, 1804. Egypt (by implica- 
tion). This description, supplied by the editor. Hammer, is regarded by 
HoUister as probably unidentifiable, and is supposed by Hilzheimer to be 
a serval, but the size, fulvous ears without white bar, and details of pattern 
make it almost certainly F. I. ocreata. 

Felis ruppelii (sic) Schinz, Cuvier's Thierreich, 4: 509, 1824 (not Felis ruppellii 
Brandt, 1832). Dongola. 

Felis maniculata Cretzschmar, in Riippell's Atlas zu d. Reise im Nordl. Afrika, 
Saugeth., p. 1, pi. 1, 1826. Nubia, west side of Nile at Ambukol. 

Felis bubastis Hemprich and Ehrenberg, Symbolse Physicse, Mamm., dec. 2, 
folio ij, Nov. 1832. Egypt (the sacred cat of ancient Egypt). 

Felis dongolana Hemprich and Ehrenberg, Symbolse Physicae, Mamm., dec. 2, 
folio jj, Nov. 1832. Egypt. 

Felis pulchella Gray, Charlesworth's Mag. Nat. Hist., 1 : 577, Nov. 1837. Egypt. 

Felis lybica rubida Schwann 

Felis ocreata rubida Schwann, Ann. Mag. Nat. Hist. (7) 13: 422, 426, June 1904. 
Monbuttu ( = Niangara), Congo Beige. 

Felis lybica rusticana Thomas 

Felis ocreata rusticana Thomas, Ann. Mag. Nat. Hist. (10) 1: 319, Feb. 1928. 
Zuurbron, Wakkerstroom district, Transvaal. 

Felis lybica tait^ Heller 

Felis ocreata taitas Heller, Smithsonian Misc. Coll., 61 : no. 13, p. 14, 16 Sep. 
1913. Voi, Kenya Colony. 

Felis lybica ugand^ Schwann 

Felis ocreata ugandx Schwann, Ann. Mag. Nat. Hist. (7) 13: 424, June 1904, 
Mulema, Uganda, 5000 ft. 

Felis lybica vernayi Roberts 

Fdis ocreata vernayi Roberts, Ann. Transvaal Mus., 15: 6, 1 Oct. 1932. Tsot- 
soroga Pan, northern Bechuanaland, South Africa. 

Felis lybica xanthella Thomas 

Felis ocreata xanthella Thomas, Proc. Zool. Soc. London, p. 291, 29 Apl. 1926. 
Ukualukasi, northwestern Ovamboland, Southwest Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 239 

Felis MARGARITA MARGARITA Loche. Sand Cat 

Felis margarita Loche, Rev. et Mag. de Zool. (2) 10: 49, pi. 1, Feb. 1858. Near 
Negonga, Algeria. 

Felis marginata Gray, Proc. Zool. Soc. London, p. 275, 1867 (errorim). 

Felis margueriUei Trouessart, Causeries Sci. Soc. Zool. de France, 1: 386, Aug. 
1905 (emendation on the ground that the spelling should conform to that 
of General Margueritte for whom the species was named). 
Range: The .AJgerian Sahara, possibly to Senegal. A distinct species, accord- 
ing to S. S. Flower, confined to sandy deserts. 

Felis margarita airensis Pocock 

Felis margarita airensis Pocock, Ann. Mag. Nat. Hist. (11) 1: 472, Apl. 1938. 
In-Abbangarit, west of Air, southwestern Sudan. 

Felis margarita meinertzhageni Pocock 

Felis margarita meinertzhageni Foeock, Ann. Mag. Nat. Hist. (11) 1: 472, 
Apl. 1938. El Golea, Algerian Sahara. 

Subgenus Leptailurus Severtzow. Servals 

Leptailurus Severtzow, Rev. et Mag. de Zool. (2) 10: 389, Sep. 1858. Genotype 

Felis serval Schreber. 
Serval Brehm, Fiihrer Zool. Garten Hamburg, 6 ed., p. 53, 1864. Genotype 

Serval niaculatus. 
Servalina Greve, Nova Acta Acad. Caes. Leop. -Carol., Halle, 63: 76, 1894. 

Genotype Felis serval Schreber. 
Galeopardus Heuglin and Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 

54: sect. 1, p. 557, 1866. Genotype Felis serval Schreber. 

Felis brachyura brachyura Wagner. Small-spotted Serval 

Felis servalina Ogilby, Proc. Zool. Soc. London, p. 94, Nov. 1839. Sierra Leone. 
Not F. servalina Jardine, 1837, see Cabrera, A., Bol. Real Soc. Espahola 
Hist. Nat., Madrid, 10: 427, Nov. 1910. 

Felis brachyura Wagner, Schreber's Saugthiere, SuppL, 2: 547, 1841. Pro- 
posed as a substitute for Felis servalina Ogilby, preoccupied by F. serva- 
lina Jardine, 1837. 

Felis ogilbyi Schinz, Synopsis Mamm., 1: 469, 1844. Sierra Leone. Renaming 
of Felis servalina Ogilby, 1839, not F. servalina Jardine, 1837, a synonym 
of F. ornata. 
Note : This is possibly merely an alternative form of the Serval rather than 

a distinct species. 

Felis brachyura liposticta Pocock 

Felis ogilbyi liposticta Pocock, Proc. Zool. Soc. London, p. 666, 9 Oct. 1907. 
Mombasa, Kenya Colony, but this is doubtless merely the port of ship- 
ment, hence the type may have come "from some place on the Mombasa 
to Uganda Railway," and is probably identical with F. o. pantasticta. 



240 bulletin: museum of comparative zoology 

Felis servalina larseni Thomas, Ann. Mag. Nat. Hist. (8) 12: 91, July 1913. 
Near Bembe, Congo district of northern Angola. Lonnberg (Rev. Zool. 
Africaine, 7: 238, 1920) says this is indistinguishable from liposticta. 

Felis brachyuka pantasticta Pocock 

Felis servalina pantasticta Pocock, Proc. Zool. Soc. London, p. 665, pi. 38, f. 3, 
9 Oct. 1907. Entebbe, Uganda. 

Felis servalina poliotricha Pocock, Proc. Zool. Soc. London, p. 665, 9 Oct. 1907. 
Monbuttu = Niangara, Congo Beige (see Allen, J. A., Bull. Amer. Mus, 
Nat. Hist., 47: 270, 11 Apl. 1924). A gray phase of F. b. liposticta (Lonn- 
berg, E., Rev. Zool. Africaine, 7: 238, 1920). 

Felis serval serval Schreber. Large-spotted Serval 

Felis serval Schreber, Saugthiere, pi. 108, 1776; text, 3: 407, 587, 1777. Cape of 

Good Hope. The name is based on Schreber's plate which is a copy of 

Buffon's plate 34, of Hist. Nat., 13: 1765. 
Felis capensis J. R. Forster, Phil. Trans. Roy. Soc. London, 71: pt. 1, p. 4, pi. 

1, 1781. Cape of Good Hope. 
Felis galeopardus Desmarest, Encyclop. Method., Mammalogie, 1: 227, 1820. 

No locality, name unidentifiable. 

Felis serval beir^ Wroughton 

Felis capensis beiras Wroughton, Ann. Mag. Nat. Hist. (8) 5: 206, Feb. 1910. 
Beira, Mozambique. 

Felis serval faradjia (J. A. Allen) 

Leptailurus serval faradjius J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 267, 
pi. 65-70, 74, f. 1, 11 Apl. 1924. Faradje, northeastern Congo Beige. 

Felis serval ferrarii de Beaux 

Felis capensis ferrarii de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. Nat. 
Milano, 63: 4, July 1924. Giumbo, Italian Somaliland. 
For Capt. G. Ferrari, coll. 

Felis serval hamiltoni (Roberts) 

Leptailurus serval hamiltoni Roberts, Ann. Transvaal Mus., 14: 227, 28 July 
1931. 'Mabbaat River, Kruger National Park, Transvaal, 

Felis serval hindei Wroughton 

Felis capensis hindei Wroughton, Ann. Mag. Nat. Hist. (8) 5: 205, Feb. 1910. 
Machakos, Kenya Colony. 

Felis serval kempi Wroughton 

Felis capenfis kempi Wroughton, Ann. Mag. Nat. Hist. (8) 5: 206, Feb. 1910. 
Kirui, Mt. Elgon, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 241 

Felis serval kivuensis Lonnberg 

Felis serval kivuensis Lonnberg, Rev. Zool. Africaine, 7: 242, 30 Dec. 1919. 
Kivu district, Congo Beige. 

Felis serval limpopoensis (Roberts) 

Leplailurus capensis limpopoensis Roberts, Ann. Transvaal Mus., 11: 248, 14 
Sep. 1926. Fairfield, Rustenburg, Transvaal. 

I 

Felis serval lonnbergi Cabrera 

Felis (Serval) togoensis niger Lonnberg, Zool. Jahrb., Syst., 10: 571, 26 Nov. 

1897. Not Felis nigra Erxleben, 1777, a melanistic jaguar. Kunene River, 

opposite mouth of the Kakulovar, southwestern Angola. 
Felis serval lonnbergi Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 10; 

427, Nov. 1910. Substitute for Felis togoensis niger Lonnberg, not Felis 

nigra Erxleben. 

Felis serval mababiensis (Roberts) 

Leplailurus serval mababiensis Roberts, Ann. Transvaal Mus., 15: 7, 1 Oct. 
1932. Mababe Flats, northern Bechuanaland, South Africa. 

Felis serval phillipsi G. M. Allen 

Felis capensis phillipsi G. M. Allen, Bull. Mus. Comp. Zool., 58: 337, July 1914. 
El Garef, Blue Nile, Anglo-Egyptian Sudan. 

Felis serval pococki Cabrera 

Felis senegalensis Lesson, Mag. de Zool., (2) 1: Mamm., pi. 10 and 2 pp. 

text, 1839. "Les bords du fleuve Senegal." Not Felis lea senegalensis 

von Meyer, 1826. 
Felis serval pococki Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., Madrid, 10: 

427, Nov. 1910. Senegal; name proposed in place of Felis senegalensis 

Lesson, preoccupied by Felis lea senegalensis von Meyer, 1826. 

Felis serval togoensis Matschie 

Felis (Serval) togoensis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p, 109, 
1 Mar. 1893. Bismarckburg, Togo, Dahomey. 



Subgenus Mi(:;rofelis Roberts 

Microfelis Roberts, Ann. Transvaal Mus., 11: 250, 14 Sep. 1926. Genotype 

Felis nigripes Burchell. See Pocock, R. I., Notes upon some African 

species of the genus Felis, based upon specimens recently exhibited in the 

Society's Gardens, Proc. Zool. Soc. London, p. 669-674, 1907. 

Although regarded by Roberts as a full genus, the characters given do not 

seem to be of more than subgeneric value. 



242 bulletin: museum of comparative zoology 

Felis nigripes nigripes Burchell 

Felis nigripes Burchell, Travels in Interior of South Africa, 2: 592, footnote, 
1824. Batlaping, South Africa. 
Range: Sand plains of the Kalahari Desert region. 

Felis nigripes thomasi Shortridge 

Felis (Microfelis) nigripes thomasi Shortridge, Records Albany Mus., Grahams- 
town, 4: 119, pi. 17, 9 Jan. 1931. Thorn Kloof (Carlisle Bridge), near 
Grahamstown, South Africa. 



Subgenus Profelis Severtzow. Golden Cat 

Profelis Severtzow, Rev. et Mag. de Zool. (2) 10: 386, 390, Sep. 1858. Type Felis 
celidogaster Temminck = F. aurata Temminck. 

Chrysailurus Severtzow, Rev. et Mag. de Zool. (2) 10: 389, 390, Sep. 1858. 
Type Felis neglecia Gray = F. aurata Temminck. 

Catopuma Severtzow, Rev. et Mag. de Zool. (2) 10: 387, 390, Sep. 1858 (sub- 
genus of Felis). Type Felis moormensis Hodgson = f. temminckii Vigors 
and Horsfield. 

Felis aurata aurata Temminck 

Felis aurata Temminck, Monographies de Mamm., 1: 120, 1827. No locality. 

Felis chrysothrix Temminck, Monographies de Mamm., 1 : 251, 1827 (substitute 

for F. aurata Temminck, not Rafinesque). 
Felis celidogaster Temminck, Monographies de Mamm., 1: 140, 1827 (the gray 

phase). Said to have come from Chile or Peru but doubtless from West 

Africa. 
Felis neglecta Gray, Ann. (Mag.) Nat. Hist. (1) 1: 27, Mar. 1838. Sierra Leone. 
Felis rutilus Waterhouse, Proc. Zool. Soc. London, for 1842, p. 130, Jan. 1843. 

Mandingo country, Sierra Leone. 

Felis aurata cottoni Lydekker 

Felis chrysothrix cottoni Lydekker, Proc. Zool. Soc. London, for 1906, 992, 994, 
pi. 70, f. 1, 11 Apl. 1907. Ituri Forest, Congo Beige. Probably a valid race 
(see Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 274, 11 Apl. 1924). 



Subgenus Panthera Oken 

Panthera Oken, Lehrbuch d. Naturgesch., pt. 3, Zool., sect. 2, p. 1058, 1816. 

Genotype Panthera vulgaris Oken = Felis pardus Linnajus. 
Leo Oken, Lehrbuch d. Naturgesch., pt. 3, Zool., sect. 2, p. 1070, 1816. Type 

Felis leo Linnaeus. 
Leopardus Gray, Proc. Zool. Soc. London, for 1857, p. 278, 28 Jan. 1858 (not 

of Gray, 1842). Genotype Leopardus hernandesii Gray. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 243 

Jaguarius Severtzow, Rev. et Mag. de Zool., (2) 10: 386, Sep. 1858. Genotype 

Panthera { = Felis) onca Linnaeus. 
Leonina Grave, Nova Acta Acad. Cses. Leop.-Carol., Halle, 63: no. 1, p. 60, 1894, 

Subgenus of Felis, type Felis leo Linnaeus. 

Review: Pocock, R. I. The leopards of Africa. Proc. Zool. Soc. London, 
p. 543-591, pi. 1-4, text-f. 1-9, 8 July 1932. 

Felis leo leo Linnaeus. The Barbary Lion 

Felis leo Linnaeus, Syst. Nat., ed. 10, 1: 41, 1758. "Africa," fixed as Constan- 

tine, Algeria (see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 222, 11 Apl. 

1924). 
Felis leo, race 1, barbaricus J. N. von Meyer, Dissertatio Inaug. Anat.-Med. de 

Genere Felium, p. 6, about 1 Aug. 1826. Barbary. 
Felis leo, a barbarus J. B. Fischer, Synopsis Mamm., p. 197, 1829. Based on 

Cuvier's "Lion de Barbarie" = f rom Constantine, Algeria. 
Leo africanus Jardine, Naturalist's Library, Mamm., 2 (Felinae): 118, 1834. 

"Africa." 
Felis leo nubicus Blainville, Osteographie, Mammiferes, Genus Felis, p. 58; 

atlas, pi. 6, 1843. Nubia. 
Felis leo nigra Loche, Exploration Sci. de I'Algerie, Zool., Mamm., p. 35, 1867. 

Algeria. 
Felis leo var. somaliensis Noack, Jahrbuch. Hamburg. Wiss. Anstalt., 9: pt. 1, 

p. 120, 1891. "Somaliland" (type in Berlin Zool. Garden). Status un- 
certain. 

Felis leo azandicus (J. A. Allen). 

Leo leo azandicus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 224, pi. 37-40, 
11 Apl. 1924. Vankerckhovenville, northeastern Congo Beige. 

Felis leo bleyenberghi Lonnberg 

Felis leo bleyenberghi Lonnberg, Rev. Zool. Africaine, 3: 273, 20 Jan. 1914. 
Katanga, Congo Beige. 

Felis leo hollisteri (J. A. Allen) 

Leo leo hollisteri J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 229, 11 Apl. 1924. 
Lime Springs, Sotik, Kenya Colony. 

Felis leo kamptzi Matschie 

Felis leo kamptzi Matschie, Sitzb. Ges. Naturf. Freimde, Berlin, p. 92, text-f. 
(skuU), 1900. Yoko, upper Sanaga, Cameroons. 

Felis leo krugbri (Roberts) 

Leo leo krugeri Roberts, Ann. Transvaal Mus., 13: 91, 6 July 1929. Brixton, 
Sabi Game Reserve, eastern Transvaal. 



244 bulletin: museum of comparative zoology 

Felis LEO MASSAiCA Neumann 

Felis leo massaicus Neumann, Zool. Jahrb., Syst., 13: 550, 10 Oct. 1900. Kibaya 
and Gurui, Tanganyika Territory. 

Felis leo sabakiensis Lonnberg, in Wiss. Ergebn. d. Schwed. Zool. Exped. nach 
Kilimandjaro, 1905-06, u. Leitung Sjostedt, p. 22, pi. 3, f. 4, 1908. Ki- 
bonoto and Leitokitok, northwest of Kilimanjaro, Tanganyika Territory. 

Felis leo roosevelti Heller, Smithsonian Misc. Coll., 61: no. 19, p. 2, 8 Nov. 1913. 
"Near Addis Ababa," Ethiopia, but "cannot be separated subspecifically" 
from F.l.massaica (Hollister, N., Bull. U. S. Nat. Mus., no. 99, pt. 1, 
p. 165, 1918). 

Felis leo melanochaitus (Hamilton Smith). Cape Lion. 

Felis leo e capensis J. B. Fischer, Synopsis Mamm., addenda, p. 565, 1829. 

Based on the South African Lion of Griffith's Cuvier, Anim. Kingdom, 5: 

163, 1827. South Africa. Not Felis capensis Forster, 1781,= Felis serval 

Schreber. 
Felis (Leo) melanochaitus Hamilton Smith, Jardine's Naturalist's Library, ed. 

2, Mam., 15: 177, 1858. Cape of Good Hope. 
Felis (Leo) inelanochxtus Hamilton Smith, Jardine's Naturalist's Library, ed. 2, 

15: pi. 10, 1858. Cape of Good Hope. 

Felis leo nyanz^ Heller 

Felis leo nyanzse Heller, Smithsonian Misc. Coll., 61: no. 19, p. 4, 8 Nov. 1913. 
Kampala, Uganda, north shore of Lake Victoria. 

Felis leo senegalensis J. N. von Meyer 

Felis leo, race 3, senegalensis J. N. von Meyer, Dissertatio Inaug. Anat.-Med. 

de Genere Felium, p. 6, about 1 Aug. 1826. Senegal. (Not Felis senegalensis 

Lesson, 1839, for a servaline cat). 
Leo gambianus Gray, List Spec. Mamm. Brit. Mus., p. 40, 13 May 1843. 

Interior of Gambia. (Renaming of Felis leo senegalensis Fischer). 

Felis parous pardus Linnaeus. Egyptian Leopard. 

Felis pardus Linnseus, Syst. Nat., ed. 10, 1: 41, 1758. "In Indiis"; but type 
locality fixed by Thomas (Proc. Zool. Soc. London, 1: 135, 1911) as Egypt. 
(See also Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 10: 422, 
1910; J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 248, 11 Apl. 1924). 

Felis fusca F. A. A. Meyer, Zool. Annalen, 1: 394, 1794. Bengal. Pocock 
(Proc. Zool. Soc. London, p. 558, 8 July 1932) writes that he "cannot find 
any differences either in size, colour, pattern, cranial, or dental characters 
between the leopards of India . . . and those of Kenya and Tanganyika," 
and adopts fusca as the subspecific name for those of both areas, with vari- 
ous races intervening. The implication is that the Indian and East 
African leopards in turn are not different from typical pardus. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 245 

Felis leopardus suahelicus Neumann, Zool. Jahrb., Syst., 13: 551, 10 Oct. 1900. 
Lake Manyara district, northern Ugogo, northeastern Tanganyika Terri- 
tory (see HoHister, Bull. U. S. Nat. Mus., no. 99, pt. 1, p. 171, 16 Aug. 1918). 

Felis pardus fortis Heller, Smithsonian Misc. Coll., 61: no. 19, p. 5, 8 Nov. 1913. 
Loita Plains, Southern Guaso Nyiro district, Kenya Colony. 

Felis parous adersi (Pocock). Zanzibar Leopard 

Panthera pardus adersi Pocock, Abstr. Proc. Zool. Soc. London, no. 347, p. 33, 

26 Apl. 1932; Proc. Zool. Soc. London, p. 563, pi. 2, 8 July 1932. Near 

Chuaka, Island of Zanzibar. 

Felis parous aousta (Pocock) 

Panthera pardus adusta Pocock, Ann. Mag. Nat. Hist. (9) 20; 214, Aug. 1927. 

Highlands of Abyssinia (Ethiopia). 
Leopardus poliopardus Brehm, Reise nach Habesch, p. 106, 1863. Not Felis 

poliopardus Fitzinger (Sitzb. K. Akad. Wiss. Wien, math.-nat. Kl., 17: 

295, 1855), said to be a menagerie hybrid between a jaguar and a black 

leopard. 
Panthera nimr, niger Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. Kl., 

58: sect. 1, p. 466, 1868. Abyssinia. Not Felis nigra Erxleben, 1777 (a 

black jaguar). 
Range: Highlands of Ethiopia. 

Felis parous antinorii de Beaux 

Felis nimr Hemprich and Ehrenberg, Symbolse Physicse, Mamm., dec. 2, 

folio gg, pi. 17, Nov. 1832 (in part, as to an Abyssinian skin). Arabia Felix. 
Felis pardus minor Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 199, 1895; 

nomen nudum. Sudan region. Not Felis onca minor Hamilton Smith, 

based on GriflBth, Cuvier's Anim. Kingdom, 2: 456, pL, 1827. 
Felis pardus antinorii de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. Nat. 

Milano, 62: 278, Feb. 1924. Keren, Bogos Country, Eritrea. 
Range: Eritrea and adjacent parts of Abyssinia. 

Felis parous brockmani (Pocock) 

Panthera pardus brockmani Pocock^, Proc. Zool. Soc. London, p. 555, 8 July 
1932. Upper Sheikh in the Golis Range, British Somaliland. 
Range: British Somaliland, "apparently as far south as Jubaland" (Pocock). 

Felis parous chui Heller 

Felis pardus chui Heller, Smithsonian Misc. Coll., 61 : no. 19, p. 6, 8 Nov. 1913. 
Gondokoro, Uganda. 



246 bulletin: museum of comparative zoology 

Felis pardus iturensis (J. A. Allen) 

Felis -pardus centralis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., Stockholm, 

(2) 58: no. 2, p. 49, text-f. 3-5, 1 Sep. 1917. Kabare, Lake Albert Edward, 

Congo Beige. Not Felis centralis Mearns, 1901, a Central American 

jaguar. 
Panthera pardus iturensis J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 259, 11 

Apl. 1924. Niapu, northeastern Congo Beige. 

Felis parous leopardus Schreber 

Felis leopardus Schreber, Saugthiere, pi. 101, 1775; text, 3: 387, 586, 1777. 
Senegal, based on Buffon's account of a leopard "du S6negal ou de Guin^e;" 
see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 250, 11 Apl. 1924. 

Felis poecilura Valenciennes, Compt. Rend. Acad. Sci. Paris, 42: 1036, 1856. 
Gaboon. 

Felis pardus melanotica Giinther. Cape Leopard 

Felis leopardus var. inelanot\ica\ Giinther, Proc. Zool. Soc. London, p. 243, pi. 
16, 1 Aug. 1885; p. 203, text-f., Aug. 1886. Hills, 40 miles from Grahams- 
town, Cape of Good Hope. 

Felis pardus melanosticta Lydekker, Game Animals of Africa, p. 430, 1908 
(errore). Grahamstown district. South Africa. 

Felis parous nanoparous Thomas 

Felis pardus nanopardus Thomas, Ann. Mag. Nat. Hist. (7) 14: 94, Aug. 1904. 
Forty miles west of Gorahai, Somaliland. 
Range: Italian Somaliland. 

Felis parous panthera Schreber. North African Leopard 

Felis panthera Schreber, Saugthiere, 3: 384, 587, 1777; pi. 99, 1776. Algeria; 
based on a copy of Buffon's plate of a female from that country. See 
Cabrera, A., Los Mamiferos de Marruecos, p. 179, 1932. 

Felis palearia F. Cuvier, Hist. Nat. des Mamm., pi. 121 and text, 1832. Al- 
geria. 

Felis pardus barbarus Blainville, Osteographie, 2: "Fehs," p. 186; atlas, pi. 6, 
1843. Barbary coast. Not Felis leo, var. barbarus J. B. Fischer, 1829. 

Felis parous puella (Pocock) 

Panthera pardus puella Pocock, Abstr. Proc. Zool. Soc. London, no. 347, p. 33, 
26 Apl. 1932; Proc. Zool. Soc. London, p. 588, text-f. 9, 8 July 1932. Okoro- 
save, central Kaokoveld, Damaraland, Southwest Africa. 

Felis parous reichenowi (Cabrera) 

Panthera pardus reichenowi Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., 
Madrid, 18: 481, Dec. 1918. Yoko, Cameroons. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 247 

Felis parous ruwenzorii Camerano 

Felis pardus ruwenzorii Camerano, Boll. Mus. Zool. ed Anat. Comp. Univ. 
Torino, 21: no. 545, p. 1, 1906. Bujungolo, Ruwenzori, Uganda, 3800 m. 

Felis pardus shortridgei (Pocock) 

Panthera pardus shortridgei Pocock, Abstr. Proc. Zool. Soc. London, no. 347, 
p. 33, 26 Apl. 1932; Proc. Zool. Soc. London, p. 584, 8 July 1932. Gan- 
gongo, Okavango River, western Caprivi, Damaraland, Southwest Africa. 

Order PINNIPEDIA 

Family OTARIIDiE. Fur Seals and Sea Lions 
Arctocephalus F. Cuvier 

Arctocephalus F. Cuvier, Dictionnaire d. Sci. Nat., 39: 554, 1826. Genotype 

Phoca ursina Linnaeus. 
Ardophoca Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 276, 1866; 

Genotype Otaria philippi Peters. 
Halardus Gill, Proc. Essex Inst., Salem, Communications, 5: 7, 11, July, 1866. 

Genotype Arctocephalus delalandii Gray. 
Euotaria Gray, Ann. Mag. Nat. Hist. (4) 4: 269, Oct. 1869. Genotype not 

specified, but intended for Otaria byronia. 
Gypsophoca Gray, Ann. Mag. Nat. Hist. (4) 4: 269, Oct. 1869. Genotype not 

specified, the Australian Fur Seal. 

Arctocephalus pusillus (Schreber). Cape Fur Seal 

Phoca pusilla Schreber, Saugthiere, 3: 314, 1776; 584, 1777; pi. 86, 1775. "In 

den levantischen, und nach dem Herrn Grafen von Buffon, im indischen 

Meere," but assumed to be South Africa. "The pertinence of the name 

pusilla to the African Fur Seal is not beyond reasonable doubt" (Allen, 

J. A., Hist. North Amer. Pinnipeds, p. 212, 1880). 
Phoca parva Boddsert, Elenchus Anim., p. 172, 1785 (based on Buffon's "Petit 

Phoque." Mediterranean Sea = South Africa. 
Phoca anlarctica Thunberg, Mem. Acad, des Sci. St. Petersbourg, 3: 222, 1811. 
Otaria peronii Desmarest, Encyclop. Method., Mammalogie, p. 250, 1820. 

Cape of Good Hope (Pages); "I'lnde" (Daubenton). 
Arctocephalus delalandii Gray, Proc. Zool. Soc. London, p. 107, pl. 69, Feb.- 

June 1859. Cape of Good Hope. 
Arctocephalus nivosus Gray, Ann. Mag. Nat. Hist. (4) 1: 219, Mar. 1868; Suppl. 

Cat. Seals and Whales Brit. Mus., p. 27, 1871. Cape of Good Hope. 
Arcto-cephalus schist- hyperoes Turner, Journ. Anat. and Phys., 3 = (2) 2: 113, 

text-f., Nov. 1868. Cape of Good Hope. 
lEuotaria compressa Gray, Hand-list Seals, Morses, Sea-lions, and Sea-bears 

Brit. Mus., p. 38, pl. 24, 1874. ?South Africa. 
Range: Coast of South Africa, from Algoa Bay west and north to Cape Cross. 



248 bulletin: museum of comparative zoology 

Family PHOCID.E. Hair Seals 

MoNACHUS Fleming. Monk Seals 

Monachus Fleming, Philosophy of Zool., 2: 187, footnote, 1822. Genotype 
Phoca monachus Hermann. 

Pelagios F. Cuvier, Mem. Mus. d'Hist. Nat., Paris, 11: 196, pi. 2, 1824. Geno- 
type Phoca monachus Hermann. 

Pelagius F. Cuvier, Dictionnaire des Sci. Nat., 39: 550, 1826. Genotype Phoca 
monachus Hermann. 

Heliophoca Gray, Ann. Mag. Nat. Hist. (2) 13: 201, Mar. 1854; Proc. Zool. 
Soc. London, for 1854, p. 43, 1855. Genotype Heliophoca atlantica Gray = 
Monachus monachus (Hermann). 

Monachus monachus (Hermann). Mediterranean Monk Seal 

Phoca monachus Hermann, Beschaftigungen d. Berlin. Ges. Naturf. Freunde, 

4: 501, pi. 12, 13, 1779. Mediterranean Sea. 
Phoca albiventer Boddsert, Elenchus Anim., p. 170, 1785. Adriatic Sea. 
Phoca bicolor Shaw, Gen. Zoology, 1: Mamm., pt. 2, p. 254, 1800. Adriatic 

seacoasts. 
Phoca leucogaster Peron, Voy. aux Terres Austr., 2: 47, footnote, 1817. Nimes, 

France. 
Phoca hermannii Lesson, Dictionnaire Class. d'Hist. Nat., 13: 416, Jan. 1828. 

Adriatic Sea. 
Monachus mediterraneus Nilsson, Kongl. Svenska Vet.-Akad. Handl., Stock- 
holm, for 1837, p. 235, 1838. Adriatic Sea and Greek Archipelago. 
Heliophoca atlantica Gray, Ann. Mag. Nat. Hist. (2) 13: 202, Mar. 1854; Proc. 

Zool. Soc. London, for 1854, p. 43, 10 Jan. 1855. Deserta Grande Island, 

Madeira group. 
Range: Mediterranean Sea and adjacent coasts of Africa, to Madeira and 

Canary Islands. 

MiROUNGA Gray. Elephant Seals 

Macrorhinus F. Cuvier, Dictionnaire des Sci. Nat., 39: 552, art. "Phoques," 
1826. Genotype Phoca proboscidea Peron. Preoccupied by Macrorhinus 
Latreille, 1825, in Coleoptera. 

Mirounga Gray, in Griffith's Cuvier, Animal Kingdom, 5: 179, 1827. As a sub- 
genus of Phoca. No type specified. Included Phoca cristatus Gmelin and 
the Elephant Seals (four species). 

Morunga Gray, List Spec. Mamm. British Mus., pp. xxiii, 103, 13 May 1843 
(misprint? for Mirounga, here made a genus). Genotype Phoca leonina 
Linnaeus. 

Rhinophoca Wagler, Natiirl. Syst. Amphib., p. 27, 1830. Genotype Phoca probo- 
scidea Peron. 



ALLEN : CHECKLIST OF AFRICAN MAMMALS 249 

MiROUNGA LEONiNA (Linnseus). Southern Elephant Seal 

Phoca leonina Linnaeus, Syst. Nat., ed. 10, 1 : 37, 1758. "ad polum Antarcticum." 

Phoca elephantina Molina, Saggio suUa Storia Nat. Chili, p. 280, 1782. "Isole 
di Gio: Fernandes, alle Coste di Araneo, all' Arcipelago di Chiloe, e verso 
lo stretto Magellanico." 

Phoca proboscidea Peron, Voy. aux Terres Austr., 2: 34, pi. 32, 1817. King 
Island, New South Wales. 

Phoca ansonii Desmarest, Encyclopedie M^thodique, Mammalogie, 1 : 239, 1820 
(in part). Falkland Ids. 

Phoca byronii Desmarest, Encyclopedie Methodique, Mammalogie, 1: 240, 
1820. Island of Tinian, Marianne Ids. 

M[irounga] patagonica Gray, in Griffith's Cuvier, Animal Kingdom, 5: 180, 
1827. Patagonia (by implication). 

Phoca dubia J. B. Fischer, Synopsis Mamm., p. 235, 1829. "In insuhs Maluinis" 
= Magellanic region, Falkland Ids. 

Clystophora] falklandica Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 
394, footnote, 1875. Falkland Ids. (by implication). 

Clystophora] kerguelensis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 
p. 394, footnote, 1875. Kerguelen Id. (by implication). 
Range: Southern Ocean, antarctic islands, north in the Pacific to Juan Fer- 
nandez, and formerly in the South Atlantic as far north as Tristan da 
Cunha, South African coast (1926), and island of St. Helena to about 1739 
(seeFraser, F. C, Proc. Linn. Soc. London, for 1934-35, pp. 34-35, 1936). 



Order CETACEA. Whales, Dolphins 
Family DELPHINID.E. Dolphins and Porpoises 

Cephalorhynchus Gray 

Cephalorhynchus Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., 36, pi. 16, 
1846. As a subgenus of Delphinus; no type specified but later fixed as 
Delphinus heavisidii Gray. 

Cephalorhynchus heavisidii (Gray) 

Delphinus {Grampus) heavisidii Gray, Spicilegia Zool., 1: 2, pi. 2, f.6, 1828. 

Cape of Good Hope. 
Delphinus capensis F. Cuvier, Hist. Nat. des Mammiferes, pt. 58, and pi., 

1829. 
D[elphinus] cephalorhyncus F. Cuvier, Hist. Nat. des Cetaces, p. 158, 1836. 

Coast of Cape of Good Hope. 
D[elphinus] hastatus F. Cuvier, Hist. Nat. des C6tac6s, p. 161, 1836. Cape of 

Good Hope. 



250 bulletin: museum of comparative zoology 

Delphinus tridens Van Beneden, ex Sunk MS., Bull. Acad. Roy. Sci., Lettres et 

Beaux-arts Belgique, Bruxelles, (2) 36: 32, pi. 36, f.l, 1873. Cape Town, 

Cape of Good Hope. 
Orca capensis Van Beneden, Bull. Acad. Roy. Sci., Lettres et Beaux-arts 

Belgique, Bruxelles, (2) 36: 32, pi. 36, f. 1, 1873. Cape Town, Cape of 

Good Hope. 

Delphinus Linneeus 

Delphinus Linnseus, Syst. Nat., ed. 10, 1: 77, 1758. Genotype Delphinus 
delphis Linnseus. 

Rhinodelphis Wagner, Schreber's Saugthiere, Suppl., 7: 281, 316, 1846. Sub- 
genus of Delphinus to include the dolphins with a beak. Of the 86 species 
mentioned Delphinus delphis may be selected as the type. 

Delphinus capensis Gray 

Delphinus capensis Gray, Spicilegia Zool., 1: 2, pi. 2, f. 1, 1828. Cape of Good 

Hope. 
D[elphinus]longirostris G. CuvieT,'Regne Animal, ed. 2, 1: 288, 1829 (not of 

Gray, 1828). Malabar coast. 

Delphinus delphis Linnaeus. Common Ocean Dolphin 

Delphinus delphis Linnseus, Syst. Nat., ed. 10, 1: 77, 1758. "In Oceano Euro- 

pseo." 
IDelphinus novse-zealandise Quoy and Gaimard, Voy. Astrolabe, Zool., 1: 149, 

1830. 
Delphinus fulvifasciatus Wagner, Schreber's Saugthiere, Suppl., 7: pi. 361, 

f. 1, 1846. Based on d'Urville's figure. 
Delphinus janira Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 41, pi. 23, 

1846. Newfoundland. 
Delphinus albimanus Peale, U. S. Explor. Exped., ed. 1, 8: Mamm. and 

Ornith., p. 33, pi. 7, f. 1, 1848. Coast of ChUe, 27° 16' S., 75° 30' W. 
Delphinus sao Gray, Cat. Cetacea Brit. Mus., p. 125, 1850; Cat. Seals and 

Whales Brit. Mus., ed. 2, p. 248, 1866. Off Madagascar. 
Delphinus algeriensis Loche, Revue et Mag. de Zool., (2) 12: 474, pi. 22, f. 1, 

Nov. 1860. Coast of Algeria. 
Delphinus moorii Gray, Proc. Zool. Soc. London, for 1865, p. 736, text-f. 1, 

Apl. 1866. South Atlantic, 34° S., 7° 3' W. 
Delphinus moorei Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 396, 1866. 
Delphinus walkeri Gray, Proc. Zool. Soc. London, for 1865, p. 737, text-f. 2, 

Apl. 1866; Cat. Seals and Whales Brit. Mus., ed. 2, p. 397, text-f. 100, 

1866. South Atlantic, 35° 38' S., 0° 10' E. 
Delphinus forsteri Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 42, 

pi. 24, 1846; Cat. Seals and Whales Brit. Mus., ed. 2, p. 248, 1866. Be- 
tween New Caledonia and Norfolk Island. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 251 

Delphinus pomeegra Owen, Trans. Zool. Soc. London, 6: 23, pi. 6, f. 3, 15 Aug. 

1866. Off coast of Madras, India. 
Delphinus major Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 396, 1866. 

No locality. 
Delphinus marginatus Lafont, Actes Soc. Linn. Bordeaux, (3) 6: 518, 15 Sep. 

1868. Coast of Arcachon, dept. Gironde, France. 
Delphinus bairdii Dall, Proc. California Acad. Sci., 5: 12, 29 Jan. 1873. Cape 

Arguello, California. 
Delphinus fusus, souverbianus, variegatus, balteatus, inoschatus Lafont, Fischer, 

P., Actes Soc. Linn. Bordeaux, (4) 5 = 35: 127, 1881. Coast of France. 



Globicephala Lesson. Blackfish 

Globicephala Lesson, Hist. Nat. Gen. et Partic. des Mamm. et Oiseaux decou- 
verts depuis 1788, (Compl. (Euvres de Buffon) 1: [276, pi. 8], 441, 1828. 
Genotype Delphinus deductor Scoreshy = Delphinus melas Traill. 

Globiocephalu^ Gray, List Spec. Mamm. Brit. Mus., p. xxii, 13 May 1843; Zool. 
Voy. Erebus and Terror, 1: Mamm., p. 32, 1846. Genotype not specified; 
three species included. 

Sphxrocephalus Gray, Proc. Zool. Soc. London, p. 244, Nov. 1864; Cat. Seals 

and Whales Brit. Mus., ed. 2, p. 323, text-f. 63, 64, 1866. Subgenus of 

Globiocephalus Gray; type Globiocephalus incrassaius Gr ay = Globicephala 

melsena (Traill). »^44 

Globicephalus Van Beneden and Gervais, Ost^ographie des Cetaces, p. 554, 1777. 
Genotype Globicephala melsena (Traill) . 

Globiceps Flower, Proc. Zool. Soc. London, for 1883, p. 508, Apl. 1884. Geno- 
type Delphinus melas Traill. 

Globicephala mel^na (Traill). North Atlantic Blackfish. 

Delphinus melas Traill, Nicholson's Journ. Nat. Philos., Chem. and Arts, 22: 

81, pi. 3, Feb. 1809. Scapay Bay, Pomona, Orkney Ids. 
Delphinus globiceps G. Cuvier, Ann. Mus. d'Hist. Nat., Paris, 19: 14, pi. 1, 2ff., 

1812. St. Brieux, France. 
Delphinus grinda Lyngbye, Tidskr. f. Naturvidensk., 4: 232, 1825. Faroe 

Islands. 
Delphinus intermedins Harlan, Journ. Acad. Nat. Sci. Philadelphia, (1) 6: 51, 

pi. 1, f. 3, June 1827. Salem Harbor, Mass. Not Delphinus intermedius 

Gray, IS27 = Orcinus orca (LinnjBus). 
Delphinus harlani J. B. Fischer, Synopsis Mamm., p. (456) 656, 1829. Coast of 

New England (substitute for D. intermedius Harlan, not Gray, 1827). 
Globiocephalus affinis Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 32, 

1846. No locality. 
Globiocephalus svineval Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 32, 

1846. East Coast of North America. 



252 bulletin: museum of comparative zoology 

Globioccphalus edwardsii A. Smith, Illiistr. Zool. South Africa, Mamm., p. 127, 
1849. Near Slangkop, Cape of Good Hope. 
Above reference is quoted with description, by Gray, Cat. Seals and Whales 

Brit. Mus., ed. 2, p. 320, 1866, but does not appear in the copy at hand. 

Globiocepholus incrassatus Gray, Proc. Zool. Soc. London for 1861, p. 309, 2 
figs., Apl. 1862. Coast of Dorsetshire, England. 

Globiocephalus propinquus Malm, Kongl. Svenska Vet.-Akad. Handl., Stock- 
holm, (2) 9: pt. 1, art. 2, p. 85, 1871. North Atlantic, near the equator. 

Grampus Gray. The Grampus 

Grampus Gray, Spicilegia Zool., 1: 2, 1828. Subgenus of Delphinus, type Del- 
phinus griseus Cuvier. 

Grampus griseus (G. Cuvier). Gray Grampus 

Delphinus griseus G. Cuvier, Ann. Mus. d'Hist. Nat., Paris, 19: 14, pi. 1, f. 1, 

1812. Brest, France. 
Delphinus rissoanus Desmarest, Encyclop. Methodique, Mammalogie, Suppl., 

p. 519, 1822. Coast of Nice, France (Mediterranean Sea). 
Grampus ciivieri Gray, Ann. Nat. Hist., (1) 17: 85, Feb. 1846. Isle of Wight. 
Grampus sakamata Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 31, 1846. 

Japan coast. 
Grampus richardsoni Gray, Cat. Spec. Mamm. Brit. Mus., Cetacea, p. 85, 1850; 

Proc. Zool. Soc. London, p. 522, Oct. 1865. Kalk's Bay, Simon's Bay, Cape 

of Good Hope. 
Globiocephalus chinensis Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 323, 

1866. China Seas, near Leuchen. 
Grampus stearnsii Dall, Proc. California Acad. Sci., 5: 13, Apl. 1873. Coast of 

California. 
Grampus souverbianus P. Fischer, Actes Soc. Linn. Bordeaux, (4) 5=35: 210, 

1881. No locality. 

Lagenorhynchus Gray 

Lagenorhynchus Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 34, 1846. 

Genotype not specified; included four species. 
Electra Gray, Suppl. Cat. Seals and Whales Brit. Mus., p. 76, 1871. Genotype 

not specified; included eight species. 
Leucopleurus Gray, Suppl. Cat. Seals and Whales Brit. Mus., p. 78, 1871. 

Genotype Leucopleurus arcticus Gray = Lagenorhrjnchus acutus Lesson. 

Lagenorhynchus obscurus (Gray) 

Delphinus {Grampus) obscurus Gray, Spicilegia Zool., 1: 2, pi. 2, f. 2-5, 1828. 

Cape of Good Hope. 
IPhocsena australis Peale, U. S. E.xplor. Exped., ed. 1,8: Mamm. and Ornith., 

p. 33, pi. 6, f. 2, 1848. South Atlantic, off coast of Patagonia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 253 

Lagenorhynchus SUPERCILIOSU8 (Schlegel) 

Delphinus superciliosus Schlegel, Abh. aus d. Gebiete Zool., pt. 1, p. 22, pi. 1, 

f. 3; 2, f. 3 (skull); 4, f. 4 (teeth), 1841. Between Cape Horn and van 

Dieman's Land. 

Note : Although the type of Neomeris phocaenoides (Cuvier) was said to have 
come from the Cape of Good Hope, this was probably not the case. It is not 
certainly known from west of the India peninsula (Allen, G. M., Bull. Mus. 
Comp. Zool., 65: 235, Jan. 1923). 



Orcinus Fitzinger. Orca, Killer Whale 

Orca Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 33, pis. 8, 9, 1846. 

Genotype Delphinus orca Linnaeus. Not Orca Wagler, 1830 = Mesoplodon. 
Orcinus Fitzinger, Wiss.-populare Naturges. Saugethiere, 6: 204, 1860; Palmer, 

Proc. Biol. Soc. Washington, 13: 24, 31 Jan. 1899 (name revived). 
Gladiator Gray, Proc. Zool. Soc. London, p. 71, June 1870. Subgenus of Orca ( = 

Orcinus); type Orca stenorhijncha Gray = Delphinus orca Linnseus. 

Orcinus orca (Linnseus) 

Delphinus orca Linnaeus, Syst. Nat., ed. 10, 1 : 77, 1758. "In Oceano Europaeo." 

Delphinus gladiator Bonnaterre, Tabl. Encyclopedique et Method., Cetologie, 

p. 23, 1789. Spitsbergen, Davis Straits, New England Coast. 
Delphinus duhameli Lacepede, Hist. Nat. des Cetacees, p. 314, 1804 (large 8vO' 

ed.). Mouth of the Loire, France. 
Orca capensis Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 34, pi. 9, 1846 

(not of Synopsis, 1868). Cape of Good Hope. 
Delphinus victorini Grill, Kongl. Svenska Vet.-Akad. Handl., Stockholm, (2) 2- 

no. 10, p. 21, pi. 1, 1858. Knysna River, west of Cape Town, South Africa- 
0[rca] schlegelii Lilljeborg, Recent Memoirs on Cetacea, Ray Society, p. 235, 

1866. West coast of Norway. 
Orca eschrichtii Steenstrup, in note by J. Reinhardt, Recent Memoirs on the 

Cetacea, Ray Soc, p. 188, text-f., 1866. Faroe Islands, Kollefjord, on 

Stromo. 
Orca magellanica Burmeister, Ann. -Mag. Nat. Hist. (3) 18: 101, pi. 9, f. 5, 

Aug. 1866; Anales Mus. PubHco Buenos Aires, 1:373, pi. 22, f. 1-3, 1869. 

South of Cape Corrientes, Argentina, mouth of Rio del Cristiano, lat. 38° 

50' S. 
Orca rectipinna Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 22, f. 15, 16, Apl. 

1869. California, southwards. 
Orca atra Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 22, 58, f. 17, Apl. 1869. 

Oregon to Aleutian Islands. 



254 bulletin: museum of compakative zoology 

Orca padfica Gray, Proc. Zool. Soc. London, p. 76, June 1870, based on the Orca 
(Ophysia) capensis Gray, in part, of Synopsis of Whales and Dolphins, p. 8, 
pi. 9, 1868. North Pacific. 

Orca stenorhyncha Gray, Proc. Zool. Soc. London, p. 74, text-f. 1, 3, June 1870. 
North Sea, Weymouth, England. 

Orca latirostris Gray, Proc. Zool. Soc. London, p. 76, June 1870. North Sea. 

Orca africana Gray, Suppl. Cat. Seals and Whales Brit. Mus., p. 90, 1871. 
Algoa Bay, South Africa. 

Orca ater, var. fusca Dall, in Scammon's Marine Mamm. North-west Coast 
North Amer., app., p. 298, pi. 17, f. 3, 1874. Coast of California and Ore- 
gon. 

Orca antarctica P. Fischer, Journ. de Zool., 5: 146, 1876. Powell and South 
Shetland Islands, Antarctic Ocean. 

Orca australis Van Beneden and Gervais, Osteographie des Cetaces, p. 540, pi. 
47, f. 1, 2, 1877. Tasmania (f. 1); Algoa Bay, Cape of Good Hope (f. 2). 

Orca gladiator (arcticus) et {europseus) Gervais and Van Beneden, Osteogr. des 
Cetaces, pi. 47, f. 3, 5, 1877. Faroe Islands and Atlantic Ocean (resp.). 

Orca gladiator var., australis Gervais and Van Beneden, Osteogr. des C6tac68, 
p. 540, pi. 47, f. 1, 1877. Tasmania. 

Orca tasmanica Gervais and Van Beneden, Osteogr. des C^tac^s, p. 543, 1877. 
Tasmania. 

Orca minor Trouessart, Cat. Mamm. Viv. Foss., ed. 1, 2: pt. 5, p. 1049, 1898, 
here the name is credited to Reinhardt, Ray Society, Memoirs on Recent 
Cetacea, where, however, only the description of a smaller variety occurs, 
in the paper by Eschricht, p. 179, in translation, without a name. 



Phoc^na G. Cuvier. Harbor Porpoises 

Phocsena G. Cuvier, Regne Animal, ed. 1, 1: 279, 1817. Genotype Delphinus 
phocuena Linnaeus. 

Phoc^ena phoccena (Linnaeus). Harbor Porpoise 

Delphinus phoccena Linnaeus, Syst. Nat., ed. 10, 1: 77, 1758. Swedish seas 
(Thomas); "In Oceano Europseo, & Balthico." 

Phocsena communis Lesson, Manuel de Mammalogie, p. 413, 1827. Atlantic 
Ocean, and French estuaries of la Loire, la Charente, la Seine. 

Phocsena homeii A. Smith, Zool. Journ., 4: 440, May 1829. South Africa, Table 
Bay and seas about Cape of Good Hope. 

Phocaena vomerina Gill, Proc. Acad. Nat. Sci. Philadelphia, p. 178, 1865. Puget 
Sound, Washington, and San Francisco, California. 

Plwcaena brachycium Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 279, 1865. 
Salem Harbor, Massachusetts. 

Phocsena lineata Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 135, 1876. East- 
port, Maine. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 255 



Prodelphinus Gervais. Ocean Dolphins 

Clymene Gray, Proc. Zool. Soc. London, p. 237, Nov. 1864. Subgenus of Del- 
phinus, type Delphinus euphrosyne Gray, raised to generic rank by Gray, 
Proc. Zool. Soc. London, p. 214, Sep. 1866. Preoccupied by Clymene Oken, 
1815, in mollusks. 

Sienella Gray, Proc. Zool. Soc. London, p. 213, Sep. 1866. As a subgenus of 
Steno; type Steno attenuatus Gray. 

Clymenia Gray, Synopsis of Whales and Dolphins, p. 6, 1868. No type speci- 
fied; eleven species included. Preoccupied by Clymenia Munster, 1839, in 
mollusks. 

Prodelphinus Gervais, Osteographie des Cetaces, p. 604, 1877. Genotype not 
specified; three species included. Substitute for Clymenia Gray. 

Prodelphinus attenuatus (Gray) 

Delphinus pseudodelphis Wiegmann, in Schreber's Saugthiere, 7: pi. 358, 

1837(?). 
Steno attenuatus Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 44, pi. 28, 

1846. No locality. 
Steno capensis Gray, Proc. Zool. Soc. London, p. 522, Oct. 1865. Cape of Good 

Hope. 
Clymene punctata Grayj Proc. Zool. Soc. London, for 1865, p. 738, text-f. 3, 

Apl. 1866. Lat. 16° 40' N., long. 21° W. 

Prodelphinus euphrosyne (Gray) 

Delphinus styx Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 39, pi. 21, 

1846. West coast of Africa. 
Delphinus euphrosyne Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 40, 

pi. 22, 1846. No locality. 
Delphinus tethyos Gervais, Bull. Soc. d'Agric. Herault, 40: 150, pi. 1, f. 1-4, 

1853. Mouth of the Orb, Herault, France. 
Delphinus marginatus Duvernoy, in Pucheran, Rev. et Mag. de Zool. (2) 8: 

545, pi. 25, Dec. 1856. Near Dieppe, France. 
Tursio dorcides Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 400, 1866. No 

locality. 
Clymene dorides Gray, Proc. Zool. Soc. London, p. 214, Sep. 1866. No locality. 
Clymene similis Gray, Proc. Zool. So^c. London, p. 146, text-f. 2, May 1868. 

Cape of Good Hope. 
Clymenia euphrosynoides Gray, Synopsis Whales and Dolphins, p. 6, 1868; 

based on Owen, Trans. Zool. Soc. London, 6: pi. 8, f. 5, 1866. No locality. 

Prodelphinus fr.enatus (F. Cuvier). Bridled Dolphin 
D[elphinus] frontalis G. Cuvier, ex Dussumier, Regne Anim., ed. 2, 1: 288, 1829. 
Cape Verde Islands. 



256 bulletin: museum of comparative zoology 

Delphinus frsenatus F. Cuvier, Hist. Nat. des C6tac6s, p. 155, pi. 10, f. 1, 1836. 

Cape Verde. 
Delphinus doris Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 39, pi. 20, 

1846. No locality. 
Delphinus clymene Gray, Cat. Spec. Mamm. Brit. Mus., Cetacea, p. 115, 1850; 

Cat. Seals and Whales Brit. Mus., ed. 2, p. 249, 1866. No locality. 
IDelphinus dubius G. Cuvier, Ann. Mus. d'Hist. Nat., Paris, 19: 14, 1812. No 

locality. 
Delphinus mediterraneus Loche, Revue et Mag. de Zool. (2) 12: 475, pi. 22, f. 2, 

Nov. 1860. Coast of Algeria. 
Clymenia normalis Gray, Proc. Zool. Soc. London, p. 214, Sep. 1866. No 

locality, 

Prodelphinus longirostris (Gray) 

Delphinus longirostris Gray, Spicilegia Zool., 1:1, 1828. No locality. 

Delphinus microps Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 42, pi. 

25, 1846. No locality. 
Delphinus alope Gray, Cat. Spec. Mamm. Brit. Mus., Cetacea, p. 118, 1850. 

No locality. 
Delphinus stenorhynchus Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 396, 

1866. No locality. 

PsEUDORCA Reinhardt. False Killer 

Pseudorca Reinhardt, Overs. K. Danske Vid. Selsk. Forhandl., Kjobenhavn 
p. 151, 1862. Genotype Phocxna crassidens Owen. 

Pseudorca crassidens (Owen) 

Phocsena crassidens Owen, British Fossil Mammals and Birds, p. 516, 1846. 

Lincolnshire Fens, near Stamford, England (subfossil). 
Orca meridionalis Flower, Proc. Zool. Soc. London, for 1864, p. 420, text-f. 1, 2, 

May 1865. Tasmania. 
Orca destructor Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 293, Oct. 1866. Off 

Paita, Peru. 
Globicephalus grayi Burmeister, Anales Mus. Pub. Buenos Aires, 1: 373, pi. 21, 

(f. 3-6 but not f. 1, 2), 1868. Coast of Argentina south of the La Plata 

River. 

SoTALiA Gray. White Dolphins 

Sotalia Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 401, 1866. Genotype 
Delphinus guianensis Van Beneden. 

Sotalia teuszii Kiikenthal 

Sotalia teuszii Kiikenthal, Zool. Jahrb., Syst., 6: 442, pi. 21 (cranium), 1892. 
Bay of Warships anchorage, Cameroons, West Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 257 

Steno Gray 

Steno Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 30, 43, pi. 26, f. 1; 

27, 28, 1846. Genotype Delphinus rostratus G. Cuvier. 
Stenopontistes Miranda-Ribeiro, Bol. Museu Nac, Rio de Janeiro, 12: 19, 42, 

Mar. 1936. Genotype Stenopontistes zamhezicus Miranda-Ribeiro = 

Steno rostratus (G. Cuvier). 

Steno rostratus (G. Cuvier) 

Delphinus rostratus G. Cuvier, in Desmarest, Nouv. Dictionnaire d'Hist. Nat., 

ed. 2, 9: 160, 1817. Paimpol, France. 
Delphinus pernettyi Desmarest, Encyclop. Method., Mammalogie, Suppl., p. 

513, 1822. No locality. 
Delphinus bredanensis G. Cuvier, Lesson, Hist. Nat. Gen. et Partic. d. Mamm. 

et Oiseaux decouverts depuis 1788, (Compl. Oeuvres de Buff on, 1) C6tac6s, 

p. 206, 1828. Coast of France. 
[Delphinus] frontatus G. Cuvier, Recherches sur les Ossemens Foss., ed. 2, 5: 

pt. 1, 278, 1823 (in part). Lisbon, Portugal. 
Delphinus planiceps Schlegel, Abh. aus d. Gebiete Zool., pt. 1, p. 27, pi. 4, f. 8, 

1841 (not Van Breda). Dutch coast. 
Delphinus reinwardtii Schlegel, Abh. aus d. Gebiete Zool., pt. 1, p. 27, pi. 3, f. 2, 

3; pi. 4, f. 7, 1841. Indian archipelago. 
Steno co7npressus Gray, Zool. Voy. Erebus and Terror, 1: Mamm., 43, pi. 27, 

1846. No locality. 
Stenopontistes zamhezicus Miranda-Ribeiro, Bol. Museu Nac, Rio de Janeiro, 

12: 20, 42, pi.. Mar. 1936. Zambesi, southeastern Africa. 

TuRSioPS Gervais. Bottle-nose Porpoises 

Tursiops Gervais, Hist. Nat. des Mamm., 2: 323, 1855. Genotype Delphinus 
tursio Fabricius. 

Review: True, F. W. On Tursiops catalania and other existing species of 
Bottlenose Porpoises of that genus. Ann. Durban Mus., 1: 10-24, 1 June 1914. 

Tursiops abusalam (Riippell) 

Delphinus aduncus Hemprich and Ehrenberg, Symbolse Physicse, Mamm., 
dec. 2, folio k, footnote, Sep. 1832. Belhosse Island, Red Sea; "highly 
probable that this species is the same as T. abusalam" — ^True. 

Delphinus abusalam Riippell, Mus. Senckenberg., 3: 140, pi. 12, f. 1-6, 1842. 
Red Sea. 

Tursiops catalania (Gray) 

Delphinus catalania Gray, Proc. Zool. Soc. London, p. 143, Sep. 1862. Cape 
Melville and Cape Flattery, northern Australia. 
Range: Southern Oceans, from South Africa to Australia. 



258 bulletin: museum of comparative zoology 

TuRSioPS TRUNCATUS (Montagu). Common Bottle-nose Porpoise 
Delphinus tursio O. Fabricius, Fauna Grcenlandica, p. 49, 1780 (in part). 

South Greenland seas. 
Delphinus truncatus Montagu, Mem. Wernerian Nat. Hist. Soc, Edinburgh, 

3: 75, pi. 3, 1821. Duncannon Pool, near Stoke Gabriel, 5 miles up the 

river Dart, Scotland. 
Delphinus compressicauda Lesson, Hist. Nat. Gen. et Partic. d. Mamm. et 

Oiseaux decouverts depuis 1788, Cetaces, (Compl. des Oeuvres de Buffon, 

1), p. 272, 1828 (on p. 441 it appears as Phocaena compressicauda). Atlantic 

Ocean, near the equator, 4° S., 26° W. 
Delphinus eurynome Gray, Zool. Voy. Erebus and Terror, 1 : Mamm., p. 38, 

pi. 17, 1846. No locality. 
Delphinus metis Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 38, pi. 18, 

1846. No locaHty. 
Delphinus cymodoce Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 38, 

pi. 19, 1846. No locality. 



Family KOGIID^E. Pygmy Sperm Whales 
KoGiA Gray. Pygmy Sperm Whale 

Kogia Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 22, 1846. Genotype 

Physeter breviceps Blainville. 
Euphysetes Wall, Mem. Australian Mus., Sydney, no. 1, p. 46, 1851. Genotype 

Euphysetes grayii Wall = Kogia breviceps (Blainville) . 
Callignathus Gill, Amer. NaturaUst, 4: 737-738, 740, footnote, Feb. 1871. 

Genotype Physeter simus Owen. Not Calignathus Costa, 1853, in fishes. 

Kogia breviceps (Blainville) 

Physeter breviceps Blainville, Ann. frang. et etrang. d'Anat. et de Physiol., 2: 

337, 1838. Cape of Good Hope. 
Euphysetes grayii Wall, Mem. Australian Mus., Sydney, no. 1, p. 37, pi. 2, 

1851; reprint 1887. Maroubra Beach, New South Wales. 
Euphysetes macleayi Krefft, Proc. Zool. Soc. London, for 1865, p. 713, text-f. 

1-6, Apl. 1866. Manly Beach, New South Wales. 
Physeter (Euphysetes) siinus Owen, Trans. Zool. Soc. London, 6: 30, pi. 10-14, 

15 Aug. 1866. Waltair, Madras Presidency, India. 
Kogia floweri Gill, Amer. Nat., 4: 736, 738, text-f. 167, Feb. 1871. Off Mazat- 

lan, Sinaloa, Mexico. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 259 

Family PHYSETERID^. Sperm Whales 

Physeter Linnaeus. Sperm Whale 

Physeter Linnaeus, Syst. Nat., ed. 10, 1: 76, 1758. Genotype Physeter catodon 

Linnseus. 
Catodon Linnaeus, Fauna Suecica, ed. 2, p. 18, 1761. Genotype Catodon 

macrocephalus Linnaeus = Physeter catodon Linnaeus. 
Physalus Lacepede, Hist. Nat. des Cetacees, p. xl, 219, 1804. Genotype 

Physalus cylindricus Lacepede = Physeter catodon Linnaeus. 
Meganeuron Gray, Proc. Zool. Soc. London, p. 440, Oct. 1865. Genotype 

Catodon (Meganeuron) krefftii Gray = Physeter catodon Linnaeus. 

Physeter catodon Linnaeus. Sperm Whale; Cachalot 

Physeter catodon Linnaeus, Syst. Nat., ed. 10, 1: 76, 1758. "In Oceano sep- 

tentrionali" =Kairston, Orkney Ids. (see Thomas, O., Proc. Zool. Soc. 

London, p. 157, 22 Mar. 1911). 
Physeter macrocephalus Linnaeus, Syst. Nat., ed. 10, 1: 76, 1758. "In Oceano 

Europaeo." 
Physeter microps Linnaeus, Syst. Nat., ed. 10, 1: 76, 1758. "In Oceano septen- 

trionali." 
Physeter tursio Linnaeus, Syst. Nat., ed. 10, 1: 77, 1758. "In Oceano septen- 

trionali." 
Catodon (Meganeuron) krefftii Gray, Proc. Zool. Soc. London, p. 440, text-f. 

1-4, Oct. 1865. Australian Seas. 
Meganeuron krefftii Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 389, 

text-f. 94-97, 1866. Australian seas. 
trumpo, gibbosus, cylindricus, orthodon, sulcatus Lacepede et al., are some 
syns. of P. catodon. 

also, polycyphus Quoy and Gaimard, Zool. Voy. Uranie, p. 77, pi. 12, 1824. 



Family ZIPHIIDiE. Ziphioid Whales 

Review: True, Frederick W. An account of the beaked whales of the family 
Ziphiidae in the collection of the United States National Museum, with remarks 
on some specimens in other American museums. Bull. U. S. Nat. Mus., no. 73 
(quarto), 89 pp., 42 pis., 1910. 

Hyperoodon Lacepede. Bottle-nose Whale 

Hyperoodon Lacepede, Hist. Nat. des Cetacees, pp. xliv, 319, 1804. Genotype 
Hyperoodon butskopf Lacepede = /f. ampullatus (Forster). 

Anarnak Lacepede, Hist. Nat. des Cetacees, pp. xxxviii, 164, 1804. Genotype 
Anarnak graenlandicus Ija,cepede= Hyperoodon ampullatus (Forster). 
Also various emendations: Anarnacus, Anarcus, Ananareus, Anarmacus. 



260 bulletin: museum of comparative zoology 

Ancylodon Illiger, Prodromus Syst. Mamm. et Avium, p. 142, 1811. Genotype 

Monodon spurius O. Fahricius = Hyperoodon ampullatus (Forster). 
Uranodon Illiger, Prodromus Syst. Mamm. et Avium, p. 143, 1811. Genotype 

Delphinus butzkopf Bonnaterre. 
Heterodon Blainville, Nouv. Dictionnaire d'Hist. Nat., ed. 2, 9: 175, 1817. Type 

not specified, eight species included; as a subgenus of Delphinus. Not 

Heterodon Beauvois, 1800, for reptiles. 
Ceto-diodon Jacob, Dublin Philos. Journ. and Sci. Rev., 1: 72, pi. [3], fig. 8, 

Mar. 1825. Genotype Ceto-diodon hunteri Jacob = Hyperoodon ampullatus 

(Forster). 
Nodus Wagler, Natiirl. Syst. der Amphibien, p. 34, 1830. Genotype Delphinus 

edentulus Schreher = H yperoodon ampullatus (Forster). 
Chsenodelphinus Eschricht, Forhandl. Skandinav. Naturforsk., 3die Mote, 

Stockholm, p. 651, 1843. Genotype Balsena rostrata Muller = Hyperoodon 

ampullatus (Forster). 
Chsenocetus Eschricht, Overs. K. Dansk. Vidensk. Selsk. Forhandl., Kjoben- 

havn, for 1845, p. 17, 1846. Genotype the "Nsebhval" or Hyperoodon 

ampullatus (Forster). 
Lagenocetus Gray, Proc. Zool. Soc. London, p. 200, Oct. 1863. Genotype Lagen- 

ocetus latifrons Gray = Hyperoodon ampullatus (Forster). Misspelled Lago- 

cetus by Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 82, 104, 1866. 

Hyperoodon ampullatus (Forster). Bottle-nose Whale 

Balxna ampullaia Forster, in Kalm's Travels into North America, 1: 18, foot- 
note, 1770. Based on Pennant's account. 

B[alsena] rostrata O. F. Muller, Zool. Danicse Prodr., p. 7, 1776. No locality; 
Danish and Norwegian seas implied. 

Monodon spurius O. Fabricius, Fauna Grcenlandica, p. 31, 1780. Greenland 
Seas. 

D[elphinus] bidentatus Bonnaterre, Tableau Encyclop. et Methodique des Trois 
Regncs de la Nature, Cetologie, p. 25, pi. 11, f. 3, 1789. Thames River, 
England (based on Hunter's account). 

D[elphinus] butskopf Bonnaterre, Tabl. Encyclop. et Method, des Trois Regnea 
de la Nature, Cetologie, p. 25, 1789. Honfleur, France. 

Delphinus edentulus Schreher, Saugthiere, 7: 360, 1846; pi. 347; 348, f. 1, 1802. 
North Atlantic. 

Anarnak grcenlandicus Lacepede, Hist. Nat. des Cetacees, p. 164, 1804. Green- 
land seas. 

Delphinus diodon Lacepede, Hist. Nat. des Cetacees, p. 309, pi. 13, f. 3, 1804. 
Near London, England. 

Delphinus bidens Turton, British Faima, p. 17, 1807, not of Sowerby. No 
locality; based on Hunter's account. 

Delphinus chemnitzianus Blainville, in Desmarest, Nouv. Dictionnaire d'Hist. 
Nat., ed. 2, 9: 175, 1817. No locality. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 261 

Delphinus anarnacus Desmarest, Encyclop. Methodique, Mammalogie, 2: 520, 

1822. Greenland Seas. 
Delphinus hunteri Desmarest, Encyclop. Methodique, Mammalogie, 2: 520, 

1822. Thames River, England. 
Delphinus hyper oodon Desmarest, Encyclop. Methodique, Mammalogie, 2: 

521, 1822. Near Honfleur, France. 
Ceto-diodon hunteri Jacob, Dublin Philosoph. Journ. and Sci. Rev., 1: 72, pi. [3], 

fig. 8, Mar. 1825. Killiney, near Dublin, Ireland. 
Heterodon dalei Lesson, Manuel de Mammalogie, p. 419, 1827. Harwich, Eng- 
land (origin of Dale's specimen); Havre, France, also mentioned. 
Hyperoodon bidens Fleming, History of British Animals, p. 36, 1828. Near 

Maldon, England. 
Hyperoodon honfloriensis Lesson, Hist. Nat. Gen. et Partic. des Mamm. et 

Oiseaux decouverts depuis 1788 (Compl. Oeuvres de Buffon, 1), Cetac^s, 

p. 137, 440 (name), 1828. Honfleur, France. 
Hyperoodon latijrons Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 27, 

pi. 4, 1846. Orkney Ids. 
Hyperoodon borealis Nilsson, Skandinav. Fauna, pt. 1, mamm., p. 622, 1847. 

Coasts of Iceland and Faroe Islands, Greenland and Spitsbergen. 

Mesoplodon Gervais. Cowfish 

Aodon Lesson, Hist. Nat. Gen. et Partic. des Mamm. et Oiseaux decouverts 
depuis 1788, (Compl. Oeuvres de Buffon, 1), Cetaces, p. 150, 440, 1828. 
Genotype Aodon dalei J^esson = Mesoplodon bidens (Sowerby). (Name said 
to be preoccupied by Anodon Lacepede, 1798, in Pisces.) 

Micropterus Wagner, Schreber's Saugthiere, 7: 281, 352, pi. 348, 1846. Sub- 
genus of Delphinus; type Delphinus micropterus Cuvier = Mesoplodon 
bidens (Sowerby). Preoccupied by Micropterus Lacepede, 1802, in Fishes. 

Mesoplodon Gervais, Ann. des Sci. Nat., zool., (3) 14: 16, July 1850. Genotype 
Mesoplodon bidens (Sowerby) . 

Dioplodon Gervais, Compt. Rend. Acad. Sci. Paris, 31: 512, 1850. Genotype 
Delphinus densirostris Blainville. 

Mesodiodon Duvernoy, Ann. des Sci. Nat., zool., (3) 15: 41, pi. 2, f. 2, 2', 1851. 
Genotype Dioplodon sowerbyi Gervais = Mesoplodon bidens (Sowerby). 

Dolichodon Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 353, text-f. 72, 
1866. Subgenus of Ziphius, type Ziphius layardii Gray. 

Callidon Gray, Ann. Mag. Nat. Hisb (4) 7: 368, text-figs.. May 1871. Geno- 
type Mesoplodon gilnlheri Krefft = M. layardii (Gray) . 

Neoziphius Gray, Suppl. to Cat. Seals and Whales Brit. Mus., p. 101, 1871. 
Genotype Dioplodon europseus Gervais, = Mesoplodon europseus (Gervais). 

Mesoplodon densirostris (Blainville) 

Delphinus densirostris Blainville, in Desmarest, Nouv. Dictionnaire d'Hist. 
Nat., ed. 2, 9: 178, 1817. Locality unknown. 



262 bulletin: museum of comparative zoology 

Ziphius sechellensis Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 28, 
1846. Seychelles, Indian Ocean. 

Mesoplodon layardii (Gray). Strap-toothed Whale 

Ziphius layardii Gray, Proc. Zool. Soc. London, p. 358, text-f., 1865. Cape of 

Good Hope. 
Mesoplodon giintheri Krefft, Ann. Mag. Nat. Hist. (4) 7: 368, 2 figs.. May 

1871. Little Bay, near Sydney, New South Wales. 

Ziphius G. Cuvier 

Epiodon Rafinesque, Precis des Decouvertes et Trav. Somiologiques entre 1800 

et 1814, p. 13, 1814. Genotype Epiodon urganantus Rafinesque = IZiphius 

cavirostris G. Cuvier. 
Ziphius G. Cuvier, Recherches sur les Ossemens Foss., ed. 2, 5: pt. 1, p. 352, 

pi. 27, f. 3, 1823. Genotype Ziphius cavirostris G. Cuvier. 
Xiphius L. Agassiz, Nomenclator Zool., Index Univers., p. 389, 392, 1846; 

emendation. 
Aliama Gray, Proc. Zool. Soc. London, p. 242, Nov. 1864. Genotype Delphinus 

desrnarestii Ilisso= Ziphius cavirostris G. Cuvier. 
Petrorhynchus Gray, Proc. Zool. Soc. London, p. 524, 2 text-f., Oct. 1865. 

Genotype Hyperoodon capensis Gray = Ziphius cavirostris G. Cuvier. 
Ziphiorrhynchus Burmeister, Revista Farmaceutica, Oct. 1865; Ann. Mag. 

Nat. Hist. (3) 17: 94, Feb.; 303, Apl. 1866. Genotype Ziphiorrhynchus 

cryptodon Burmeister = Zip/iz us cavirostris G. Cuvier. 

Ziphius cavirostris G. Cuvier. Cuvier's Beaked Whale 

Epiodon urganantus Rafinesque, Precis des Decouvertes et Trav. Somiologiques 

entre 1800 et 1814, p. 13, 1814. Mediterranean Sea. Nomen nudum. 
Ziphius cavirostris G. Cuvier, Rech. sur Ossemens Foss., ed. 2, 5: pt. 1, p. 352, 

pi. 27, f. 3, 1823. Between Fos and the mouth of the Galegeon, dept. des 

Bouches-du-Rhone, France. 
Delphinus desrnarestii Risso, Hist. Nat. Anim. Europe Merid., 3: 24, pi. 3, f. 3, 

1826. 
Delphinus philippii Cocco, Arch. f. Naturgesch., 12: sect. 1, p. 104, pi. 4, f. C, 

1846. Straits of Messina. 
The article is quoted from Maurolico, Journ. Lit. Kabinets v. Messina (2) 1 : 
pt. 6, Oct. 1841. 
Hyperoodon doumetii Gray, Cat. Spec. Mamm. Brit. Mus., Cetacea, p. 68, 1850. 

Corsica. 
Hyperoodon gervaisii Duvernoy, Ann. des Sci. Nat., zool., (3) 15: 49, 67, 1851. 

Coast of Aresquiers, near Frontignan, departement of I'Herault, France. 
Ziphius indicus Van Beneden, Mem. Couronnes et Autrcs Mem. Acad. Roy. d. 

Sci., Lettres et Beaux- Arts Belgique, Bruxelles, 16: art. 1, p. 23, pi. 1, June 

1864. Cape of Good Hope, probably on Indian Ocean side. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 263 

Delphinorhynchus australis Burmeister, Allgemein. Zeitschr. f. Ges. Naturw., 

Dresden, 26: 262, Aug. 1865; An. Mus. Buenos Aires, 1: 312, pi. 15-20, 

1865. Argentina, near Buenos Aires. 
Hyperodon semijundus Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 280, Dec. 

1865. Charleston, South Carolina. 
Hyperoodon capensis Gray, Proc. Zool. Soc. London, p. 359, text-f., Oct. 1865. 

Cape of Good Hope. 
Ziphiorrhynchus cryptodon Burmeister, Ann. Mag. Nat. Hist. (3) 17: 94, pi. 3, 

Feb.; 303, Apl. 1866. Buenos Aires, Argentina. 
Petrorhynchus mediterraneus Gray, Suppl. to Cat. Seals and Whales Brit. Mus., 

p. 98, 1871. Mediterranean Sea. 
Epiodon chathamiensis Hector, Trans, and Proc. New Zealand Inst., 5: 164, 

text-f. a, b, pis. 4, 5, May 1873. Chatham Islands. 
Delphinus savii Richiardi, Arch, per Zool., pi. 3, 7, 8, 1873. 
Ziphius (Epiodon) rwvse-zealandise Haast, Trans, and Proc. New Zealand Inst., 

12: 241, pi. 8, May 1880. Lyttelton Harbor, New Zealand. 
Ziphius grebnitzkii Stejneger, Proc. U. S. Nat. Mus., 6: 77, 22 June 1883. 

Commander Ids., Bering Sea. 



Family BAL.ENID.E. Right Whales 
EuBAL.ENA Gray. Right Whales 

Balsena Linnaeus, Syst. Nat., ed. 10, 1: 75, 1758, in part. Genotype Balsena 

mysticetus Linnaeus. 
Leiobalsena Eschricht, K. Dansk. Vid. Selsk. Skrifter, Kjobenhavn, nat.-math. 

Afd., (5) 1: 108, 1849. Genotype the "Glathvaler." 
Eubalaena Gray, Proc. Zool. Soc. London, p. 201, 1864. Genotype Euhalxna 

australis (Desmoulins). 
Hunterus Gray, Ann. Mag. Nat. Hist. (3) 14: 349, Nov. 1864. Genotype Hunt- 

erus temminckii Gray = Eubalsena glacialis Bonnaterre. Spelling emended 

to Hunterius in Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 78, 1866. 
Macleayius Gray, Proc. Zool. Soc. London, for 1864, p. 589, text-f. 1, 2, May 

1865. Genotype Macleayius australiensis Gray. 

EuBAL^NA AUSTRALIS (Desmoulins). Southern Right Whale. 

Balsena australis Desmoulins, Dictionnaire Class. d'Hist. Nat., 2: 161, 1822. 

Cape of Good Hope, Algoa Bay. Doubtfully distinct from glacialis; see 

Lonnberg, E. 
Balsena mysticetus antarctica Schlegel, Abh. aus dem Gebiete d. Zool., pt. 1, 

p. 37, 1841 (in part). Cape of Good Hope. 
Balsena sieboldii Gray, Ann. Mag. Nat. Hist. (3) 14: 349, Nov. 1864. North 

Pacific. 



264 bulletin: museum of comparative zoology 

Hunterus iemminckii Gray, Ann. Mag. Nat. Hist., (3) 14: 349, Nov. 1864; 
Hunterius temminckii Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 98, 
1866. Cape of Good Hope. 

EuBAL^NA GLACiALis (Borowski). Atlantic Right Whale 

Balxna gladalis Borowski, Gemeinniizzige Naturgesch. d. Thierreichs, (Berlin), 

2: pt. 1, p. 18, 1781. North Cape, Norway, also southern Atlantic Ocean, 

off coast of Africa, and near Antilles. 
B[alsena\ gladalis Bonnaterre, Tabl. Encycl. et Method, des Trois Regnes de la 

Nature, Cetologie, p. 3, 1789. Iceland. 
Balsena nordcaper Lacepede, Hist. Nat. des Cetacees, p. 103, pis. 2, 3, (4to ed.), 

1804. North Atlantic, between Spitsbergen, Norway and Iceland. 
Balsena biscayensis Eschricht, Revue et Mag. de Zool. (2) 12: 229, May 1860. 

San Sebastian, Bay of Biscay, Spain. 
Balsena dsarctica Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 168, Aug. 1865. 

Opposite Philadelphia, Pennsylvania. 
Hunterius svedenborgii Lilljeborg, Nova Acta Reg. Soc. Sci. Upsala, (3) 6: pt. 2, 

no. 1, p. 35, pi. 9-11, 1867. W&nga, West Gothland, Sweden (subfossil). 
Balsena britannica Gray, Ann. Mag. Nat. Hist. (4) 6: 198, Sep. 1870. Off Lyme 

Regis, England. 
Balsena eubalsena Gray, Suppl. Cat. Seals and Whales Brit. Mus., p. 44, 1871 

(not of Flower, 1864). 
Balsena tarentina Capellini, Mem. Accad. Sci. dell'Ist. Bologna, (3) 8: 9, pi. 1, 

1877. Gulf of Taranto, Italy. 



Family BAL^ENOPTERID^. Finner Whales 
Bal^noptera Lacepede. Finback Whales 

Balsenoptera Lacepede, Hist. Nat. des Cetacees, 4to ed., p. xxxvi, 114, 1804. 
Genotype not named, but fixed by Flower (1864) as Balsena rostrata 
MilWeT = Balsenoptera acuto-rostrata Lacepede. 

Rorqual G. Cuvier, Regne Animal, ed. 2, 1: 298, 1829. Genotype not speci- 
fied, but included Balsena boops Linnaeus and B. musculus Linnaeus. Form 
here used probably not intended as a generic term, but altered to Ror- 
qualus F. Cuvier, Hist. Nat. des Cctaces, p. 303, pi. 20, 1836. 

Pterobalsena Eschricht, Kongl. Danske Vid. Selsk. Skr., Kjobenhavn, nat.- 
math. Afd., (5) 1 : 108, 1849. Genotype the Finhval, = Balsenoptera physalus 
Linnaeus. 

Benedenia Gray, Proc. Zool. Soc. London, p. 211, Nov. 1864. Genotype 
Benedenia knoxii Gr&y = Balsenoptera physalus (Linnaeus). 

Rudolphius Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 170, 1866. 
As a subgenus of Sibbaldius, type Sibbaldius laticeps Gray = Balsenoptera 
borealis Lesson. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 265 

Bal^noptera acuto-rostrata Lac6pede. Little Piked Whale; Least Ror- 
qual; Bay Whale 

Balsena rostrata O. Fabricius, Fauna Grcelandica, p. 40, 1780 (not of P. L. S. 
Miiller, 1776). Greenland Seas. 

Balxnoptera acuto-rostrata Lacepede, Hist. Nat. des Cetacees, 4to ed., pp. 
xxxvii, 134, pi. 8, 1804. Cherbourg, France. 

Balsena minimus borealis Knox, Cat. Prep. Whale, p. 14, 1828. 

Balsena minhna Rapp, Die Cetaceen zool.-anat. dargestellt, p. 52, 1 837. 

Balsenoptera eschrichtii Rasch, Nyt Mag. f. Naturvid., Christiania, 4: 123, 

1845. Swedish coasts. 

Pterobalsena minor grcenlandica and bergensis Eschricht, Kong. Dansk. Vid. 

Selsk. Skrift., Kjobenhavn, (5) 1: 109, 1849. Greenland and Norway, 

respectively. 
Balsenoptera microcephala Gray (ex Brandt, MS.), Cat. Seals and Whales 

Brit. Mus., ed. 2, p. 188, 1866, in syn. of B. rostrata. 
Sibbaldius mondinii Capellini, Mem. Accad. Sci. dell' 1st. Bologna, (3) 7: 

423, 1877. Adriatic Sea. 
Balsenoptera edeni Anderson, Anat. and Zool. Researches Yunnan Exped., p. 

551, pi. 44, 1879. Gulf of Martaban, between Sittang and Beeling Rivers, 

India. 

Bal^noptera BOREALIS Lesson. Lesser Rorqual; Sei Whale 

Balxna rostrata Rudolphi, Abh. K. Wiss. Akad. Berlin for 1820-21, p. 27-40, 

pi. 1-5, 1822 (not of P. L. S. Miiller, 1776; not of O. Fabricius, 1780). 

North Sea. 
Balsenoptera borealis Lesson, Hist. Nat. G6n. et Partic. des Mamm. et Oiseaux 

decouverts depuis 1788 (Compl. Oeuvres de Buff on, 1), Cetaces, p. 342, 

1828. North Sea and Isle of Oleron, France.. 
Balsenoptera laticeps Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 20, 

1846. North Sea, coast of Holstein (ex Rudolphi). 

Balsenoptera brydei Olsen, Tidens Tegn, 12 Nov. 1912 (Norwegian newspaper); 
Proc. Zool. Soc. London, p. 1074, pi. 109, 110, 17 Dec. 1913. Saldahna Bay, 
Cape Colony. 

BaljENOPTera physalus (Linnseus). Common Finback Whale 

Balsena physalus Linnseus, Syst. Nat., ed. 10, 1: 75, 1758. European seas. 

Balsena boops Linnseus, Syst. Nat., ^d. 10, 1: 76, 1758 (young of B. physalus). 

"In Oceano septentrionali." 
Balsenoptera gibbar Lacepede, Hist. Nat. des Cetacees, (4to ed.) p. xxxvi, 114, 

1804. Arctic Ocean, near Greenland. 
Balsenoptera rorqual Lac^p^de, Hist. Nat. des Cetacees, (4to ed.) p. xxxvii, 

126, 1804. Eastern North Atlantic, Scotland to Mediterranean. 
Balsena sulcata Neill, Mem. Wernerian Nat. Hist. Soc, Edinburgh, 1 : 212, 1811. 

Near Alloa, Scotland. 



266 bulletin: museum of comparative zoology 

Balsenoptera mediterraneensis Lesson, Hist. Nat. Gen. et Partic. des Mamm. et 
Oiseaux decouverts depuis 1788 (Compl. Oeuvres de Buffon, 1), Cetaces, 
pp. 361, 442 (name), 1828. Renaming of B. rorqual Lacepede. Mediter- 
ranean Sea. 

Physalis vulgaris Fleming, History of Brit. Animals, p. 32, 1828. No locality. 

Balsena antiquorum J. B. Fischer, Synopsis Mamm., p. 525, 1829. Mediter- 
ranean Sea. 

Balsenoptera aragous Farines and Carcassonne, Mem. sur un Cetace 6chou6 27 
nov. 1928 sur la Cote de Saint-Cyprien, 2 pp., 1829 (Perpignan). Coast of 
St. Cyprien, Corsica. 

Balsenoptera tenuirostris Sweeting, Charlesworth's Mag. Nat. Hist. (2) 4: 343, 
July 1840. Charmouth Beach, England. 

Balsena sulcata ardica Schlegel, Abhandl. aus d. Gebiete d. Zool., pt. 1, p. 38, 
pi. 6, f. 1-5, 1841. Coast of Holland. 

Physalus duguidii Heddle, Proc. Zool. Soc. London, p. 187, pi. 44, 45 (Mam- 
mals), 11 Dec. 1856. Island of Laman (or Lambholm), Orkneys. The 
name occurs on the plates only. 

Balsenoptera robusta Lilljeborg, Forhandl. ved Skandinaviske Naturforskeres 
Ottende M0de, Kjobenhavn, for 1860, 8: 602, 1861. Sweden. 

Benedenia knoxii Gray, Proc. Zool. Soc. London, p. 212, text-f. 8-8b, Nov. 1864. 
Coast of North Wales. 

S[ibbaldius] tuberosus Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 16, 20 July 
1869. Coast of eastern North America. 

Sibbaldius tectirostris Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 17, 20 July 
1869. Sinepuxent Bay, Maryland. 

Balsenoptera blythii Anderson, Anat. and Zool. Researches Yunnan Exped., 
p. 564, 1879. Indian Coast. 

Dubertus rhodinsulensis Trumbull, in G. B. Goode, Fisheries and Fishery In- 
dustries of U.'S., sect. 1,1: 29, 1884 (nomen nudum). Rhode Island, U. S. 
Amer. 

Megaptera Gray. Humpback Whale 

Kyphobalsena Eschricht, Kongl. Danske Vid. Selsk. Skr., Kjobenhavn, nat.- 

math. Afd., (5) 1: 108, 1849. Genotype Kyphobalsena boops (O. Fabricius) 

= Megaptera nodosa (Bonnaterre). 
Megaptera Gray, Ann. Mag. Nat. Hist. (3) 14: 350, Nov. 1864. Genotype 

Balsena longimana Rudolphi = Batena nodosa Bonnaterre. 
PoBscopia Gray, Proc. Zool. Soc. London, p. 207, text-f. 3, Nov. 1864. Subgenus 

of Megaptera, type Balsena lalandii J. B. Fischer. 

Megaptera nov^anglle (Borowski). Humpback Whale 
Balsena boops O. Fabricius, Fauna Groenlandica, p. 36, 1780, and later authors, 
but not of Linnaeus. Greenland seas, especially Pamiut and Pissukbik. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 267 

Balsena novae anglise Borowski, Gemeinniizzige Naturgesch. des Thierreichs 

(Berlin), 2: pt. 1, p. 21, p. 1781. New England coast. 
For use of this name see Kellogg, R., Proc. Biol. Soc. Washington, 45: 148, 
9 Sep. 1932. 
B[al3ena] nodosa Bonnaterre, Tabl. Encycl. et Method, d. Trois Regnes de la 

Nature, Cetologie, p. 5, 1789. New England coast. 
Balaenoptere poeskop Desmoulins, Dictionnaire Class. d'Hist. Nat., 2: 164, 1822. 

Cape of Good Hope. Not Latinized. 
Balaenoptera australis Lesson, Hist. Nat. Gen. et Partic. des Mamm. et Oiseaux 

decouverts depuis 1788 (Compl. Oeuvres de Buff on, 1), Cetaces, p. 372, 

1828. Cape of Good Hope. 
Balaena lalandii J. B. Fischer, Synopsis Mamm., p. 525, 1829. Cape of Good 

Hope. 
Balsena longimana Rudolphi, Abh. K. Akad. Wiss. Berlin, p. 133, pi. 1-5, 1832. 
Balsenoptera capensis A. Smith, South African Quart. Journ., 2: 130, 1834. 

Cape of Good Hope. 
Balaenoptera leucopteron Lesson, Nouv. Tabl. Regne Anim., Mamm., p. 202, 

1842, nomen nudum. High southern latitudes. 
Megaptera americana Gray, Zool. Voy. Erebus and Terror, 1: Mamm., p. 17, 

1846. Bermuda Islands. 
Balaenoptera syncondylus A. Midler, Schrift. K. Phys.-oekonom. Ges. Konigs- 

berg, 4: 38, pi. 1-3, 1863. "Ostsee an die kurische Nehrung," Germany. 
Megaptera gigas Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 179, Dec. 1865. 
Megaptera osphyia Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 180, Dec. 1865. 

Forty miles off Petit Manan Lighthouse, Maine. 
Megaptera expansa Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 193, Dec. 1868. 

Nomini Cliffs, Westmoreland County, Virginia (?Miocene). 
Megaptera longimana var. moorei Gray, Cat. Seals and Whales Brit. Mus., ed. 

2, p. 122, 1866. Estuary of the Dee, Scotland. 
Kyphobalaena keporkak Van Beneden, Bull. Acad. Roy. Sci., Lettres et Beaux- 
Arts Belgique, Bruxelles, (2) 25: 116, 1868. No locality. 
Megaptera hellicosa Cope, Proc. Amer. Phil. Soc, 12: 107, text-f. 21-27, 1871. 

St. Bartholomew's, West Indies. 
Megaptera novse-angliae Trouessart, Cat. Mamm. Viv. Foss., pt. 5, p. 1085, 1898, 

based on Brisson. New England coast. 



SiBBALDUs Gray 

Sibbaldus Gray, Proc. Zool. Soc. London, p. 222, text-f. 16-18, Nov. 1864 
Genotype Sibbaldus borealis Gray (not hesson) = Balaena musculus Lin- 
nseus. Spelling changed to Sibbaldius by Flower, Proc. Zool. Soc. London, 
p. 391, 1864. 

Cuvierius Gray, Cat. Seals and Whales Brit. Mus., ed. 2, p. 114, 1866. Geno- 
type Physalus latirostris Flower = Sibbaldus musculus (Linnaeus). 



268 bulletin: museum of comparative zoology 

Flowerius Lilljeborg, Nova Acta Reg. Soc. Sci., Upsala, (3) 6: art. 6, p. 11^ 
1867. Genotype "Flowerius gigas (Eschricht), Sibbaldius borealis Gray" = 
Balsena musculus Linnseus. 

Balsenoptera in part, of authors. 

SiBBALDUS MUSCULUS (Linnseus). Blue Whale 

Balsena musculus Linnaeus, Syst. Nat., ed. 10, 1 : 76, 1758. "In Mare Scotico" = 

Frith of Forth, Scotland. 
Balsenoptera jubartes Lacepede, Hist. Nat. des Cetacees (4to ed.), pp. xxxvii, 

120, 1804. Greenland Seas to Iceland. 
Balsena maximus borealis Knox, Cat. Prep. Whale, p. 5, 1828. ' 

Physalus (Rorqualus) sibbaldii Gray, Proc. Zool. Soc. London, p. 92, 20 July 

1847. Coast of Yorkshire, England. 
Balaenoptera gigas Reinhardt, in Rink's Gronland Geografisk og Statistisk 

Beskrevet, Bidrag, 1 : pt. 2, p. 10, 1857. Greenland Seas. 
Balaenoptera indica Blyth, Journ. Asiatic Soc. Bengal, (2) 29: 488, 1859. Sondip, 

Bay of Bengal. 
Balaenoptera patachonica Burmeister, Proc. Zool. Soc. London, p. 191, text-f. 

1-11, June 1865. Banks of La Plata River, "some leagues from Buenos 

Ayres," Argentina. 
Balaenoptera carolinae Malm, Nagra blad om Hvaldjur i Allmaenhet og Balaenop- 
tera carolinae i Synnerhet, Goeteborg, 1866. 
Sibbaldius sulfureus Cope, Proc. Acad. Nat. Sci. Philadelphia, p. 20, 20 July 

1869. Northwest coast of United States. 
Pterobalaena gryphus Munter, Mitth. Naturw. Verein von Neu-Vorpommern u. 

Rligen, 9: 1, pi. 1-2, 1877. Wieck bei Greifswald, Germany. 



Order NOMARTHRA. Pangolins 

Although this term originally included both the pangolins and the aard- 
varks, the latter have now been removed to a separate order, so that either the 
name must be used in the restricted sense, or a new one proposed. 
Lepidota Lane, Science, new ser., 31: 913, 1910; to replace Pholidota Weber, 
1904, not of Merrem, 1820. Not Squamata Huxley, 1872, which is ante- 
dated by the same term proposed by Oppel for reptiles. Not Lepidota 
Vogt, 1851; see Gill, Science, new ser., 32: 56, 8 July 1910. 

Family MANIDiE. Pangolins 

Review: Pocock, R. I. The external characters of the pangolins (Manidae). 
Proc. Zool. Soc. London, p. 707-723, text-f. 6-13, 22 Sep. 1924. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 269 

Subfamily SMUTSIIN.E 
Smutsia Gray 

^Smutsia Gray, Proc. Zool. Soc. London, p. 369, Oct. 1865. Genotype Manis 

temminckii Smuts. 

Smutsia gigantea (Illiger). Giant Ground Pangolin 

Manis gigantea Illiger, Abh. K. Akad. Wiss., Berlin, p. 84, 1815. 

Pholidotus africanus Gray, Proc. Zool. Soc. London, p. 368, pi. 17, text-f., Oct. 

1865. River Niger. 
Manis wagneri Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 65: sect. 

1, p. 48, 1872. 

Smutsia temminckii (Smuts) 

Manis temminckii Smuts, Enumerat. Mamm. Capensium, p. 54, pi. 3, f. 1-2, 

1832. Cape of Good Hope, from beyond Latukou. 
Phatages hedenborgii Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. CI., 

65: sect. 1, p. 69, 1872. Sennaar, and elsewhere in northeastern and 

central Africa. 

Subfamily UR0MANINJ5. Arboreal Pangolins 
Phataginus Rafinesque 

Phataginus Rafinesque, Ann. Gen. Sci. Phys. de Bruxelles, 7: 214, 1820. As a 
subgenus, type Manis tricuspis Rafinesque. 

Phatagin Gray, Proc. Zool. Soc. London, p. 363, 2 text-f., Oct. 1865. As a sub- 
genus of Manis; type Manis tricuspis Rafinesque. 

Triglochinopholis Fitzinger, Sitzb. K. Akad. Wiss., Wien, math.-nat. CI., 65: 
sect. 1, p. 27, 1872. Genotype Manis tricuspis Rafinesque. 

Phataginus tricuspis (Rafinesque) 

Manis tricuspis Rafinesque, Ann. Gen. Sci. Phys. de Bruxelles, 7: 214, 1820 

West Africa. 
Manis multiscutata Gray, Proc. Zool. Soc. London, p. 22, July 1843. "Western 

Africa." 
M[anis] tridentata Focillon, Revue et Mag. de. Zool., (2) 2: 472, pi. 11, Sep. 

1850. Coast of Mozambique (?). 

Uromanis Pocock. Long-tailed Pangolin 

Uromanis Pocock, Proc. Zool. Soc. London, pp. 722, 723, 22 Sep. 1924. Geno- 
type Manis longicaudata Brisson. 

Uromanis longicaudata (Brisson) 

Pholidotus longicaudatus Brisson, Regne Anim., 3: Quadr., p. 19, 1762. 



270 bulletin: museum of comparative zoology 

Manis tetradactyla Linnseus, Syst. Nat., ed. 12, 1 : 53, 1766. "America australi." 

= West Africa. 
Manis macroura Erxleben, Syst. Regni Animalis, p. 101, 1777. "Guinea?" 
Manis ceonyx Rafinesque, Ann. Sci. Phys. Bruxelles, for 1820, 7: 215, 1821. 
Manis africana Desmarest, Encyclop. Methodique, Mammalogie, 2: 376, 1822. 

Senegal and Guinea. 
Manis guineensis and senegalensis Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.- 

nat. CI., 65: sect. 1, p. 16, 17 resp., 1872. Guinea and Senegambia. 
Manis hessi Noack, Zool. Jahrb., Syst., 4: 100, pi. 1; 3, f. 1-3, 28 Feb. 1889. 

Banana, north bank of the mouth of the Congo. 



Order TUBULIDENTATA. Aard-varks 
Family ORYCTEROPODID^ 

Review: Pocock, R. I. Some external characters of Orycteropus afer. Proc. 
Zool. Soc. London, p. 697-706, text-f. 1-5, 22 Sep. 1924. 

Orycteropus E. GeoflProy 

Orycteropus E. Geoffroy, Decade Philosophique, 1795; Bull. Sci. Soc. Philomati- 
que, Paris, (1) 1: 102, Apr.-Je. 1796. Genotype Myrmecophaga capensis 
Gmelin = M. afra Pallas. 

Note: the status of most of the named forms is subject to revision. 

Orycteropus afer afer (Pallas). South African Aard-vark 
Myrmecophaga afra Pallas, Miscellanea Zool., p. 64, 1766. Cape of Good Hope. 
Myrmecophaga capensis Gmelin, Linn. Syst. Nat., ed. 13, 1: pt. 1, p. 53, 1788. 
Cape of Good Hope. 

Orycteropus afer adametzi Grote 

Orycteropus afer adametzi Grote, Arch. f. Naturgesch., 87: sect. A, pt. 7, p. 126, 
f. 2, June 1921. Bamessing, northwestern Cameroons. 

Orycteropus afer ^thiopicus Sundevall 

Orycteropus sethiopicus Sundevall, Kongl. Svenska Vet. -Akad. Handl., Stock- 
holm, for 1842, p. 236, pi. 3, f. 1-5, 1843. Near Bahr-el-Abiad, Sennaar. 

Orycteropus afer albicaudus W. Rothschild 

Orycteropus afer albicaudus W. Rothschild, Novitates Zool., 14: 506, 1 Nov. 
1907. Former German Southwest Africa. 

Orycteropus afer erikssoni Lonnberg 

Orycteropus erikssord Lonnberg, Arkiv f. Zool., 3: no. 3, p. 1, pi. 1, 15 Jan. 
1906. Northern Congo. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 271 

Orycteropus afer faradjius Hatt 

Orycteropus erikssoni faradjius Hatt, Amer. Mus. Novitates, no. 535, p. 1, 6 
June 1932. Faradje, Haut-Uele, Congo Beige. 

Orycteropus afer haussanus Matschie 

Orycteropus haussanus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 102, 
104, 1900. Haussaland, interior of Togo, West Africa. 

Orycteropus afer kordofanicus W. Rothschild 

Orycteropus afer kordofanicus W. Rothschild, Ann. Mag. Nat. Hist. (9) 19: 
512, Apl. 1927. Kordofan. 

Orycteropus afer lademanni Grots 

Orycteropus afer lademanni Grote, Arch. f. Naturgesch., 87: sect. A, pt. 7, 
p. 123, June 1911. Wassi, north of Kondoa-Irangi, Masai Plains, Tan- 
ganyika Territory. 

Orycteropus afer leptodon Hirst 

Orycteropus leptodon Hirst, Ann. Mag. Nat. Hist. (7) 17: 383, Mar. 1906. 
Efulen, Cameroons. 

Orycteropus afer matschiei Grote 

Orycteropus afer matschiei Grote, Arch. f. Naturgesch., 87: sect. A, pt. 7, p. 
122, f. 1, June 1921. Mikindani, coast of Tanganyika Territory. 

Orycteropus afer observandus Grote 

Orycteropus afer observandus Grote, Arch. f. Naturgesch., 87: sect. A, pt: 7, 

p. 123, f. 1, June 1921. Ussangire, northwest of Ssongea, northeast coast 

of Lake Nyasa, Tanganyika Territory. 

Orycteropus afer ruvanensis Grote 

Orycteropus afer ruvanensis Grote, Arch. f. Naturgesch., 87: sect. A, pt. 7, 

p. 123, f. 1, June 1921. Ruwana region, southeast coast of Victoria 

Nyanza, Tanganyika Territory. 

Orycteropus afer senegalensis Lesson 

Orycteropus senegalensis Lesson, Revue Zoologique par Soc. Cuvierienne, 3: 

225, Aug. 1840. Senegal. 
Orycteropus senegalensis Schinz, Syst. Verzeichn. d. Saugeth., 2: 319, 1845. 

Podor River, Senegambia. 
These two are apparently independent namings of the same animal. 

Orycteropus afer somalicus Lydekker 

0[rycteropus] a[fer] somalicus Lydekker, Game Animals of Africa, p. 466, 1908. 
Somaliland. 



272 bulletin: museum of comparative zoology 

Orycteropus afer wardi Lydekker 

Olrycteropus] a[fer] ivardi Lydekker, Game Animals of Africa, p. 467, 1908. 
Northeastern Rhodesia. 

Orycteropus afer wertheri Matschie 

Orycteropus wertheri Matschie, in Werther, C. W., Die mittleren Hochlander 

des nordUchen Deutsch-Ost-Afrika, (Berlin), p. 266, 1898. Inland of 

Bagamoyo, Tanganyika Territory. 



Order RODENTIA. Gnawing Mammals 

St. Leger, Jane. A key to the families and genera of African Rodentia. 
Proc. Zool. Soc. London, p. 957-997, pi. 1-10 (col.), 30 Sep. 1931. 



Suborder DUPLICIDENTATA 
Family LEPORID^E. Hares and Rabbits 

Bunolagus Thomas. Kopje Hares 

Bunolagus Thomas, Proc. Zool. Soc. London, p. 109, 30 Apl. 1929. Genotjrpe 
Lepus monticularis Thomas. 

Bunolagus monticularis (Thomas) 

Lepus monticularis Thomas, Ann. Mag. Nat. Hist. (7) 11: 78, Jan. 1903. 
Deelfontein, central Cape Colony. 

Lepus Linnreus. Hares 

Lepus Linnaeus, Syst. Nat., ed. 10, 1 : 57, 1758. Genotype Lepus timidus 

Linnaeus. 
Chionobates Kaup, Skizzirte Entw.-Gesch. Naturl. Syst. Europ. Thierwelt, 1: 

170, 1829. Genotypes Lepus variabilis and L. borealis. 
Eulagos Gray, Ann. Mag. Nat. Hist. (3) 20: 222, Sep. 1867. No type specified; 

included Eulagos niediterraneus (Wagner) and Eulagos judsex Gray. 
Eulepus Acloque, Faune de France, Mamm., p. 52, 1900. As a subgroup of 

Lepus, to include L. timidus Linnaeus and L. variabilis Pallas, of which 

the former is now designated as type. 

Lepus ^egyptius Desmarest 

Lepus aegyptius Desmarest, Encyclop. Methodique, Mammalogie, suppl., p. 

350, 1822 (in part). Egypt. 
Lepus sethiopicus Hemprich and Ehrenberg, Symbolae Physicae, Mamm., dec. 

2, folio n, page 2, pi. 13, Oct. 1832. Nubia and Dongola, Upper Nile. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 273 

LePUS ANGOLENSI8 ANGOLENSIS ThomaS 

Lepus angolensis Thomas, Ann. Mag. Nat. Hist. (7) 13: 420, text-f., June 1904. 
Ambaca, Angola. 

Leptjs ANGOLENSIS MERiDiONALis Monard 

Lepus angolensis meridionalis Monard, Bull. Soc. Neuchateloise des Sci. Nat., 

Neuchatel, 57: 61, 1932. Kubango region, Angola (Rio Mbale, Vila da 

Ponte, Caquindo). 

Lepus ansorgei ansorgei Thomas and Wroughton 

Lepus ansorgei Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 16: 176, 
Aug. 1905. Caiala, Bihe, Angola, 6000 ft. alt. 

Lepus ansorgei zairensis Hatt 

Lepus ansorgei zairensis Hatt, Revue Zool. et Bot. Africaines, 26: 423, fig., 15 
June 1935. Camp de la Lufu, lower Congo, Congo Beige. 

Lepus atlanticus De Winton 

Lepus atlanticus De Winton, Proc. Zool. Soc. London, for 1897, p. 960, text-f. 
3, 5, Apl. 1898. Ras-el-Ain, in Haha, Morocco. 
Range: Southwestern Morocco. 

Lepus barc^us Ghigi 

Lepus barcseus Ghigi, Mem. R. Accad. Sci. 1st. Bologna, (7) 7: 81, Dec. 1920. 
Woods of Frejna, near Merg, Egypt. 

Lepus berberanus Heuglin 

Lepus berberanus Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 28: 
no. 10, p. 5, 1861. No locality; Berbera, Somaliland, implied. 

Lepus canopus Thomas and Hinton 

Lepus canopus Thomas and Hinton, Novitates Zool., 28: 12, May 1921. 
Farniso, near Kano, northern Nigeria. 

Lepus capensis capensis Linnaeus 

Lepus capensis Linnaeus, Syst. Nat., ed. 10, 1: 58, 1758. Cape of Good Hope. 

Lepus arenarius I. Geoffroy, Dictionnaire Class. d'Hist. Nat., 9: 383, Feb. 1826 

(young). "Dans les sables du pays des Hottentots." 
Lepus capensis var. major Grill, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 

(2) 2: no. 10, p. 19, 1858. Roodeval, Karroo, Cape of Good Hope. 

Lepus capensis abbotti Hollister 

Lepus capensis abbotti Hollister, Proc. Biol. Soc. Washington, 31: 35, 16 May 
1918. Plains east of Mt. Kilimanjaro, Kenya Colony. 



274 bulletin: museum of comparative zoology 

Lepus capensis aquilo Thomas and Wroughton 

Lepus capensis aquilo Thomas and Wroughton, Proc. Zool. Soc. London, p. 
297, 1 Aug. 1907. Coguno, Inhambane, Mozambique. 

Lepus capensis bedfordi Roberts 

Lepus capensis bedfordi Roberts, Ann. Transvaal Mus., 15: 14, 1 Oct. 1932. 
Kalkbank, Pietersburg district, Transvaal. 

Lepus capensis centralis Thomas 

Lepus capensis centralis Thomas, Ann. Mag. Nat. Hist. (7) 12: 344, Sep. 1903. 
Deelfontein, Cape of Good Hope. 

Lepus capensis crawshati De Winton 

Lepus crawshayi De Winton, Proc. Zool. Soc. London, p. 416, pi. 24, 1 Aug. 

1899. Kitwi, east of Athi River and northeast of Machakos, Kenya 

Colony. 

Lepus capensis ermeloensis Roberts 

Lepus capensis ermeloensis Roberts, Ann. Transvaal Mus., 15: 14, 1 Oct. 1932. 
South Africa. 

Lepus capensis granti Thomas and Schwann 

Lepus capensis granti Thomas and Schwann, Abstr. Proc. Zool. Soc. London, 

no. 2, p. 6, 9 Feb. 1904; Proc. Zool. Soc. London, 1 : 182, 9 June 1904. Port 

Nolloth, Little Namaqualand, Southwest Africa. 

Lepus capensis hartsensis Roberts 

Lepus capensis hartsensis Roberts, Ann. Transvaal Mus., 15: 14, 1 Oct. 1932. 
Fourteen Streams, Orange Free State, South Africa. 

Lepus capensis kalaharicus Dollman 

Lepus ochropus kalaharicus Dollman, Ann. Mag. Nat. Hist. (8) 6: 400, Oct. 
1910. Lehutitung, Bechuanaland. 

Lepus capensis langi Roberts 

Lepus capensis langi Roberts, Ann. Transvaal Mus. ,15: 15, 1 Oct. 1932. Nkate, 
northern Bechuanaland. 

Lepus capensis mandatus Thomas 

Lepus capensis mandatus Thomas, Proc. Zool. Soc. London, 1 : 309, 29 Apl. 
1926. Berseba, Great Namaqualand, Southwest Africa. 

Lepus capensis narranus Thomas 

Lepus capensis narranus Thomas, Proc. Zool. Soc. London, 1: 309, 29 Apl. 
1926. Rooibank, Kuiseb River, Namib Desert, Southwest Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 275 

Lepus capensis ochropoides Roberts 

Lepus capensis ochropoides Roberts, Ann. Transvaal Mus., 13: 121, 6 July 
1929. Burghersdorp, Cape Province. 

Lepus capensis ochropus Wagner 

Lepus ochropus Wagner, Schreber's Saugthiere, Suppl., 4: 96, 1844. Cape of 
Good Hope. 
"The yellow-naped High Veldt representative" of L. capensis (Thomas). 

Lepus capensis sal^ Jentink 

Lepus salse Jentink, Notes Leyden Mus., 2: 57, Apl. 1880. Mossamedes, Ben- 
guela, Angola. 
See Thomas, O., Proc. Zool. Soc. London, p. 309, 29 Apl. 1926, as to status 
of this form. 

Lepus capensis vernayi Roberts 

Lepus capensis vernayi Roberts, Ann. Transvaal Mus., 15: 13, 1 Oct. 1932. 
Geluk, Kroonstad, Bechuanaland. 

Lepus chadensis Thomas and Wroughton 

Lepus chadensis Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 384, 
May 1907. Kadde, Lake Chad. 

Lepus cordeauxi Drake-Brockman 

Lepus cordeauxi Drake-Brockman, Ann. Mag. Nat. Hist. (8) 8: 276, Aug. 1911. 
Sheikh Mahomed, eastern Arussi, Abyssinia. 

Lepus crispii Drake-Brockman 

Lepus crispii Drake-Brockman, Ann. Mag. Nat. Hist. (8) 8: 275, Aug. 1911. 
Obbia, Italian Somaliland. 

Lepus fagani Thomas 

Lepus fagani Thomas, Proc. Zool. Soc. London, for 1902, 2: 315, Apl. 1903. 
Zegi, Lake Tana, Ethiopia, 4000 ft. 

Lepus gungunyan^ Roberts 

Lepus gungunyanse Roberts, Ann. Transvaal Mus., 4: 184, 22 Aug. 1914. 
Jabwielu, between Bubye and Nuanetsi Rivers, southeastern Rhodesia. 

Lepus habessinicus Hemprich and Ehrenberg 

Lepus habessinicus Hemprich and Ehrenberg, Symbolae Physicae, Mamm., 

dec. 2, folio p, 2d page, pi. 15, f. 2, Oct. 1832. East coast of Abyssinia, near 

Arkiko. 



276 bulletin: museum of comparative zoology 

Lepus harterti Thomas 

Lepus harterti Thomas, Novitates ZooL, 10: 301, 25 Aug. 1903. Rio de Oro. 

Lepus hawkeri Thomas 

Lepus hawkeri Thomas, Ann. Mag. Nat. Hist. (7) 8: 277, Oct. 1901. Kaka, 
Anglo-Egyptian Sudan. 

Leptjs isabellinus Cretzschmar 

Lepus isabellinus Cretzschmar, in Riippell's Atlas zu d. Reise im NordHche 

Afrika, Saugeth., p. 52, pi. 20, 1826. Deserts southwest of Ambukol, 

Sudan. 

Lepus kabylicus De Winton 

Lepus kabylicus De Winton, Ann. Mag. Nat. Hist. (7) 1: 155, Feb. 1898. 
Algiers, Algeria. 

Lepus pallidior Barrett-Hamilton 

Lepus pallidior Barrett-Hamilton, Ann. Mag. Nat. Hist. (7) 2: 422, Nov. 1898. 
Aures Mts., near Biskra, Algeria. 

Lepus raineyi Heller 

Lepus raineyi Heller, Smithsonian Misc. Coll., 59: no. 16, p. 18, 5 July 1912. 
Longaya Water, 30 miles south of Mt. Marsabit, Kenya Colony. 

Lepus rothschildi De Winton 

Lepus rothschildi De Winton, Novitates Zool., 9: 444, 25 July 1902. Giza 

Province, Egypt. 
Lepus innesi De Winton, Novitates Zool., 9: 445, 25 July 1902. Gattah, Fay- 

otam Province, Egypt. 

Lepus saxatilis saxatilis F. Cuvier 

Lepus saxatilis F. Cuvier, Dictionnaire des Sci. Nat., 26: 309, 1823. Cape of 

Good Hope. 
Lepus rufinucha A. Smith, Zool. Journ., 4 : 440, 1829. "Rocky and mountainous 

situations in South Africa." 
Lepus longicaudatus Gray, Charlesworth's Mag. Nat. Hist. (2) 1 : 586, Sep. 

1837. "Magellan Land," but doubtless South Africa. 
Lepus fumigatus Wagner, Schreber's Saugthiere, Suppl., 4 : 98, pi. 233 G, 1844. 

"Kaffernland." 

Lepus saxatilis albaniensis Roberts 

Lepus saxatilis albaniensis Roberts, Ann. Transvaal Mus., 15: 12, 1 Oct. 1932. 
Kleinpoort, Fish River Valley, Albany district. Cape Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 277 

Lepus saxatilis aurantii Thomas and Hinton 

Lepus saxatilis aurantii Thomas and Hinton, Proc. Zool. Soc. London, p. 497, 
3 Sep. 1923. Louisvale, Orange River, Transvaal. 

Lepus saxatilis chiversi Roberts 

Lepus saxatilis chiversi Roberts, Ann. Transvaal Mus., 13: 119, 6 July 1929. 
Rouxville, Orange Free State. 

Lepus saxatilis chobiensis Roberts 

Lepus saxatilis chobiensis Roberts, Ann. Transvaal Mus., 15: 12, 1 Oct. 1932. 
Kabulabula, Chobe River, Transvaal. 

Lepus saxatilis damarensis Roberts 

Lepus zuluensis damarensis Roberts, Ann. Transvaal Mus., 11: 262, 14 Sep. 

1926. Quickborn, Okahandja, Damaraland, Southwest Africa. 
Lepus saxatilis hechuame Roberts, Ann. Transvaal Mus., 15: 12, 1 Oct. 1932. 

Gaberones, Bechuanaland. 

Lepus saxatilis herero Thomas 

Lepus capensis herero Thomas, Proc. Zool. Soc. London, 1: 308, 29 Apl. 1926. 
Lukualukasi, Ovamboland, Southwest Africa. 

Lepus saxatilis megalotis Thomas and Schwann 

Lepus saxatilis megalotis Thomas and Schwann, Proc. Zool. Soc. London, 1 : 271, 
10 Aug. 1905. Klipfontein, Little Namaqualand, Cape of Good Hope. 

Lepus saxatilis micklemi Chubb 

Lepus zuluensis micklemi Chubb, Ann. Mag. Nat. Hist. (8) 1: 466, June 1908. 
Bulawayo, Southern Rhodesia. 

Lepus saxatilis ngamiensis Roberts 

Lepus saxatilis ngamiensis Roberts, Ann. Transvaal Mus., 15: 12, 1 Oct. 1932. 
Maun, Ngamiland, Bechuanaland. 

Lepus saxatilis nigrescens Roberts 

Lepus saxatilis nigrescens Roberts, Ann. Transvaal Mus., 15: 13, 1 Oct. 1932. 
Vila Pery, inland from Beira, Mozambique. 

Lepus saxatilis subrufus Roberts 

Lepus zuluensis subrufus Roberts, Ann. Transvaal Mus., 4: 100, 20 Oct. 1913. 
Wakkerstroom, Transvaal. 

Lepus saxatilis zuluensis Thomas and Schwann 

Lepus saxatilis zuluensis Thomas and Schwann, Proc. Zool. Soc. London, 1 : 
270, 10 Aug. 1905. Umvolosi, Zululand, Natal. 



278 bulletin: museum of comparative zoology 

Lepus schlumbergeri schlumbergeri Saint-Loup 

Lepus schlumbergeri Saint-Loup, Bull. Soc. Zool. de France, 19: 168, Dec. 1894. 
Yerf-el-Acaab, El Fahs, Morocco (Cabrera). 
Range : Northeastern Morocco from the Straits of Gibraltar to Yebala, and 
perhaps to Um-er-Rbia. 

Lepus schlumbergeri maroccanus Cabrera 

Lepus maroccanus Cabrera, Bol. R. Soc. Espanola Hist. Nat., Madrid, 7: 178, 
Apl. 1907. Marrakesh, Morocco. 
Range: Southern plateau of the interior of Morocco, south of Um-er-Rbia; 
southeast to Seksaua. 

Lepus schlumbergeri pedi^us Cabrera 

Lepus schlumbergeri pediseus Cabrera, Bol. Soc. Espaiiola Hist. Nat., Madrid, 

23: 332, 30 July 1923; Trab. Mus. Nac. de Cienc. Nat., Madrid, zool., no. 

57, p. 304, pi. 10, f. B (col.), 30 Dec. 1932. Ayeddin, eastern Rif, Morocco. 

Lepus schlumbergeri sefranus Thomas 

Lepus sefranus Thomas, Novitates Zool., 20: 590, 21 Oct. 1913. Ain Sefra, 
Morocco, 1100 m. 
Range: Extreme southeastern Morocco and adjacent parts of the Algerian 
Sahara. 

Lepus schlumbergeri sherif Cabrera 

Lepus sherif Cabrera,, Bol. Soc. Espanola Hist. Nat., Madrid, 6: 366, July 1906. 

Mogador, Morocco. 
Lepus schlumbergeri sherif Cabrera, Trab. Mus. Nac. de Cienc. Nat., Madrid, 

zool., no. 57, p. 303, pi. 10, f. 1, 30 Dec. 1932. 
Range: Southwestern Morocco, between the Great Atlas and the sea. 

Lepus senegalensis (Rochebrune) 

Cuniculus senegalensis Rochebrune, Actes Soc. Linn, de Bordeaux, (4) 7: 120, 
1883. Cayor and Oualo, dunes of right bank of the Senegal, Senegambia. 

Lepus somalensis Heuglin 

Lepus somalensis Heuglin, Nova Acta Acad. Ca^s. Leop.-CaroL, Halle, 28: no. 
10, p. 5, 1861. No locality; SomaUland implied. 

Lepus tigrensis Blanford 

Lepus tigrensis Blanford, Ann. Mag. Nat. Hist. (4) 4: 330, Nov. 1869. Tigre, 

Takonda, highlands of Abyssinia (Ethiopia), 2500 m. alt. 
Lepus abyssinicus Lefebvre, Voyage en Abyssinie, zool., 6: atlas, pi. 5, f. 1, 1850. 

Lepus tunet^e De Winton 

Lepus tunetx De Winton, Ann. Mag. Nat. Hist. (7) 1 : 157, Feb. 1898. Tunis. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 279 

Lepus victoria victoria Thomas 

Lepus victorise Thomas, Ann. Mag. Nat. Hist. (6) 12: 268, Oct. 1893. Nassa, 
Speke Gulf, Victoria Nyanza, Tanganyika Territory. 

Lepus victoria kakumeg^ Heller 

Lepus kakumegoe Heller, Smithsonian Misc. CoU., 59: no. 16, p. 19, 5 July 1912. 
Lukosa River, Kakumega Forest, Kenya Colony. 

Lepus victoria microtis Heuglin 

Lepus microtis Heuglin, Leopoldina, 5: 32, June 1865, in Nova Acta Acad. 
Caes. Leop.-Carol., Halle, 24: 1865. "Lande der Ridj," Bahr-el-Ghazal, 
Anglo-Egyptian Sudan. 
As to status of this hare, see Hollister, N., Bull. U. S. Nat. Mus., no. 99, 
pt. 2, p. 169, 1919. 

Lepus whitakeri Thomas 

Lepus whitakeri Thomas, Proc. Zool. Soc. London, 2: 12, pi. 1 (col.), Oct. 1902. 
Wadi Agarib, northwest of Sokna, Tripoli. 

Lepus whytei Thomas 

Lepus whyiei Thomas, Proc. Zool. Soc. London, p. 142, June 1894; for 1896, p. 
795, Apl. 1897. Palombi River, Shirwa plain, Nyasaland. 



Oryctolagus Lilljeborg 

Orydolagus Lilljeborg, Sverig. och Norges Ryggradsdjur, 1: 417, 441, 1873. 

Subgenus of Lepus, type Lepus cuniculus Linnaeus. 
Cuniculus Meyer, Mag. f. Thiergesch., 1: 52, 1790. No type specified; not of 

Brisson, 1762; not of Wagler, 1830. 

Oryctolagus cuniculus cuniculus (Linnaeus) 

Lepus cuniculus Linnaeus, Syst. Nat., ed. 10, 1: 58, 1758. "Europa australi." 
Introduced under domestication, and sometimes feral as in lower Egypt. 

Oryctolagus cuniculus algirus (Loche) 

Cuniculus algirus Loche, Cat. Manam. et Oiseaux observes en Algerie, p. 27, 
1858; Exploration Sci. de 1' Algerie, Zool., Mammiferes, p. 122, 1867. 
Algeria. 
The name is credited to Gervais, ex LerebouUet, but the former gives no 
Latin name. 

Range: Morocco, east to northern Algeria, in the western part of department 
of Constantine. See Cabrera, A., Sobre los conejos de Marruecos. Bol. 
Soc. Espanola Hist. Nat., Madrid, 23: 356-367, 29 Oct. 1923. 



280 bulletin: museum of comparative zoology 

Oryctolagus cuniculus habetensis Cabrera 

0[ryctolagus] cunicidus habetensis Cabrera, Bol. Soc. Espanola Hist. Nat., 
Madrid, 23: 366, 29 Oct. 1923. Dar Amezuk, Morocco. 
Range: Straits of Gibraltar to Lau, Yebala. 

Oryctolagus ctjniculus oreas Cabrera 

Oryctolagus cuniculus oreas Cabrera, Bol. Soc. Espanola Hist. Nat., Madrid , 
22: 112, 27 Feb. 1922. Xauen, Morocco. 
Range: Southeastern part of Yebala, Morocco, east to the high Rif. 



PoELAGUS St. Leger 

Poelagus St. Leger, Proc. Zool. Soc. London, p. 119, text-f. lb, 2b, 3b, 30 Mar. 
1932. Genotype Lepus marjorita St. Leger. 

Poelagus marjorita marjorita (St. Leger). Grass Hare 
Lepus marjorita St. Leger, Ann. Mag. Nat. Hist. (10) 4: 292, Sep. 1929. Near 
Masindi, Bunyoro, Uganda, 4000 ft. 

Poelagus marjorita larkeni St. Leger 

Poelagus marjorita larkeni St. Leger, Ann. Mag. Nat. Hist. (10) 16: 598, Dec. 

1935. Yambio district, 100 miles north of Diawo, southwestern Bahr-el- 

Ghazal, Anglo- Egyptian Sudan. 



Pronolagus Lyon. Rock Hares 

Pronolagus Lyon, Smithsonian Misc. Coll., 45: 416, 15 June 1904. Genotype 
Pronolagus crassicaudatus Lyon (not Geoff roy) = P. ruddi Thomas and 
Schwann (see Lyon, M. W., jr., Proc. Biol. Soc. Washington, 19: 95, 4 
June 1906). 

Pronolagus caucinus Thomas 

Pronolagus caucinus Thomas, Proc. Zool. Soc. London, p. 109, 30 Apl. 1929. 
Karibib, Kaokoveld, Southwest Africa. 

Pronolagus crassicaudatus crassicaudatus (L Geoff roy). Rock Hare 
Lepus crassicaudatus I .Geoffroy, Guerin's Mag. de Zool., 2: cl. 1, pi. 9 and 

text, 1832. Port Natal, Natal. 
Lepus rupestris A. Smith, South African Quart. Journ., 2: 174, 1834. South 

Africa. 
Lepus melanurus Riippell, Mus. Senckenberg., Frankfurt a. M., 3: pt. 2, 

p. 137, 1842. Cape Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 281 

Pronolagus crassicaudatus australis Roberts 

Pronolagus crassicaudatus australis Roberts, Ann. Transvaal Mus., 15: 270, 1 
Aug. 1933. Clan William, Cape of Good Hope. 

Pronolagus crassicaudatus bowkeri Hewitt 

Pronolagus crassicaudatus bowkeri Hewitt, Records Albany Mus., 3: 437, pi. 

26, f. 1; pi. 27, f. 7, 31 July 1927. Gaika's Kop, Amatola Mts., Cape 

Colony. 

Pronolagus crassicaudatus curryi (Thomas) 

Oryctolagus crassicaudatus curryi Thomas, Ann. Mag. Nat. Hist. (7) 10: 245, 
Sep. 1902. Boshof, Orange Free State. 

Pronolagus crassicaudatus nyik^e (Thomas) 

Oryctolagus crassicaudatus nyikse Thomas, Ann. Mag. Nat. Hist. (7) 10: 244, 
Sep. 1902. Nyika plateau, northern Nyasaland, highest peaks, at 7000 ft. 

Pronolagus crassicaudatus saundersi^ Hewitt 

Pronolagus crassicaudatus saundersise Hewitt, Records Albany Mus., 3: 434, 
pi. 26, f. 2, 31 July 1927. Albany district. Cape Colony. 

Pronolagus crassicaudatus vallicola Kershaw 

Pronolagus crassicaudatus vallicola Kershaw, Ann. Mag. Nat. Hist. (9) 13: 
80, Jan. 1924. Rift Valley, Kenya Colony. 

Pronolagus randensis randensis Jameson 

Pronolagus ruddi randensis Jameson, Ann. Mag. Nat. Hist. (7) 20: 404, Novj 
1907. Observatory, Johannesburg, Transvaal, 5900 ft. 

Pronolagus randensis capricornis Roberts 

Pronolagus randensis capricornis Roberts, Ann. Transvaal Mus., 11: 262, 14 
Sep. 1926. Newgate, on the Zoutpansberg, Transvaal. 

Pronolagus randensis makapani Roberts 

P[ronolagus] randensis makapani Roberts, Ann. Transvaal Mus., 10: 75, 31 
Jan. 1924. Makapan's Caves, near Piet Potgietersrust, Transvaal. 

Pronolagus randensis powelli Roberts 

Pronolagus randensis powelli Roberts, Ann. Transvaal Mus., 10: 75, 31 Jan. 
1924. Rooikrans, Rustenburg district, Transvaal. 

Pronolagus ruddi ruddi Thomas and Schwann 

Pronolagus ruddi Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 

18, p. 23, 25 Apl. 1905; Proc. Zool. Soc. London, 1: 272, pi. 16, f. 4, 5, 10 

Aug. 1905. Sibudeni, Zululand, Natal. 



282 bulletin: museum of comparative zoology 

Pronolagus ruddi karieg^ Hewitt 

Pronolagus ruddi kariegx Hewitt, Records Albany Mus., 3: 433, pi. 27, f. 5-6, 
31 Jiily 1927. Bathurst district, Kariega River, Cape Colony. 

Pronolagus ruddi lebombo Roberts 

Pronolagus ruddi lebombo Roberts, Ann. Transvaal Mus., 18: 240, 21 Feb. 

1936. Ubombo, Mozambique. 
Pronolagus ruddi lebomboensis Roberts, Ann. Transvaal Mus., 18: 173, 21 Feb. 

1936. Accidental renaming of P. ruddi lebombo. 



Suborder SIMPLICIDENTATA 
Family ANOMALURID^E. Spiny-tailed Squirrels 

Subfamily ANOMALURIN^ 

Review: Allen, J. A. Sciuridse, Anomaluridse, and Idiuridae collected by the 
American Museum Congo Expedition. Bull. Amer. Mus. Nat. Hist., 47: 
39-71, pi. 5, 27 Oct. 1922. 

Anomalurella Matschie 

AnomalureUa Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 351, July 
1914. Subgenus? of Anomalurus, type Anomalurus pusillus Thomas. 

Anomalurella pusilla pusilla (Thomas) 

Anomalurus pusillus Thomas, Ann. Mag. Nat. Hist. (5) 20: 440, Nov. 1887. 
Bellima and Tingasi, Monbuttu ( = Niangara), Congo Beige. 

Anomalurella pusilla batesi (De Winton) 

Anomalurus batesi De Winton, Ann. Mag. Nat. Hist. (6) 20: 524, Dec. 1897. 
Como River, 75 miles from Gaboon. 

Anomalurops Matschie 

Anomalurops Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 351, July 1914. 
Subgenus of Anomalurus, type A. beecrofti Fraser. 

Anomalurops beecrofti beecrofti (Fraser) 

Anomalurus beecrofti Fraser, Proc. Zool. Soc. London, for 1852, p. 17, pi. 32, 

14 Nov. 1853. Island of Fernando Po. 
Anomalurus laniger Temminck, Esquisses Zool. sur la Cote de Guine, p. 149, 

1853 (young). Guinea coast. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 283 

Anomalurops beecrofti argenteus (Schwann) 

Anomalurus beecrofti argenteus Schwann, Ann. Mag. Nat. Hist. (7) 13:70, 
Jan. 1904. Abutschi, Niger River, about 150 miles from coast. 

Anomalurops beecrofti chapini J. A. Allen 

Ano7tialurops beecrofti chapini J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 
p. 65, 27 Oct. 1922. Medje, Congo Beige. 

.Anomalurops beecrofti citrinus (Thomas) 

Anomalurus beecrofti citrinus Thomas, Ann. Mag. Nat. Hist. (8) 18: 236, Aug. 
1916. Benito River, Spanish Guinea. 
Range: Lowlands from southern Cameroons to Gaboon. 



Anomalurus Waterhouse 

Anomalurus Waterhouse, Proc. Zool. Soc. London, for 1842, p. 124, Jan. 1843. 

Genotype Anomalurus fraseri Waterhouse. 
Arosethrus Waterhouse, Proc. Zool. Soc. London, for 1842, p. 124, footnote, 

Jan. 1843. Provisional substitute for Anomalurus Waterhouse. 

Anomalurus cinereus Thomas 

Anomalurus cinereus Thomas, Ann. Mag. Nat. Hist. (6) 15: 188, Feb. 1895. 
Upper Rovuma River, toward Lake Nyasa, Tanganyika Territory. 

Anomalurus erythronotus Milne-Edwards 

Anomalurus erythronotus Milne-Edwards, Compt. Rend. Acad. Sci., Paris, 

89: 771, 1879; Huet, Nouv. Arch. Mus. d'Hist. Nat., Paris, (2) 6: 285, 

pi. 21, 1884. Gaboon. 

Anomalurus fraseri fraseri Waterhouse 

Anomalurus fraseri Waterhouse, Proc. Zool. Soc. London, for 1842, p. 124, 
Jan. 1843. Fernando Po. 

Pteromys squamicaudus Schinz, Syst. Verzeichn. Saugeth., 2: 58, 1845 (re- 
naming of Anomalurus fraseri Waterhouse). Fernando Po. 

Anomalurus fraseri laticeps d'Aguilar-Amat, Butlleti Inst. Catalana d'Hist. 
Nat., Barcelona, 2: 53, pi. 1,"!. 1-lb, Mar. 1922. Pico de Santa Isabel, 
Fernando Po. (Syn. of A. fraseri, see Cabrera, Bol. Real Soc. Espanola 
Hist. Nat., Madrid, 23: 68, 28 Feb. 1923.) 

Anomalurus fraseri beldeni Du ChaiUu 

Anomalurus beldeni Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 303, July 
1860. Gaboon. 



284 bulletin: museum of comparative zoology 

Anomalurus fraseri chrysoph.entjs Dubois 

AnomaluTus chrysophsenus Dubois, Bull. Soc. Zool. de France, Paris, 13: 23, 
Jan. 1888. Landana, southern Congo. 

Anomalurus fraseri derbianus (Gray) 

Pteromys derbianus Gray, Ann. Mag. Nat. Hist. (1) 10: 262, Dec. 1842. Sierra 
Leone. s 

Anomalurus fraseri griselda Dollman 

Anomalurus fraseri griselda Dollman, Ann. Mag. Nat. Hist. (8) 14: 490, Dec. 
1914. Bitye, southern Cameroons. 

Anomalurus fraseri nigrensis Thomas 

Anomalurus fraseri nigrensis Thomas, Abstr. Proc. Zool. Soc. London, no. 10, 

p. 12, 22 Nov. 1904; Proc. Zool. Soc. London, for 1904, 2: 190, 18 Apl. 

1905. Abutschi, lower Niger River, Nigeria. 

Anomalurus fulgens Gray 

Anomalurus fidgens Gray, Ann. Mag. Nat. Hist. (4) 3: 467, June 1869; Alston, 
Proc. Zool. Soc. London, p. 96, pi. 21 (col.), June 1875. Gaboon. 

Anomalurus imperator Dollman 

Anomalurus imperator Dollman, Ann. Mag. Nat. Hist. (8) 8: 257, Aug. 1911. 
Bibianaha, 60 miles west of Kumasi, Gold Coast. 

Anomalurus jacksoni jacksoni De Winton 

Anomalurus jacksoni De Winton, Ann. Mag. Nat. Hist. (7) 1: 251, Mar. 1898. 
Entebbe, Uganda. 

Anomalurus jacksoni fortior Lonnberg 

Anomalurus jacksoni fortior Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 58: no. 2, p. 66, 1 Sep. 1917. Masisi, near Kivu, Congo 

Beige. 

Anomalurus jacksoni jordani St. Leger 

Anomalurus jacksoni jordani St. Leger, Novitates Zool., 39: 251, 30 Aug. 1935. 
Fazenda Congulu, Amboin district, Angola, 7-800 m. 

Anomalurus jacksoni perustus Thomas 

Anomalurus jacksoni perustus Thomas, Ann. Mag. Nat. Hist. (8) 18: 235, Aug. 
1916. River Lubefu, 75 miles north of Lusambo, South Congo. 

Anomalurus neavei Dollman 

Anomalurus neavei Dollman, Ann. Mag. Nat. Hist. (8) 3: 351, Apl. 1909. 
Kambove, Katanga, Congo. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 285 

Anomalurus orientalis Peters 

Anomalurus orientalis Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 
164, pi. (col.), 1880. Zanzibar. 

Subgenus Anomalurodon Matschie 

Anomalurodon Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 350, July 
1914. Subgenus of Anomalurus, type A. auzembergeri Matschie. 

Anomalurus auzembergeri Matschie 

A[nomaluriis] auzembergeri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 

350, July 1914. Near Patokla, middle Cavalle River, near boundary 

between Liberia and Ivory Coast. 

Anomalurus pelii (Schlegel and S. Miiller) 

Pteromys [Anomalurus) pelii Schlegel and S. Miiller, Verhandeel. over Natuurl. 
Geschiedn. Nederland. Bezittingen, Zool., publ. by Natuurkundige 
Commissie in Oostindie, C. J. Temminck, ed., 1: pt. 2, p. 109, 1843-5. 
Daboeram (=Dabacrom), Gold Coast. 

Subfamily IDIURIN.E 
Idiurus Matschie. Pygmy Scaly-tailed Squirrels 

Idiurus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 194, 1894. Genotype 
Idiurus zenkeri Matschie. 

Idiurus langi J. A. Allen 

Idiurus langi J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 69, pi. 5, 27 Oct. 
1922. Medje, Congo Beige. 

Idiurus macrotis Miller 

Idiurus macrotis Miller, Proc. Biol. Soc. Washington, 12: 73, text-f. 15-19, 
24 Mar. 1898. Efulen, Cameroons. 

Idiurus panga J. A. Allen 

Idiurus panga J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 70, 27 Oct. 1922. 
Panga, Congo Beige. 

Idiurus zenkeri zenkeri Matschie 

Idiurus zenkeri Matschie, Sitzb. Ges. Naturf. Freunde, p. 197, text-f., 1894; 
p. 25, text-f. 3, 1898. Yaunde, Cameroons. 

Idiurus zenkeri kivuensis Lonnberg 

Idiurus zenkeri kivuensis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., Stock- 
holm, (2) 58: no. 2, p. 67, 1 Sep. 1917. Masisi, Kivu district, Congo Beige. 



286 bulletin: museum of comparative zoology 



Zenkerella Matschie 

Zenkerella Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 23, text-f. 1, 2, 17 

May 1898. Genotype Zenkerella insignis Matschie. 
Aethurus De Winton, Proc. Zool. Soc. London, p. 450, pi. 34, 35, 1 Oct. 1898, 

Genotype Aethurus glirinus De 'Winton = Zenkerella insignis Matschie. 

Zenkerella insignis Matschie. Non-flying Scaly-tailed Squirrel. 
Zenkerella insignis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 24, text-f. 

1, 2, 17 May 1898. Yaunde, Cameroons. 
Aethurus glirinus De Winton, Proc. Zool. Soc. London, p. 451, pi. 34, 35, 1 Oct- 

1898. Benito River, French Congo. 



Family SCIURID^. Tree and Ground Squirrels 

See: Thomas, Oldfield. The generic arrangement of the African squirrels. 
Ann. Mag. Nat. Hist. (8) 3: 467^75, June 1909. 



iETHOsciURUs Thomas 

Mthosciurus Thomas, Ann. Mag. Nat. Hist. (8) 17: 271, Mar. 1916. Genotype 
Sciurus poensis A. Smith. 

^THOSCIURUS BYATTI BYATTI (Kcrshaw) 

Funisciurus byatti Kershaw, Ann. Mag. Nat. Hist. (9) 11: 592, May 1923. 

Moshi, Kilimanjaro, Tanganyika Territory. 
^thosciurus byatti Kershaw, Ann. Mag. Nat. Hist. (9) 11: 708, June 1923. 

^THOSciTjRUS BYATTI L^TUS Allen and Loveridge 

jEthosciurus byatti Iseius G. M. Allen and Loveridge, Bull. Mus. Comp. Zool., 

75: 96, Feb. 1933. Madehani, Ukinga Mts., north end of Lake Nyasa, 

Tanganyika Territory. 

iETHOSCiURUS LUCIFER (Thomas) 

Xerus (Paraxerus) lucifer Thomas, Proc. Zool. Soc. London, p. 430, 1 Oct. 

1897. Kombe Forest, Masuku Mts., Nyasaland, 7000 feet. 
Funisciurus lucifer Thomas, Proc. Zool. Soc. London, for 1897, p. 932, pi. 54 

(col.), 1 Apl. 1898. 

iETHOSCIURUS POENSIS POENSIS (A. Smith) 

Sciurus poensis A. Smith, South African Quart. Journ., 2: 128, 1834. Island of 
Fernando Po. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 287 

iExHOSciURUS poENSis MUSCULiNUS (Temminck) 

Sciurus musculinus Temminck, Esquisses Zool. sur la Cote de Guine, p. 142, 
1853. Guinea coast. 

^THOSCIURTJS POENSIS STJBVIRIDESCENS (LeCOnte) 

Sciurus subviridescens Leconte, Proc. Acad. Nat. Sci. Philadelphia, p. 11, Jan. 

1857. "Western Africa" = Gaboon. 
Sciurus olivaceus Milne-Edwards, Rev. et Mag. de Zool., (2) 19: 228, July 1867. 

Between Cape Palmas and the Calabar coast. West Africa. 
Mthosciurus poensis subviridescens Cabrera, Mem. Real Soc. Espanola Hist. 

Nat., Madrid, 16: 75, 10 July 1929. 

iExHOSCiURUS RUWENzoRii RUWENzORii (Schwann) 

Sciurus rufobrachiatus ruwenzorii Schwann, Ann. Mag. Nat. Hist. (7) 13: 71, 

Jan. 1904. Wimi Valley, Ruwenzori, Uganda border. 
See Thomas, Ann. Mag. Nat. Hist. (7) 19: 121, Jan. 1907 — not one of 
rufobrachiatus group. 
Sciurus ruwenzorii Thomas and Schwann, Trans. Zool. Soc. London, 19: 497, 

Mar. 1910. 

.SIthosciurus RUWENZORII vuLCANixjs (Thomas) 

Sciurus ruwenzorii vulcanius Thomas, Ann. Mag. Nat. Hist. (8) 4: 476, Nov. 

1909. Vulcan Forest, north of Lake Kivu, between Bajaka and Kassongo, 

Congo Beige. 

jEthoscitjrus vexillarius (Kershaw) 

Funisciurus vexillarius Kershaw, Ann. Mag. Nat. Hist. (9) 11, 591, May 1923. 

Wilhelmsthal, Lushoto, Usambara, Tanganyika Territory. 
j^thosdurus vexillarius Kershaw, Ann. Mag. Nat. Hist. (9) 11: 708, June 1923. 



Atlantoxerus Forsyth Major 

Atlantoxerus Forsyth Major, Proc. Zool. Soc. London, p. 189, pi. 8, f. 9; pi. 9, 
f. 9, June 1893. Subgenus of Xerus, type Xerus getulus (Linnaeus). 

Atlantoxerus getulus (Linnaeus) , 

Sciurus getulus Linnaeus, Syst. Nat.,ed. 10, 1: 64, 1758. "Africa;" type locality 

fixed as Agadir, Morocco, by Cabrera (Trab. Mus. Nac. Cienc. Nat., 

Madrid, zool. ser., no. 57, p. 217, pi. 8, 30 Dec. 1932). 
Xerus trivitiatus Gray, Ann. Mag. Nat. Hist. (1) 10: 264, Dec. 1842. "? India." 
Range : All the Grand Atlas from the Atlantic coast between Uad Tensif t and 

Uad Sus, at the extreme east of the chain extending to the middle Atlas 

and the Algerian Sahara. 



288 bulletin: museum of comparative zoology 



Epixerus Thomas 

Epixerus Thomas, Ann. Mag. Nat. Hist. (8) 3: 472, June 1909. Genotype 
Sciurus wilsoni Du Chaillu. 

Epixerus ebii (Temminck) 

Sciurus ebii Temminck, Esquisses Zool. sur la Cote de Guine, p. 129, 1853. 
Forests of Guinea: Dabocrom. 

Epixerus wilsoni (Du Chaillu) 

Sciurus wilsoni Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 364, Nov. 1860. 
Gaboon, headwaters of the Ovenga River. 



EuxERUS Thomas 

Euxerus Thomas, Ann. Mag. Nat. Hist. (8) 3: 473, June 1909. Genotype 
Sciurus erythropus E. Geoffroy. 

Euxerus erythropus erythropus (E. Geoflfroy) 

Sciurus erythropus E. Geoffroy, Cat. Mamm. Mus. d'Hist. Nat. Paris, p. 178, 

1803. Probably Senegal. 
Sciuriis alhovittatus Desmarest, Nouv. Dictionnaire d'Hist. Nat., ed. 2, 10: 110, 

1817. ? East Indies in part; probably western Africa. 
Sc[iums] marabutus Lesson, Complements de Buffon, ed. 2, Paris, 1 : 467, 1838. 

Senegal. 
Scliurus] prestigiator Lesson, Complements de Buffon, ed. 2, Paris, 1 : 468, 

1838. Senegal. 
Sciurus lessonii Fitzinger, Sitzb. K. Akad. Wiss., Wien, math.-nat. CI., 55: 

sect. 1, p. 488, 1867. New name for Sciurus marabutus Lesson. 

Euxerus erythropus agadius Thomas and Hinton 

Euxerus erythropus agadius Thomas and Hinton, Novitates Zool., 28 : 6, 14 May 
1921. Agades, northern Nigeria. 

Euxerus erythropus chadensis (Thomas) 

Xerus erythropus chadensis Thomas, Ann. Mag. Nat. Hist. (7) 15: 387, Apl. 
1905. Yo, Lake Chad. 

Euxerus erythropus fulvior (Thomas) 

Xerus microdon fulvior Thomas, Ann. Mag. Nat. Hist. (7) 15: 389, Apl. 1905. 
Fort Hall, Kenya Colony. 

Euxerus erythropus lacustris (Thomas) 

Xerus erythropus lacustris Thomas, Ann. Mag. Nat. Hist. (7) 15 : 388, Apl. 1905. 
Masindi, Unyoro, Uganda. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 289 

EUXERUS ERYTHROPUS LEUCOUMBRINUS (Riippell) 

Sciurus leucoumbrinus Riippell, Neue Wirbelth. zu d. Fauna Abyssinien 
gehorig, Saugeth., p. 38, 1835. Abyssinia, Sennaar, and Kordofan; re- 
stricted to Abyssinia, by Thomas (Proc. Zool. Soc. London, p. 256, foot- 
note, 6 July 1923). 

EuxERUs ERYTHROPUS LiMiTANEUs Thomas and Hinton 

Euxerus erythropus limitaneus Thomas and Hinton, Proc. Zool. Soc. London, 
p. 255, 6 July 1923. Zalingei, Darfur. 

Euxerus erythropus m^stus Thomas 

Euxerus erythropus msestus Thomas, Ann. Mag. Nat. Hist. (8) 5:419, May 1910. 
Daru, Sierra Leone, 150 miles inland. 

Euxerus erythropus microdon (Thomas) 

Xerus microdon Thomas, Ann. Mag. Nat. Hist. (7) 15: 389, Apl. 1905. Kitui, 
Kenya Colony. 

FuNisciURUS Trouessart 

Funisciurus Trouessart, Le Naturaliste, 2: 293, 1 Oct. 1880. As a subgenus 
of Sciurus; type by monotypy Sciurus isabella Gray = Funisciurus 
isabella (Gray); see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 51, 27 
Oct. 1922. Raised to generic rank by Thomas, Proc. Zool. Soc. London, 
p. 932, 1897. 

Funisciurus anerythrus anerythrus (Thomas) 

Sciurris pyrrhopus anerythrus Thomas, Proc. Zool. Soc. London, p. 447, pi. 40, 
Oct. 1890. Buguera, south of Lake Albert. 

Funisciurus anerythrus bandarum Thomas 

Funisciurus anerythrus bandarum Thomas, Ann. Mag. Nat. Hist. (8) 16: 146, 
Aug. 1915. Ba-mingui River, upper Shari, northeastern Congo Beige. 

Funisciurus anerythrus niapu J. A. Allen 

Funisciurus anerythrus niapu J. A. Allen, Bull. Amer. Mus. Nat. Hist., 47: 52, 
27 Oct. 1922. Niapu, Congo Beige. 

Funisciurus auriculatus auriculatus (Matschie) 

Sciurus auriculatus Matschie, Arch. f. Naturgesch., 75: sect. 1, pt. 3, p. 353, 
Oct. 1891. Kribi, Cameroons. 

Funisciurus auriculatus beatus Thomas 

Funisciurus auriculatus beatus Thomas, Ann. Mag. Nat. Hist. (8) 5: 196, Feb. 
1910. Benito River, 15 miles from the mouth, French Congo. 



290 bulletin: museum of comparative zoology 

FuNisciuRUS AURicuLATUs BOYDi Thomas 

Funisciurus auriculatus boydi Thomas, Ann. Mag. Nat. Hist. (8) 5: 196, Feb. 

1910. Mussaka, lower Mongo River, east of Cameroon Mt., Cameroons. 

Funisciurus auriculatus olivle Dollman 

Funisciurus auriculatus oliviae Dollman, Ann. Mag. Nat. Hist. (8) 8: 733, Dec. 

1911. Southern Nigeria, Oban district. 

Funisciurus bayonii (Bocage) 

Sciurus hayonii Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 2: 3, Sep. 
1890. Duque de Braganga, Angola. 

Funisciurus carruthersi carruthersx Thomas 

Funisciurus carruthersi Thomas, ^^Jin. Mag. Nat. Hist. (7) 18: 140, Aug. 1906. 
Kuwenzori East, 6500 ft., Uganda. 

Funisciurus carruthersi birungensis Gyldenstolpe 

Funisciurus carruthersi birungensis Gyldenstolpe, Ark. f. Zool., 19B: no. 6, p. 1, 

14 Dec. 1927. Mt. Karissimbi, Birunga Volcanoes, eastern Congo Beige, 

3600 m. 

Funisciurus carruthersi chrysippus Thomas 

Funisciurus carruthersi chrysippus Thomas, Ann. Mag. Nat. Hist. (9) 11: 522, 

Apl. 1923. Wabembe region, west of north end of Lake Tanganyika, 

Congo Beige. 

Funisciurus carruthersi tanganyik.e Thomas 

Funisciurus carruthersi tanganyikse Thomas, Ann. Mag. Nat. Hist. (8) 4: 477, 

Nov. 1909. Usumbura, north end of Lake Tanganyika, Urundi, Congo 

Beige. 

Funisciurus congicus congicus (Kuhl) 

Sciurus congicus Kuhl, Beitrage zur Zool., p. 66, 1820. "Congo."' 
Sciurus prsetextus Wagner, Schreber's Saugthiere, SuppL, 3: 216, 1843. No 
locahty. 

Funisciurus congicus flavinus Thomas 

Funisciurus congicus flavinus Thomas, Ann. Mag. Nat. Hist. (7) 13: 411, June 
1904. Capangombi, southern plateau of Angola. 

Funisciurus congicus interior Thomas 

Funisciurus congicus interior Thomas, Ann. Mag. Nat. Hist. (8) 18: 236, Aug. 
1916. Inkongo, or Ikongo, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 291 

FuNisciuRUS coNGicus cENONE Thomas 

Funisciurus congicus oenone Thomas, Proc. Zool. Soc. London, p. 297, 29 Apl. 
1926. Cunene Falls, Cunene River, Southwest Africa. 

Funisciurus congicus olivellus Thomas 

Funisciurus congicus olivellus Thomas, Ann. Mag. Nat. Hist. (7) 13: 410, 
June 1904. Cunga, Angola. 
May be a seasonal phase of the typical race (see Thomas, O., Ann. Mag. Nat. 
Hist. (8) 18: 237, footnote, Aug. 1916). 

Funisciurus Isabella (Gray) 

Sciurv^ isabella Gray, Proc. Zool. Soc. London, p. 180, pi. 24, Sep. 1862. Cam- 
eroon Mts., Cameroons, 7000 ft. 

Funisciurus lemniscatus (Leconte) 

Sciurus lemniscatus Leconte, Proc. Acad. Nat. Sci. Philadelphia, p. 11, Jan. 

1857. "Western Africa" = Gaboon. 
Sciurus sharpei Gray, Ann. Mag. Nat. Hist. (4) 12: 265, Sep. 1873. Gaboon. 

Funisciurus leucogenys (Waterhouse) 

Sciurus leucogenys Waterhouse, Ann. Mag. Nat. Hist. (1) 10: 202, footnote, 

Nov. 1842. Fernando Po. 
Sciurus erythrogenys Waterhouse, Proc. Zool. Soc; London, for 1842, p. 129, 

Jan. 1843. Fernando Po. 
See Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 45, 27 Oct. 1922, as to 
priority of <S. leucogenys Waterhouse. 

Funisciurus leucostigma leucostigma (Temminek) 

Sciurus leucostigma Temminek, Esquisses de Zool. sur la C6te de Guin6, p. 133, 

1853. Boutry River to borders of Ashanti country, "Guinea" = Gold 

Coast. 

Funisciurus leucostigma niveatus Thomas 

Funisciurus leucostigma niveatus Thomas, Ann. Mag. Nat. Hist. (9) 11: 522, 
Apl. 1923. Beoumi, 12 miles east of Bandama, Ivorj^ Coast. 

Funisciurus leucostigma talboti Thomas 

Funisciurus leucostigma talboti Thomas, Ann. Mag. Nat. Hist. (8) 4: 478, Nov. 
1909. Oban, southern Nigeria. 

Funisciurus mandingo mandingo Thomas 

Fuhisciurus mandingo Thomas, Ann. Mag. Nat. Hist. (7) 11: 79, Jan. 1903. 
Nianimaru, Gambia. Probably a race of F. leucostigma. 



292 bulletin: museum of comparative zoology 

FUNISCIURUS MANDINGO NIGRENSIS ThomaS 

Funisciurus mandingo nigrensis Thomas, Ann. Mag. Nat. Hist. (8) 4: 477, 
Nov. 1909. Abutschi, 150 miles up the Niger, Nigeria. 

Funisciurus mayumbicus Kershaw 

Funisciurus mayumbicus Kershaw, Revue Zool. Africaine, 11: 363, 27 Dec. 
1923. Ganda Sundi, Mayximbe Province, mouth of Congo, Congo Beige. 

Funisciurus mystax mystax De Winton 

Fimisciurus mystax De Winton, Ann. Mag. Nat. Hist. (7) 2: 9, July 1898. 
Benito River, Rio Muni. 

Funisciurus mystax ochrogaster Cabrera and Ruxton 
Funisciurus mystax ochrogaster Cabrera and Ruxton, Ann. Mag. Nat. Hist. (9) 
17: 597, May 1926. St. Joseph de Luluabourg, Congo Beige. 

Funisciurus poolii (Jentink) 

Sciurus poolii Jentink, Notes Leyden Mus., 28: 139, July 1906. Stanley Falls, 
Congo River, Congo Beige. 

Funisciurus pyrrhopus pyrrhopus (F. Cuvier) 

[Sciurus] pyrrhopus F. Cuvier, Hist. Nat. des Mammiferes, 4: pt. 66, pi. 240 
and text, Feb. 1833. "Fernando Po," but doubtless the adjacent main- 
land, for it is not like the race leucogenys of that island, but since the type 
"had been kept alive as a pet, it may easily have been taken to the island 
by natives" (see Thomas, O., Proc. Zool. Soc. London, p. 447, footnote, 
Oct. 1890). 

Funisciurus pyrrhopus akka De Winton 

Sciurus emini De Winton, Ann. Mag. Nat. Hist. (6) 16: 197, Aug. 1895. Tin- 
gasi, Monbuttu, Congo Beige. Not (SawrttsemimStuhlmann 1894, =Tam- 
isciis etnini (Stuhlmann). 

Funisciurus akka De Winton, Ann. Mag. Nat. Hist. (7) 4: 357, Nov. 1899. 
Tingasi, Monbuttu, Congo Beige. New name for Sciurus emini De Win- 
ton, 1895, not of Stuhlmann, 1894, which is Tamiscus emini (Stuhlmann). 

Funisciurus wintoni Neumann, Zool. Jahrb., Syst., 13: 547, 10 Oct. 1900. 
New name for Sciurus emini De Winton, 1895, not of Stuhlmaim, 1894. 

Funisciurus pyrrhopus leonis Thomas 

Funisciurus pyrrhopus leonis Thomas, Ann. Mag. Nat. Hist. (7) 15: 79, Jan. 
1905. Bo, Sierra Leone. 

Funisciurus pyrrhopus pembertoni Thomas 

Funisciurus pembcrtoniThomsiS, Arm.. Mag. Nat. Hist. (7) 14:201, Sep. 1904. 
Dondo, Cuanza River, Angola. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 293 

FuNisciURUS PYRRHOPUS RUBRiPES (Du Chaillu) 

Sciurus rubripes Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 366, Nov. 1860. 
Gaboon, "in the country watered by the rivers Nazareth, Ogobai, Fernand- 
Vaz, Rembo, and Ovenga . . . two degrees south of the equator." 

Macroxus pyrrhopus, var. erythrops Gray, Ann. Mag. Nat. Hist. (3) 20: 330, 
Nov. 1867. Gaboon. 

FuNisciuRTJS RAPTORUM Thomas 

Funisciurus raptorum Thomas, Ann. Mag. Nat. Hist. (7) 11:80, Jan. 1903. 
Forgados, lower Nigeria. 

Funisciurus substriatus De Winton 

Funisciurus substriatus De Winton, Ann. Mag. Nat. Hist. (7) 4: 357, Nov. 1899. 
Kintampo, Gold Coast hinterland. 



Geosciurus a. Smith 

Geosciurus A. Smith, South African Quart. Journ., 2: 128, Jan.-Mar. 1834. 
Genotj^je Xerus capensis A. Smith (see Thomas, Proc. Zool. Soc. London, 
p. 933, 1897). 

Geosciurus inauris inauris (Zimmermann) 

Sciurus inauris Zimmermann, Geogr. Geschichte, 2: 344, 1780. Kaffirland, 

100 miles north of Cape of Good Hope (based on Pennant). 
Sciurus capensis Kerr, Linnajus's Animal Kingdom, p. 266, 1792. Karoo north 

of Sneeuwberg, near Graaff Reinet. "Near the mountain of Sneeberg, 

eight hundred miles north from the Cape of Good Hope." 
Sciurus ginginianus Shaw, Gen. Zool., 2: pt. 1, p. 147, 1801. Gingi, East 

Indies, = South Africa. 
Sciurus dschinschicus Gmelin, in Linne's Syst. Nat., ed. 13, 1: pt. 1, p. 151, 

1788. "In provincia Indise Dschinschi"; but probably South Africa. 
Myoxus africanus Shaw, Gen. Zool., 2: pt. 1, p. 172, 1801. In mountains of 

Sneeburgh, 800 miles above Cape of Good Hope; (based on Pennant). 
Sciurus levaillantii Kuhl, Beitriige zur Zool., p. 67, 1820. "In Africa meri- 

dionali." 
Sciurus setosus Smuts, Enumeratio Mamm. Capensium, p. 33, 1832 (in part). 

Southern part of Cape of Good Hope. 

Geosciurus inauris namaquensis (Lichtenstein) 

Sciurus namaqxiensis Lichtenstein, Cat. Rer. Nat., p. 2, 1793. Great Namaqua- 

land, west of Fish River, Cape of Good Hope. 
Geosciurus capensis namaquensis Thomas and Hinton, Proc. Zool. Soc. London, 

p. 489, 3 Sep. 1923 (valid race). 



294 bulletin: museum of comparative zoology 

Geosciurus inauris princeps Thomas 

Geosciurus princeps Thomas, Proc. Zool. Soc. London, p. 106, 30 Apl. 1929. 
Otjitundua, central Kaokoveld, Southwest Africa. 



Heliosciurus Trouessart 

Heliosciurus Trouessart, Le Naturaliste, 2: no. 37, p. 292, 1 Oct. 1880. Sub- 
genus of Sciurus, type Scmrus annulatus Desmarest (see Thomas, Proc. 
Zool. Soc. London, p. 933, 1897). 

Review: Ingoldby, C. M. Proc. Zool. Soc. London, p. 471-487, pi. 1-4 
(col.), 12 July 1927. 

Thomas, O. The West African forms of Heliosciurus rufobrachium. Ann. Mag. 
Nat. Hist. (9) 11: 523-526, Apl. 1923. 

Heliosciurus bongensis bongensis (Heuglin) 

Sciurus bongensis Heuglin, Reise in Nordost-Afrika, 2: 59, 1877. Bahr-el- 
Ghazal, Anglo-Egj^Dtian Sudan. 

Heliosciurus bongensis canaster Thomas and Hinton 
Heliosciurus bongensis canaster Thomas and Hinton, Proc. Zool. Soc. London, 
p. 256, 6 July 1923. Foothills of Jebel Marra, Darfur. 

Heliosciurus gambianus gambianus (Ogilby) 

Sciurus annulatus Desmarest, Encyclop. Methodique, Mammalogie, 2: 338, 

1822. No locality. 
If identifiable, this name may replace gambianus. 
Sciurus gambianus Ogilby, Proc. Zool. Soc. London, p. 103, 9 Oct. 1835. 

Gambia, probably near Ft. St. Mary. 
Sc[iurus] simplex Lesson, Complements de Buffon, ed. 2, Paris, 1: 467, 1838. 

Senegal. Probably a sjmonym of H. gambianus. 
Sciurus annularis Schinz, Syst. Verzeichn. Saugethiere, 2: 14, 1845. No 

locality; perhaps not African. 
Macroxus annulatus, var. albina Gray, Ann. Mag. Nat. Hist. (3) 20: 329, 

Nov. 1867, nomen nudum. West Africa. 

Heliosciurus gambianus limbatus Schwarz 

Heliosciurus gambianus limbatus Schwarz, Jahrb. d. Nassauischen Ver. f. 
Naturk., Wiesbaden, 68: 65, 1915. Bosum, Cameroons. 

Heliosciurus gambianus senescens Thomas 

Heliosciurus gambianus senescens Thomas, Ann. Mag. Nat. Hist. (8) 4: 544, 
Dec. 1909. Thies, Senegal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 295 

Heliosciurus multicolor multicolor (Riippell) 

Sciurus multicolor Riippell, Neue Wirbelthiere z. d. Fauna Abyssinien gehorig, 

Saugeth., p. 38, pi. 13 (col.), 1835. Valleys of Kulla and east slope of the 

coast range, Abyssinia. 

Heliosciurus multicolor abassensis (Neumann) 

Sciurus abassensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 57, 1902. 
Forests on south shore of Lake Abassi, Ethiopia. 

Heliosciurus multicolor cocnosus Thomas 

Heliosciurus multicolor ccenosus Thomas, Ann. Mag. Nat. Hist. (8) 4: 104, 
Aug. 1909. Ubangi River, central Africa, 19° 30' east. 

Heliosciurus multicolor dysoni St. Leger 

Heliosciurus multicolor dysoni St. Leger, Ann. Mag. Nat. Hist. (10) 19: 527, 

May 1937. Lodwar, about forty miles above mouth of Turkwel River, 

Lake Rudolf region, Uganda. 

Heliosciurus multicolor elegans Thomas 

Heliosciurus multicolor elegans Thomas, Ann. Mag. Nat. Hist. (8) 4: 103, 
Aug. 1909. Mt. Elgon, Kenya Colony. 

Heliosciurus multicolor eaffensis (Neumann) 

Sciurus kaffensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 57, 1902. 
Anderatscha, Kaffa, (Abyssinia) Ethiopia. 

Heliosciurus multicolor lateris Thomas 

Heliosciurus multicolor lateris Thomas, Ann. Mag. Nat. Hist. (8) 4: 102, Aug. 
1909. Lado. 

Heliosciurus multicolor madog.e Heller 

Heliosciurus multicolor madogae Heller, Smithsonian Misc. Coll., 56: no. 17, p. 1, 
28 Feb. 1911. Uma, 50 miles north of Nimule, Uganda. 

Heliosciurus multicolor omensis Thomas 

Heliosciurus multicolor omensis Thomas, Ann. Mag. Nat. Hist. (8) 4: 104, 

Aug. 1909. Kum Dingani, east of north end of Lake Rudolf, Lower Omo, 

Abyssinia. 

Heliosciurus mutabilis mutabilis (Peters) 

Sciurus mutabilis Peters, Reise nach Mossambique, Saugeth., p. 131, pi. 30; 

pi. 32, f. 2, 1852; Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 273, 1852. 

Boror, Mozambique, 17° S. 



296 bulletin: museum of comparative zoology 

Heliosciurus mutabilis beir^ Roberts 

Heliosciurus mutabilis beirse Roberts, Ann. Transvaal Mus., 4: 78, 20 Oct. 1913. 
Zimbiti, Beira, Mozambique. 

Heliosciurus mutabilis shirensis (Gray) 

Macroxus shirensis Gray, Ann. Mag. Nat. Hist. (3) 20: 327, Nov. 1867. Shire 
River, Rhodesia. 

Heliosciurus mutabilis chirindensis Roberts 

Heliosciurus mutabilis chirindensis Roberts, Ann. Transvaal Mus., 4: 78, 20 
Oct. 1913. Chirinda Forest, eastern Rhodesia. 

Heliosciurus mutabilis vumb^ Roberts 

Heliosciurus mutabilis vumbse Roberts, Ann. Transvaal Mus., 19: 100, 8 Nov. 
1937. Vumba, Southern Rhodesia. 

Heliosciurus punctatus punctatus (Temminck) 

Sciurus punctatus Temminck, Esquisses Zool. sur la Cote de Guin6, p. 138, 
1853. Guinea coast; no exact locality. 

Heliosciurus punctatus savannius Thomas 

Heliosciurus punctatus savannius Thomas, Ann. Mag. Nat. Hist. (9) 11: 521, 
Apl. 1923. Beoumi, Ivory Coast. 

Heliosciurus rhodesi^ rhodesi.e (Wroughton) 

Funisciurus annulatus rhodesise Wroughton, Mem. and Proc. Manchester Lit. 

and Phil. Soc, 51 : no. 5, p. 15, 13 Mar. 1907. Road to Chewalla's, high 

plateau west of Mchinga escarpment, Rhodesia. 

Heliosciurus rhodesise loandicus Thomas 

Heliosciurus rhodesise loandicus Thomas, Ann. Mag. Nat. Hist. (9) 11: 521, 
Apl. 1923. N'Dalla Tando, northern Angola. 

Heliosciurus rufobrachium rufobrachium (Waterhouse) 
Sc[iurus] rufo-brachium Waterhouse, Ann. Mag. Nat. Hist. (1) 10: 202, foot- 
note, Nov. 1842. Island of Fernando Po. 
Sciurus rufo-brachiatus Waterhouse, Proc. Zool. Soc. London, for 1842, p. 128, 
1843. Island of Fernando Po. 
See Thomas, O., Ann. Mag. Nat. Hist. (9) 11: 518, footnote, Apl. 1923; 
Allen, J. A., Bull. Amer. Mus. Nat. Hist., 47: 45, footnote, 27 Oct. 1922. 

Heliosciurus rufobrachium acticola Thomas 

Heliosciurus rufobrachium acticola Thomas, Ann. Mag. Nat. Hist. (9) 11: 525, 
Apl. 1923. Santa Isabel, Fernando Po. 



ALLEN: CHECKLIST OF AFKICAN MAMMALS 297 

Heliosciurus rufobrachium arrhenii Lonnberg 

Heliosciurus rufobrachiatus arrhenii Lonnberg, Kungl. Svenska Vet.-Akad. 

Handl., Stockholm, (2) 58: no. 2, p. 68, 1 Sep. 1917. Masisi, near Kivu 

Congo Beige. 

Heliosciurus rufobrachium aubryi (Milne-Edwards) 

Sciurus aubryi A. Milne-Edwards, Revue de Zool. Pure et Appliqu^e, (2) 19: 
228, July 1867. Gaboon. 

Heliosciurus rufobrachium benga Cabrera 

Heliosciurus rufobrachiatus benga Cabrera, Bol. Real Soc. Espanola Hist. Nat., 
Madrid, 17: 517, Nov. 1917. Cape San Juan, Spanish Guinea. 

Heliosciurus rufobrachium brauni St. Leger 

Heliosciurus riifobrachiatus brauni St. Leger, Novitates Zool., 39: 252, 30 Aug. 
1935. Fazenda Congulu, Amboin district, Angola, 800 m. 

Heliosciurus rufobrachium caurinus Thomas 

Heliosciurus rufobrachium caurinus Thomas, Ann. Mag. Nat. Hist. (9) 11: 
523, Apl. 1923. Gunnal, Portuguese Guinea. 

Heliosciurus rufobrachium emissus Thomas 

Heliosciurus rufobrachium emissus Thomas, Ann. Mag. Nat. Hist. (9) 11: 520, 
Apl. 1923. Kumbo, southeastern Nigeria, about 6° 50' north, 10° 50' east. 

Heliosciurus rufobrachium hardyi Thomas 

Heliosciurus rufobrachium hardyi Thomas, Ann. Mag. Nat. Hist. (9) 11: 519, 
Apl. 1923. Beoumi, 250 miles north of Grand Bassam, Ivory Coast. 

Heliosciurus rufobrachium isabellinus (Gray) 

Macroxus isabellinus Gray, Ann. Mag. Nat. Hist. (3) 20:329, Nov. 1867. "West 
Africa," probably lower Niger. 

Heliosciurus rufobrachium keni^e (Neumann) 

Sciurus kenise Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 176, 1902. 
West slope of Mt. Kenya, Kenya Colony, 8000 ft. 

Heliosciurus rufobrachium leonensis Thomas 

Heliosciurus rufobrachium leonensis Thomas, Ann. Mag. Nat. Hist. (9) 11: 523, 
Apl. 1923. Mano, Sierra Leone. 

Heliosciurus rufobrachium lualab^ Thomas 

Heliosciurus rufobrachium lualabse Thomas, Ann. Mag. Nat. Hist. (9) 11:520, 
Apl. 1923. Lodja, upper Lukenye, southern Congo Beige. 



298 bulletin: museum of comparative zoology 

Heliosciurus rufobrachium maculatus (Temminck) 

Sciurus maculatus Temminck, Esquisses Zool. sur la Cote de Guine, p. 130, 
1853. Guinea Coast. 

Macroxus rufobrachiatus, var. waterhousii Gray, Ann. Mag. Nat. Hist. (3) 
20: 328, Nov. 1867. Ashanti. 

Sciurus {Heliosciurus) rufobrachiatus libericus Miller, Proc. Washington Acad. 
Sci., 2: 633, text-f. 39a, 28 Dec. 1900. Mount Coffee, Liberia. 

Sciurus aschantiensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 175, 
1902. Ashanti. Synonym of H. rufobrachium tnaculatus (Temminck) ac- 
cording to Thomas (1923). 

Heliosciurus rufobrachium medjianus J. A. Allen 

Heliosciurus rufobrachium medjianus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 
47: 48, 27 Oct. 1922. Medje, Congo Beige. 

Heliosciurus rufobrachium nyans^ (Neumann) 

Sciurus nyansse Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 56, 1902. 
Kwa Kitoto, Kavirondo, Uganda. 

Heliosciurus rufobrachium obfuscatus Thomas 

Heliosciurus rufobrachima obfuscatus Thomas, Ann. Mag. Nat. Hist. (9) 11: 
526, Apl. 1923. Ekkonnanakku, Oban district, southeastern Nigeria. 

Heliosciurus rufobrachium pasha (Schwann) 

Sciurus rufobrachiatus pasha Schwann, Ann. Mag. Nat. Hist. (7) 13: 72, Jan. 
1904. BeUima, Monbuttu (=Niangara), Congo Beige. 

Heliosciurus rufobrachium rubricatus J. A. Allen 

Heliosciurus rufobrachium rubricatus J. A. Allen, Bull. Amer. Mus. Nat. Hist., 

47: 50, 27 Oct. 1922. Near Lubila River, 50 miles southwest of Avakubi, 

Congo Beige. 

Heliosciurus rufobrachium semlikii (Thomas) • 

Sciurus rufobrachiatus semlikii Thomas, Ann. Mag. Nat. Hist. (7) 19: 120, 
Jan. 1907. Beni, Semliki River, Congo Beige. 

Heliosciurus undulatus undulatus (True) 

Sciurus undulatus True, Proc. U. S. Nat. Mus., 15: 465, text-f. 3, 26 Oct. 1892. 

Mt. Kilimanjaro and Kahe, south of it, Tanganyika Territory. 
Heliosciurus undulatus marwitzi F. Miiller, Zool. Anzeiger, 37: 82, 76, text-f. 1, 

4, 31 Jan. 1911. Mt. Kilimanjaro, Tanganyika Territory. 

Heliosciurus undulatus daucinus Thomas 

Heliosciurus undulatus daucinus Thomas, Ann. Mag. Nat. Hist. (8) 4: 101, 
Aug. 1909. Mombasa, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 299 

Heliosciurus undulatus dolosus Thomas 

Heliosciurus undulatus dolosus Thomas, Ami. Mag. Nat. Hist. (8) 4: 100, Aug. 
1909. Mafia Id., off Tanganyika Territory. 

Helioscitjeus tjndulatxjs shindi Heller 

Heliosciurus rufobrachiattis shindi Heller, Smithsonian Misc. Coll., 63: no. 7, 

p. 7, 24 June 1914. Summit of Mt. Umengo, Taita Hills, Kenya Colony, 

6000 ft. 

Myrsilus Thomas 

Myrsilus Thomas, Ann. Mag. Nat. Hist. (8) 3: 470, June 1909. Genotype 
Macroxus aubinnii Gray. 

Myrsilus aubinnii aubinnii (Gray) 

Macroxus aubinnii Gray, Arm. Mag. Nat. Hist. (4) 12: 65, July 1873. Fanti. 

Myrsilus aubinnii sal^ (Jentink) 

Sciurus salse Jentink, Notes Leyden Mus., 3: 63, Apl. 1881. Bavia and Sofore- 
Place, St. Paul's River, Liberia. 



Myosciurus Thomas. Pygmy Squirrels 

Myosciurus Thomas, Ann. Mag. Nat. Hist. (8) 3: 474, June 1909. Genotype 
Sciurus minutus Du Chaillu = Sciurus pumilio Leconte. 

Myosciurus pumilio (Leconte) 

Sciurus 'pumilio Leconte, Proc. Acad. Nat. Sci. Philadelphia, p. 11, Jan. 1857. 

"Western Africa" = Gaboon. 
Sciurus minutus Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 366, Nov. 1860 

(not of Lartet, 1851, see Hollister, Proc. Biol. Soc. Washington, 34: 135, 

30 June 1921). Headwaters of the Ovenga River, Gaboon. 
Myosciurus minutulus Hollister, Proc. Biol. Soc. Washington, 34: 135, 30 June 

1921. Substitute for M. minutus (Du Chaillu). 



Paraxerus Forsyth Major. Bush Squirrels 

Paraxerus Forsyth Major, Proc. Zaol. Soc. London, p. 189, pi. 8, f. 22; pi. 9, 
f. 23, I June 1893. A subgenus of Xerus, type Xerus cepapi (A. Smith). 

Paraxerus cepapi cepapi (A. Smith) 

Sciurus cepapi A. Smith, Report Exped. for Exploring Central Africa, from 

Cape of Good Hope, June 23, 1834, p. 43, 1836. "Banks of the Marikwa," 

i.e., Marico River, Rustenburg district, Transvaal. 
Sciurus cepate Gray, List Spec. Mamm. Brit. Mus., p. 140, 13 May 1843. 



300 bulletin: museum of comparative zoology 

Paraxerus cepapi auriventris Roberts 

Paraxerus cepapi auriventris Roberts, Ann. Transvaal Mus., 11: 250, 14 Sep, 
1926. Magudi, Mozambique. 

Paraxerus cepapi chobiensis Roberts 

Paraxerus cepapi chobiensis Roberts, Ann. Transvaal Mus., 15: 9, 1 Oct. 1932. 
Kabulabula, Chobe River, Bechuanaland. 

Paraxerus cepapi kalaharicus Roberts 

Paraxerus cepapi kalaharicus Roberts, Ann. Transvaal Mus., 15: 10, 1 Oct, 
1932. Mabeleapudi, Bechuanaland. 

Paraxerus cepapi maunensis Roberts 

Paraxerus cepapi maunensis Roberts, Ann. Transvaal Mus., 15: 9, 1 Oct. 1932, 
Maun, Ngamiland, Bechuanaland. 

Paraxerus cepapi phal^na Thomas 

Paraxerus cepapi phalxna Thomas, Proc. Zool. Soc. London, p. 296, 29 ApL 

1926. Forest between Ukuambi and Ondongwa, Ovamboland, Southwest 

Africa. 

Paraxerus cepapi quotus Wroughton 

Paraxerus cepapi quotus Wroughton, Ann. Mag. Nat. Hist. (8) 3: 516, June 
1909. Katanga district, Congo Beige. (Key.) 

Paraxerus cepapi sindi Thomas and Wroughton 

Paraxerus cepapi sindi Thomas and Wroughton, Proc. Zool. Soc. London, p, 
543, 30 Oct. 1908. Tette, Zambesia, Mozambique. 

Paraxerus cepapi soccatus Wroughton 

Paraxerus cepapi soccatus Wroughton, Ann. Mag. Nat. Hist. (8) 3: 515, June 
1909. Vwaza, Hewe River, northern Angoniland, Mozambique. 

Paraxerus flavivittis flavivittis (Peters) 

Sciurus flavivittis Peters, Reise nach Mossambique, Saugeth., p. 129, pi. 29; 32, 
f. 1, 1852. Mossimboa, 11° S., Mozambique (designated as type locality 
by Thomas, Ann. Mag. Nat. Hist. (9) 4: 32, July 1919). 

Paraxerus flavivittis exgeanus Hinton 

Paraxerus flavivittis exgeanus HLnton, Ann. Mag. Nat. Hist. (9) 5: 311, Mar. 
1920. Kilwa Kisiwani, Tanganyika Territory. 

Paraxerus flavivittis ibeanus Hinton 

Paraxerus flavivittis ibeanus Hinton, Ann. Mag. Nat. Hist. (9) 5: 312, Mar. 
1920. Mombasa, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 301 

Paraxerus flavivittis mossambicus Thomas 

Paraxerus flavivittis mossambicus Thomas, Ann. Mag. Nat. Hist. (9) 4: 31, 
July 1919. Lumbo, Mozambique. 
On seasonal variation in color see Hinton, M. A. C, Ann. Mag. Nat. Hist. 
(9) 5: 308-310, Mar. 1920. 

Paraxerus ochraceus ochraceus (Huet) 

Sciurus ochraceus Huet, Nouv. Arch. Mus. d'Hist. Nat. Paris, (2) 3: 154, pi. 7, 
f. 2, 1880. Bagamoyo, Tanganyika Territory. 

Paraxerus ochraceus affinis (Trouessart) 

Xerus (Paraxerus) poensis sp. affinis Trouessart, Cat. Mamm. Viv. Foss., pt. 3, 
p. 406, 1897. Based on Sciurus sp. Rhoads (Proc. Acad. Nat. Sci. Philadel- 
phia, p. 521, 1896), an immature Paraxerus from Marsabit, Kenya Colony. 
It is not clear here whether Trouessart intended to name this form or to 
indicate its relationship to Xerus poensis, but probably the former, as the 
next reference shows. 

Funisciurus poensis affinis Trouessart (ex Rhoads), Cat. Mamm. Viv. Foss., 
Suppl., pt. 2, p. 304, 1904. Marsabit, Kenya Colony. This name evidently 
supersedes percivali and angustus for a race of Paraxerus from the same 
locality. 

Paraxerus percivali Dollman, Ann. Mag. Nat. Hist. (8) 8: 653, Nov. 1911. 
Marsabit, Kenya Colony. 

Paraxerus ochraceus augustus Dollman, Ann. Mag. Nat. Hist. (8) 8: 654, Nov. 
1911. Marsabit, Kenya Colony. A synonym of Paraxerus percivali as 
shown by Hollister (Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 20, 1919). 

Paraxerus ochraceus animosus Dollman 

Paraxerus ochraceus animosus Dollman, Ann. Mag. Nat. Hist. (8) 8: 655, Nov. 
1911. Mt. Nyiro, Kenya Colony. 

Paraxerus ochraceus aruscensis (Pagenstecher) 

Sciurus cepapi var. aruscensis Pagenstecher, Jahrb. Hamburg. Wiss. Anstalt., 

2: 42, 1885. Pangani River near the coast and Great Aruscha, Meru Mts., 

Tanganyika Territory. 

Paraxerus ochraceus electus Thomas 

Paraxerus ochraceus electus Thomas, Ann. Mag. Nat. Hist. (8) 4: 106, Aug. 1909. 
Elgeyo, Kenya Colony. 

Paraxerus ochraceus ganana (Rhoads) 

Sciurus ganana Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 522, 8 Dec. 1896. 
Ganana River at Bar Madu, Abyssinia. 



302 bulletin: museum of comparative zoology 

Paraxerus ochraceus jacksoni (De Winton) 

Sciurus jacksoni De Winton, Ann. Mag. Nat. Hist. (6) 19: 574, May 1897> 

Kikuyu, Kenya Colony. 
Paraxerus jacksoni capitis Thomas, Ann. Mag. Nat. Hist. (8) 4: 105, Aug. 1909.. 

Nairobi forest, Kenya Colony. 

Paraxerus ochraceus kahari Heller 

Paraxerus kahari Heller, Smithsonian Misc. Coll., 56: no. 17, p. 2, 28 Feb. 1911.. 
Meru Boma, northeast of Mt. Kenya, Kenya Colony. 

Paraxerus ochraceus salutans Thomas 

Paraxerus ochraceus salutans Thomas, Ann. Mag. Nat. Hist. (8) 4: 106, Aug. 
1909. Dar-es-Salaam, Tanganyika Territory. 

Paraxerus palliatus palliatus (Peters) 

Sciurus palliatus Peters, Monatsber. K. Preuss. Akad. Wiss., Berlin, p. 273, 
May 1852; Reise nach Mossambique, Zool., 1: Saugethiere, p. 134, pi. 31, 
f. 1; pi. 32, f. 3, 1852. Mainland near Mossambique Island, 15° S., and 
Mossimboa, 11° S., Portuguese East Africa. 

Paraxerus palliatus barawensis (Neumann) 

Sciurus palliatus baraivensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 178, 1902. Barawa, coast of southern Somaliland. 

Paraxerus palliatus frerei (Gray). Zanzibar Bush Squirrel 

Macroxus annulatus, var. frerei Gray, Ann. Mag. Nat. Hist. (4) 12: 265, Sep. 

1873. Zanzibar. 
Funisciurus palliatus lastii Thomas, Ann. Mag. Nat. Hist. (7) 18: 297, Oct. 

1906. Zanzibar Island. 
Sciurus palliatus suahelicus Neumann, Sitzb. Ges. Naturf. Freunde, p. 178, 

1902. Tanganyika Territory, opposite Zanzibar and Kenya Colony to 

Mombasa; no type locality assigned. 
According to Thomas and Wroughton (Proc. Zool. Soc. London, p. 293-294, 
1 Aug. 1907) this becomes a synonym of P. p. frerei (Gray). 

Paraxerus palliatus ornatus (Gray) 

Sciurus ornatus Gray, Proc. Zool. Soc. London, p. 13, pi. 1, July 1864. Natal. 

Paraxerus palliatus swynnertoni (Wroughton) 

Funisciurus palliatus swynnertoni Wroughton, Ann. Mag. Nat. Hist. (8) 1: 305, 
Apl. 1908. Chirinda Forest, northeastern Transvaal. 

Paraxerus palliatus tan^ (Neumann) 

Sciurus palliatus tanx Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 178,, 
1902. Middle Tana River, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 303 

Paraxerus paxjli (Matschie) 

Sciurus pauli Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 256, 1894. 
Derema, Tanga district, Tanganyika Territory. 

Paraxerus sponsxjs sponsus (Thomas and Wroughton) 
Funisdurus sponsus Thomas and Wroughton, Proc. Zool. Soc. London, p. 292, 
1 Aug. 1907. Coguno, Inhambane, Mozambique. 

Paraxerus sponsus bridgemani Dollman 

Paraxerus bridgemani Dollman, Ann. Mag. Nat. Hist. (8) 14: 152, Aug. 1914. 
Indook, Panda, Mozambique. 

Paraxerus sponsus tongensis Roberts 

Paraxerus sponsus tongensis Roberts, Ann. Transvaal Mus., 14: 229, 28 July 
1931. Mangusi Forest, northern Zululand, Natal. 

Paraxerus yulei (Thomas) 

Funisdurus yulei Thomas, Proc. Zool. Soc. London, 1: 120, 1 June 1902. 
Muezo, near Lake Mweru, Nyasaland. 



Protoxerus Forsyth Major. Giant Squirrels 

Protoxerus Forsyth Major, Proc. Zool. Soc. London, p. 189, pi. 8, f. 7; pi. 9, f. 
7, 1 June 1893. Subgenus of Xerus, type Sciurus stangeri Waterhouse (see 
Thomas, Proc. Zool. Soc. London, for 1897, p. 933, 1 Apl. 1898). 

Protoxerus stangeri stangeri (Waterhouse) 

Sciurus stangeri Waterhouse, Proc. Zool. Soc. London, for 1842, p. 127, Jan. 

1843. Fernando Po. 
Sciurus nordhoffi Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 363, Nov. 1860. 

Gaboon; "near the Ashira prairies," 150 miles from the coast, 1° S. 
Sciuru.s subalbidus Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 365, Nov. 1860. 

Island of Fernando Po. 

Protoxerus stangeri bea Heller 

Protoxerus stangeri bea Heller, Smithsonian Misc. Coll., 59: no. 16, p. 2, 5 July 
1912. Lukosa River, Kakamega Forest, Kenya Colony. 

Protoxerus stangeri centricola (Thomas) 

Sciurus stangeri centricola Thomas, Ann. Mag. Nat. Hist. (7) 18: 295, 297, Oct. 
1906. Entebbe, Uganda. 

Protoxerus stangeri dissonus Thomas 

Protoxerus stangeri dissonus Thomas, Ann. Mag. Nat. Hist. (9) 11: 527, Apl. 
1923. Bitye, Ja River, Cameroons. 



304 bulletin: museum of comparative zoology 

Protoxerus stangeri eborivorus (Du Chaillu) 

Sciurus eborivorus Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 363, Nov. 1860. 

Gaboon, mountains near source of Ovenga River, about 140 miles from 

coast, 1° S. 
Sciurus calliurus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 707, 1874. 

Mungo, Cameroons. 

Protoxerus stangeri loand.e (Thomas j 

Sciurus stangeri loandse Thomas, Ann. Mag. Nat. Hist. (7) 18: 295, Oct. 1906. 
Canhoca, northern Angola. 

Protoxerus stangeri mcerens Thomas 

Protoxerus stangeri moerens Thomas, Ann. Mag. Nat. Hist. (9) 11: 527, Apl. 
1923. Lobi, near Angu, middle Uele River, Congo Beige, about 24" east. 

Protoxerus stangeri Nigeria (Thomas) 

Sciurus stangeri nigerise Thomas, Ann. Mag. Nat. Hist. (7) 18: 295, 296, Oct. 
1906. Abutschi, lower Niger, Nigeria. 

Protoxerus stangeri notabilis Thomas 

Protoxerus stangeri notabilis Thomas, Ann. Mag. Nat. Hist. (9) 11: 528, Apl. 
1923. Avakubi, Ituri region, Congo Beige. 

Protoxerus stangeri personatus Kershaw 

Protoxerus stangeri personatus Kershaw, Revue Zool. Africaine, 11 : 364, 27 Dec. 

1923. Makaia N'tete, lower Mayumbe, near mouth of Congo, north bank, 

Congo Beige. 

Protoxerus stangeri signatus Thomas 

Protoxerus stangeri signatus Thomas, Ann. Mag. Nat. Hist. (8) 5: 85, Jan. 1910. 
Lodja, upper Lukenye River, southern Congo Beige. 

Protoxerus stangeri temminckii (Anderson) 

Sciurus caniceps Temminck, Esquisses Zool. sur la Cote de Guin6, p. 127, 1853. 

Near borders of Fanti country. Not Sciurus caniceps Gray. 
Sciurus temminckii Anderson, Zool. Researches Two Exped. Western Yunnan, 

p. 229, footnote, 1879. New name for »S. caniceps Temminck, not Gray. 

Protoxerus stangeri torrentium Thomas 

Protoxerus stangeri torrentium Thomas, Ann. Mag. Nat. Hist. (9) 11: 529, Apl. 
1923. East of Stanley Falls, eastern Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 305 



Tamiscus Thomas. African Striped Squirrels 

Tamiscus Thomas, Ann. Mag. Nat. Hist. (9) 1: 33, Jan. 1918. Genotype 
Sciurus emini Stuhlmann. 

Thomas, O. On the striped squirrels hitherto referred to the genus Paraxerus. 
Ann. Mag. Nat. Hist. (9) 1 : 33-38, Jan. 1918. 

Tamiscus alexandri (Thomas and Wroughton) 

Funisciurus alexandri Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 

376, May 1907. Gudima, River Iri, upper Uele, Congo Beige. 
Range: Uele and Ituri Rivers, eastward into Uganda. 

Tamiscus antoni^e (Thomas and Wroughton) 

Funisciurus antonise Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 

377, May 1907; Trans. Zool. Soc. London, 19: 498, pi. 23, upper f. (col.), 
Mar. 1910. Ponthierville, above Stanley Falls, Congo Beige. 

Tamiscus boehmi (Reichenow) 

Sciurus boehmi Reichenow, Zool. Anzeiger, 9: 315, 17 May 1886. Marungu, 
Tanganyika Territory. 

Tamiscus emini emini (Stuhlmann) 

Sciurus emini Stuhlmann, Mit Emin Pascha ins Herz von Afrika, p. 320, 1894. 

Atyangara, Semliki River, Uganda. 
Sciurus emini ugandx Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 180, 

1902. Kampala and Entebbe, Uganda. 
Syn. of T. emini, see Thomas, Trans. Zool. Soc. London, 19: 498, Mar. 1910; 
Dollman, Revue Zool. Africaine, 4: 79, 25 July 1914. 

Tamiscus emini gazelle Thomas 

Tamiscus emini gazellse Thomas, Ann. Mag. Nat. Hist. (9) 1 : 34, Jan. 1918. 
Meridi, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 

Tamiscus vulcanorum vulcanorum Thomas 

Tamiscus vulcanorum vulcanorum Thomas, Ann. Mag. Nat. Hist. (9) 1 : 35, 
Jan. 1918. Buhamba, near Lake Kivu, Congo Beige. 

Tamiscus vulcanorum lunaris Thpmas 

Tamiscus vulcanorum lunaris Thomas, Ann. Mag. Nat. Hist. (9) 1 : 36, Jan. 
1918. Mubuku Valley, Ruwenzori East, Uganda, 6500 ft. 

Tamiscus vulcanorum tanganyik^e Thomas 

Tamiscus vulcanorum tanganyikae Thomas, Ann. Mag. Nat. Hist. (9) 1 : 36, Jan. 

1918. Burton Gulf, 10 miles west of Baraka, Lake Tanganyika, Congo 

Beige. 



306 bulletin: museum of comparative zoology 



Xerus Hemprich and Ehrenberg. Ground Squirrels 

Xerus Hemprich and Ehrenberg, Symbolse Physicse, Mamra., dec. 1, fol. ee, 
pi. 9, Aug. 1832. Subgenus of Sciurus, type Sciurus (Xerus) brachyotus 
Hemprich and Ehrenberg = »SCT'Mr us rutilus Cretzschmar. 

Spermosciurus Lesson, Nouv. Tabl. du Regne Anim., Mamm., p. 110, 1842. 
Subgenus of Sciurus, type Sciurus rutilus Cretzschmar. 

Xebus rutilus rutilus (Cretzschmar) 

Sduriis abessinicus Gmelin, Linne's Syst. Nat., ed. 13, 1: pt. 1, p. 149, 1788. 

Abyssinia. Based on the "ecureuil de 1' Abyssinie" of Thevenot, Voyage, 

5: 34, 1665. Identity uncertain. 
Sciurus rutilus Cretzschmar, in Rlippell's Atlas zu d. Reise im Nordhche 

Afrika, Saugeth., p. 59, pi. 24, 1826. Eastern slope of Abyssinia. 
Sciurus (Xerus) brachyotus Hemprich and Ehrenberg, Symbolae Physicse, 

Mamm., dec. 1, fol. ee, pi. 9, Aug. 1832. Gedam near Arkiko and Taranta 

near Eilet, Abyssinia. 
Sciurus (Xerus) fusc^ls Huet, Nouv. Arch. Mus. d'Hist. Nat. Paris, (2) 3: 

139, pi. 6, f. 1, 1880. Mountains of Adel, Abyssinia. 

Xerus rutilus dabagala Heuglin 

Xeru^ dabagala Heuglin, Nova Acta Acad. Cses. Leop.- Carol., Halle, 28: art. 10, 
p. 4, pi. 2, f. 3, 1861. No locality; probably Red Sea coast. 

Xerus rutilus dorsalis Dollman 

Xerus dabagala dorsalis Dollman, Ann. Mag. Nat. Hist. (8) 7: 519, May 1911. 
Baringo, Kenya Colony, 4000 ft. 

Xerus rutilus intensus Thomas 

Xerus rutilus intensus Thomas, Ann. Mag. Nat. Hist. (7) 14: 100, Aug. 1904. 
Gerlogobi Wells, British Somaliland. 
Range: Red sandy country of Central SomaU. 

Xerus rutilus rupifrons Dollman 

Xerus dabagala rufifrons Dollman, Ann. Mag. Nat. Hist. (8) 7: 518, May 1911. 
Northern Guaso Nyiro, Kenya Colony. 

Sciurus (Xerus) flavus A. Milne-Edwards, Revue de Zool. Pure et Appliqu^e, 
(2) 19: 229, July 1867; Huet, Nouv. Arch. Mus. d'Hist. Nat. Paris, (2) 
3: pi. 6, f. 2 (type, in color), 1880. "Gaboon," but shown to be probably 
Gnelidi or Ras Hafoun, Somaliland, by Oustalet, E., Mem. Soc. Zool. de 
France, Paris, 7: 76-77, 1894. Possibly a synonym but not Sciurus flavus 
Linnaeus, 1766. 

Xerus rutilus stephanicus Thomas 

Xerus rutilus stephanicus Thomas, Ann. Mag. Nat. Hist. (7) 18: 301, Oct. 
1906. Lake Stephanie, Abyssinia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 307 

Xerus rtjtilus saturatus Neumann 

Xerus saturatus Neumann, Zool. Jahrb., Syst., 13: 546, 10 Oct. 1900. Kibwezi, 
southern Ukamba, Kenya Colony. 



Family MUSCARDINIDiE. Dormice 

Aethoglis G. M. Allen. Large Gray Dormice 

Aethoglis G. M. Allen, Joiu-n. Mammalogy, 17: 292, 14 Aug. 1936. Genotype 

Graphiurus nagtglasii Jentink. 

Aethoglis hueti hueti (Rochebrune) 

Graphiurus hueti Rochebrune, Actes Soc. Linn, de Bordeaux, (4) 7: 110, 
pi. 6, f. 1, 1883. Environs of St. Louis, Sorres, Senegal. 

Aethoglis hueti argenteus G. M. Allen 

Aethoglis hueti argenteus G. M. Allen, Journ. Mammalogy, 17: 293, 14 Aug, 
1936. Lolodorf, Cameroons. 

Aethoglis hueti monardi (St. Leger) 

Claviglis monardi St. Leger, Ann. Mag. Nat. Hist. (10) 17: 465, Apl. 1936. 
Tyihumbwe River, 15 km. above Dala, Angola. 

Aethoglis hueti nagtglasii (Jentink) 

Graphiurus nagtglasii Jentink, Notes Leyden Mus., 10: 38, 39, Apl. 1888. 
Liberia, DuQueah and Farmington Rivers. 

Claviglis Jentink. Tree JDormice 

Claviglis Jentink, Notes Leyden Mus., 10: 41, Apl. 1888. Genotype Claviglis 
crassicaudatus Jentink. Characters keyed, Thomas and Hinton, Proc. 
Zool. Soc. London, p. 232, 3 Apl. 1925. 

Claviglis alticola Roberts 

Claviglis alticola Roberts, Ann. Transvaal Mus., 13: 98, 6 July 1929. "Kastrol 
Nek," Wakkerstroom, Transvaal, 6500 ft. 

Claviglis ansorgei ansorgei (Dollman) 

Graphiurus ansorgei Dollman, Ann. Mag. Nat. Hist. (8) 9: 317, Mar. 1912. 
Tala Kilau, Dongwenna, Mossamedes, southern Angola. 

Claviglis ansorgei cuanzensis Hill and Carter 

Claviglis ansorgei cuanzensis Hill and Carter, Amer. Mus. Novitates, no. 913, 
p. 9, f. 8, 8 Mar. 1937. Chitau, Angola, 4930 ft. 



308 bulletin: museum of comparative zoology 

Claviglis brockmani brockmani (Dollman) 

Graphiurus brockmani Dollman, Ann. Mag. Nat. Hist. (8) 5: 287, Mar. 1910. 
Burao, Somaliland, 4000 ft. 

Claviglis brockmani internus (Dollman) 

Graphiurus brockmani internus Dollman, Ann. Mag. Nat. Hist. (8) 9: 318, 

Mar. 1912. Nyama Nyango, Northern Guaso Nyiro, Kenya Colony, 

3200 ft. 

Claviglis butleri (Dollman) 

Graphiurus butleri Dollman, Ann. Mag. Nat. Hist. (8) 9: 319, Mar. 1912. 
Jebel Ahmed Aga, Anglo-Egyptian Sudan. 

Claviglis christyi (Dollman) 

Graphiurus christyi Dollman, Revue Zool. Africaine, 4: 80, 25 July 1914. 
Mambaka and Mambo, Congo Beige. 

Claviglis coupeii (E. Geoffroy and F. Cuvier) 

[Myoxus] coupeii E. Geoffroy and F. Cuvier, Hist. Nat. des Mammiferes, pt. 
37, col. pi. and leaf of text ( = pl. 251), Nov. 1822. Senegal. 

Claviglis crassicaudatus crassicaudatus Jentink 

Claviglis crassicaudatus Jentink, Notes Leyden Mus., 10: 41, Apl. 1888. Du 
Queah River, western Liberia. 

Claviglis crassicaudatus Dorothea (Dollman) 

Graphiurus crassicaudatus dorolhese Dollman, Ann. Mag. Nat. Hist. (8) 9: 
312, Mar. 1912. Oban district, southern Nigeria. 

Claviglis foxi (Dollman) 

Graphiurus foxi Dollman, Ann. Mag. Nat. Hist. (8) 13: 196, Jan. 1914. Kabwir, 
Bauchi Province, northern Nigeria. 

Claviglis griselda (Schwann) 

Graphiurus griselda Schwann, Proc. Zool. Soc. London, 1: 105, 7 June 1906. 
Kuruman, Bechuanaland. 

Claviglis h^dulus (Dollman) 

Graphiurus hsedulus Dollman, Ann. Mag. Nat. Hist. (8) 9: 316, Mar. 1912; 

Assobam, Bumba River, Cameroons. 
Graphiurus schwabi G. M. Allen, Bull. Mus. Comp. Zool., 54: 441, Apl. 1912. 

Kribi, Cameroons. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 309 

Claviglis kelleni (Reuvens) 

Eliomys kelleni Reuvens, Die Myoxidse oder Schlafer, p. 35, pi. 1, f. 1; pi. 3, 
f. 3, a, b, 1890. Damaraland, Southwest Africa. 

Claviglis littoralis Roberts 

Claviglis littoralis Roberts, Ann. Transvaal Mus., 13: 97, 6 July 1929. Masiene, 
north of mouth of Limpopo River, Mozambique. 

Claviglis lorraineus (Dollman) 

Graphiurus lorraineus Dollman, Ann. Mag. Nat. Hist. (8) 5: 285, Mar. 1910. 
Molegbwe, south of Setema Rapids, Uele River, Congo Beige. 

Claviglis murinus murinus (Desmarest) 

Myoxus murinus Desmarest, Encyclop. Methodique, Mammalogie, Suppl., 

p. 542, 1822. Cape of Good Hope. 
Myoxus lalandianus Schinz, Cuvier's Thierreich, 4: 393, 1825. Cape of Good 

Hope. 
Myoxus erythrobronchus A. Smith, Zool. Journ., 4: 438, May 1829. South 

Africa. 
Myoxus cineraceus Riippell, Mus. Senckenberg., Frankfurt a. M., 3: pt. 2, 

p. 136, 1842. Port Natal. 
Myoxus cinerascens Schinz, Syst. Verzeichn., Synopsis Mamm., 2: 80, 1845 

(emendation of M. cineraceus Riippell). 
Eliomys microtis Noack, Zool. Jahrb., 2: 248, pi. 9, f. 19-22, 7 May 1887. 

"Qua Mpala, Marungu," Tanganyika Territory. 

Claviglis murinus griseus (G. M. Allen) 

Graphiurus microtis griseus G. M. Allen, Bull. Mus. Comp. Zool., 54: 440. 

Apl. 1912. Northern Guaso Nyiro, Kenya Colony. 
Graphiurus murinus johnstoni Heller, Smithsonian Misc. Coll., 59: no. 16, p. 2, 

5 July 1912. Mt. Gargues, Mathews Range, Kenya Colony. 

Claviglis murinus isolatus (Heller) 

Graphiurus murinus isolatus Heller, Smithsonian Misc. Coll., 59: no. 16, p. 3, 5 
July 1912. Mt. Umengo, Taita Mts., Kenya Colony, 5000 ft. 

Claviglis murinus Pretoria (Roberts) 

Graphiurus pretorisp Roberts, Ann. Transvaal Mus., 4: 79, 20 Oct. 1913. Little 
Wonderboom, Pretoria, Transvaal. 

Claviglis murinus raptor (Dollman) 

Graphiurus raptor Dollman, Ann. Mag. Nat. Hist. (8) 5: 96, Jan. 1910. West 
slope of Mt. Kenya, Kenya Colony. 



310 bulletin: museum of comparative zoology 

Claviglis murinus saturatus (Dollman) 

Graphiurus microtis saturatus Dollman, Ann. Mag. Nat. Hist. (8) 5: 204, Feb. 

1910. South face of Mt. Elgon, Kenya Colony, 9000 ft. 

Graphiurus murinus saturatus Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 
153, 1919. 

Claviglis murinus selindensis Roberts 

Claviglis murinus selindensis Roberts, Ann. Transvaal Mus., 19: 100, 8 Nov. 
1937. Mt. Selinda, Southern Rhodesia. 

Claviglis murinus tzaneenensis (Roberts) 

Graphiurus murinus tzaneenensis Roberts, Ann. Transvaal Mus., 4: 79, 20 Oct. 
1913. Tzaneen Estate, Transvaal. 

Claviglis nanus (De Winton) 

Myoxus (Eliomys) nanus De Winton, Proc. Zool. Soc. London, for 1896, p. 799, 
Apl. 1897. Mazoe, Mashonaland, Southern Rhodesia. 

Claviglis olga Thomas 

Claviglis olga Thomas, Ann. Mag. Nat. Hist. (9) 16: 191, July 1925. Tehsiderak, 
Asben, (Air), French West Africa. 

Claviglis orobinus (Wagner) 

Myoxus orobinus Wagner, Arch. f. Naturgesch., 11: sect. 1, p. 149, 1845. 
Sennaar. 

Claviglis parvulus (Monard) 

Graphiurus parvulus Monard, Bull. Soc. Neuchateloise des Sci. Nat., Neuchfitel, 
57: 54, 1932. Rio Mbale, Tumbole, and Vila da Ponte, Angola. 

"Claviglis parvus parvus (True) 

Eliomys parvus True, Proc. U. S. Nat. Mus., 16: 601, 25 Oct. 1893. Tana River, 
between coast and Hameye, Kenya Colony. 

Claviglis parvus dollman: (Osgood) 

Graphiurus parvus dollmani Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: no. 3, p. 15, 7 Apl. 1910. Lukenya Mt., Ulukenya Hills, Kenya Colony, 

Claviglis personatus (Heller) 

Graphiurus personatus Heller, Smithsonian Misc. Coll., 56: no. 17, p. 2, 28 Feb. 

1911. Rhino Camp, Lado Enclave. 

Claviglis smithii (Thomas) 

Myoxus (Eliomys) smithii Thomas, Ann. Mag. Nat. Hist. (6) 12: 267, Oct. 1893. 
Nassa, Speke Gulf, Lake Victoria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 311 

Claviglis soleatus soleatus (Thomas and Wroughton) 
Graphiurus soleatus Thomas and Wroughton, Trans. Zool. Soc. London, 19: 
499, Mar. 1910. Mubuku Valley, Ruwenzori East, Uganda, 5-6000 ft. 

Claviglis soleatus collaris G. M. Allen and A. Loveridge 

Claviglis soleatus collaris G. M. Allen and Loveridge, Bull. Mus. Comp. Zool., 

75: 122, Feb. 1933. Madehani, Ukinga Mts., north end of Lake Nyasa, 

Tanganyika Territory. 

Claviglis spurrelli (Dollman) 

Graphiurus spurrelli Dollman, Ann. Mag. Nat. Hist. (8) 9: 315, Mar. 1912. 
Bibianaha, 60 miles west of Kumasi, Gold Coast. 

Claviglis streeteri (Roberts) 

Graphiurus streeteri Roberts, Ann. Transvaal Mus., 4: 80, 20 Oct. 1913. Hector 
Spruit, Transvaal. 

Claviglis surdus (Dollman) 

Graphiurus surdus Dollman, Ann. Mag. Nat. Hist. (8) 9: 314, Mar. 1912. Ben- 
ito River, French Congo. 

Claviglis tasmani Roberts 

Claviglis tasmani Roberts, Ann. Transvaal Mus., 13: 95, 6 July 1929. Driefon- 
tein Mission Station, near Gwelo, Southern Rhodesia. 

Claviglis vandami Roberts 

Claviglis vandami Roberts, Ann. Transvaal Mus., 13: 97, 6 July 1929. Guja, 
lower Olifants River Valley, Portuguese Southeast Africa (Mozambique). 

Claviglis vulcanicus (Lonnberg and Gyldenstolpe) 

Graphiurus vulcanicus Lonnberg and Gyldenstolpe, Arkiv f . Zool., 17B: no. 9, 
p. 2, 14 May 1925. Mt. Karissimbi, Bii'unga Volcanoes, Congo Beige. 

Claviglis woosnami (Dollman) 

Graphiurus woosnami Dollman, Arm. Mag. Nat. Hist. (8) 6: 393, Oct. 1910. 
North of Okwa, Kalahari Desert. 

Claviglis ztjluensis Roberts 

Claviglis zuluensis Roberts, Ann. Transvaal Mus., 14: 229, 28 July 1931i 
Ubombo Bush, northern Zululand, South Africa. 



312 bulletin: museum of comparative zoology 



Eliomys Wagner 

Eliomys Wagner, Abh. K. Bayer. Akad. Wiss., Miinchen, math.-phys. CI., 3: 
176, pi. 2, f. 1-4; pi. 3, f. 1, 1843. Genotype Eliomys (Myoxus) melanurus 
Wagner. 

Bifa Lataste, Le Naturaliste, 7: 61, 15 Apl. 1885. Genotype Bifa lerotina 
Lataste. 



Eliomys munbyanus munbyanus (Pomel) 

Myoxus munbyanus Pomel, Compt. Rend. Acad. Sei. Paris, 42: 653, 1856. 

Province of Oran, Algeria. 
Eliomys lerotinus tunetae Thomas, Ann. Mag. Nat. Hist. (7) 11: 495, May 1903. 

Karouana, Tunis. See Cabrera, A., Trab. Mus. Nac. de Cien. Nat. Madrid, 

zool. ser., no. 57, p. 226, 30 Dec. 1932. 

Eliomys munbyanus cyrenaicus Festa 

Eliomys cyrenaicus Festa, Boll. Mus. Zool. ed Anat. Comp. Univ. Torino, 36: 
no. 740, p. 4, 15 Dec. 1921. Gheminez, Cirenaica (Barke). 

Eliomys munbyanus lerotina (Lataste) 

Bifa lerotina Lataste, Le Naturaliste, 7: 61, 15 Apl. 1885. Near Ghardaia, 
Mzab, Algeria. 

Eliomys munbyanus occidentalis Thomas 

Eliomys lerotinus occidentalis Thomas, Novitates Zool., 10: 300, 25 Aug. 1903. 
Rio de Oro. 
A race of E. munbyanus according to Cabrera (Trabajos del Mus. Nac. Cien. 
Nat., Madrid, zool. ser., no. 57, p. 226, 30 Dec. 1932). 



Indeterminate 

Myoxus cattoirii J. B. Fischer, Synopsis Mamm., p. 310, 1829. Senegal (based 
on Autre loir of F. Cuvier, Diet, des Sci. Nat., 27: 124, 1823, collected by 
Cattoir) . 

Gliriscus Thomas and Hinton. Rock Dormice 

Gliriscus Thomas and Hinton, Proc. Zool. Soc. London, p. 232, 3 Apl. 1925. 
Genotype Graphiurus platyops Thomas. 

Gliriscus angolensis angolensis (De Winton) 

Graphiurus angolensis De Winton, Ann. Mag. Nat. Hist. (7) 20: 320, Sep. 1897. 
Caconda, Angola. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 313 

Gliriscus angolensis jordani Roberts 

Gliriscus angolensis jordani Roberts, Ann. Transvaal Mus., 13: 95, 6 July 1929. 
Isoka, Northern Rhodesia. 



Gliriscus platyops (Thomas) 

Graphiurus platyops Thomas, Ann. Mag. Nat. Hist. (6) 19: 388, Apl. 1897. 
Enkeldorn, Mashonaland, Southern Rhodesia. 

Gliriscus rupicola rupicola Thomas and Hinton 

Gliriscus rupicola Thomas and Hinton, Proc. Zool. Soc. London, p. 232, 3 Apl. 
1925. Karibib, inland highlands of Southwest Africa, 3842 ft. 

Gliriscus rupicola montosus Thomas and Hinton 

Gliriscus rupicola montosus Thomas and Hinton, Proc. Zool. Soc. London, p. 

233, 3 Apl. 1925. Great Brukaros Mountain, Southwest Africa, 3600- 

5000 ft. 



Graphiurus Smuts. Cape Dormice 

Graphiurus Smuts, Enumer. Mamm. Cap., p. 32, 1832. Genotype Graphiurus 
capensis Smuts = Sciuru^ ocularis A. Smith. 



Graphiurus eastwood^ Roberts 

Graphiurus eastwoodse Roberts, Ann. Transvaal Mus., 4: 80, 20 Oct. 1913. 
Woodbush, Transvaal. Probably a race of (?. ocularis. 

Graphiurus ocularis ocularis (A. Smith) 

Sciurus ocularis A. Smith, Zool. Journ., 4: 439, May 1829. Near Plettenberg's 

Bay, Cape of Good Hope. 
Graphiurus typicus A. Smith, South African Quart. Journ., 2: 145, 1834. Forests 

about Knysna, South Africa. 
"Graphiure" [Myoxus] capensis F. Cuvier, Hist. Nat. des Mamm., pt. 60, and 

text, pi. 254, Sep. 1829. 
Graphiurus elegans Ogilby, Proc. Zool. Soc. London, p. 5, July 1838. ? Cape of 

Good Hope (see Thomas, O., Proc. Zool. Soc. London, p. 298, 29 Apl. 

1926). 



314 bulletin: museum of compael\tive zoology 

Family CRICETIDiE. Hamster-like Rodents 
Subfamily CRICETIN^ 
See Miller and Gidley: Journ. Washington Acad. Sci., 8:436, 19 July 1918. 



Mystromys Wagner 

Mystromys Wagner, Gelehrte Anzeigen, Mtinchen, 12: no. 54, col. 434, 17 Mar. 
1841. Genotype Mystromys albipes Wagner = Oiomys albicaudatus A. 
Smith. 

Mystromys ALBicAtrDATus albicaudatus (A. Smith) 

Otomys albicaudatus A. Smith, South African Quart. Journ., 2: 148, 1834; 

lUustr. Zool. South Africa, Mamm., pi. 33 and text, Sep. 1841. District 

of Albany, South Africa. 
Arvicola albicaudatus Desmarest, Encyclop. MethodiqueJ Mammalogie, pt. 2, 

p. 281, 1822. Locality unknown; based on a specimen given the name 

Lemmus albicaudatus by Geoffroy in Catalogue of the Museum d'Hist. 

Nat. Paris. Although referred to Malacothrix by Trouessart, this may not 

be the same as Otomys albicaudatus A. Smith. 
Mystromys albipes Wagner, Gelehrte Anzeigen, Miinchen, 12: no. 54, col. 435, 

17 Mar. 1841; Arch. f. Naturgesch., 7: pt. 1, p. 133, 1841. South Africa. 
Mystromys lanuginosus Lichtenstein, Verzeichn. d. Doubletten Mus. Berlin, 

p. 10, 1842. 

Mystromys albicaudatus fumosus Thomas and Schwann 
Mystromys albicaudatus fumosus Thomas and Schwann, Proc. Zool. Soc. Lon- 
don, 1: 137, 1 June 1905. Wakkerstroom, Transvaal. 

Mystromys longicaudatus Noack 

Mystromys longicaudatus Noack, Zool. Jahrb., 2: 246, pi. 9, f. 16-18, 7 May 
1887. Gonda, Tanganyika Territory? 



Subfamily LOPHIOMYIN^. Maned Rats 
See Miller and Gidley: Journ. Washington Acad. Sci., 8: 437, 19 July 1918. 

LoPHiOMYS Milne-Edwards. Maned Rat 

Lophiomys Milne-Edwards, L'Institut, Paris, 35: 46, 6 Feb. 1867; Compt. 
Rend. Acad. Sci. Paris, 64: 812, 1867. Genotype Lophiomys imhausii 
Milne-Edwards. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 315 

Phradomys Peters, Zeitschr. f. Ges. Naturw., Berlin, 29: 195, 1867; Sitzb. 
Ges. Naturf. Freunde, Berlin, p. 19, 1867. Genotype Phradomys sethiopicus 
7eters=Lophiomys i7nhausii Milne-Edwards. Phragmomys in second ref. 

See Thomas, O. The species of Lophiomys. Ann. Mag. Nat. Hist. (8) 6: 
222-223, Aug. 1910, with key. 

Lophiomys bozasi Oustalet 

Lophiomys bozasi Oustalet, Bull. Mus. d'Hist. Nat., Paris, 8: 400, 1902. Goba, 
country of the Arussi Gallas, southern Abyssinia, 3000 m. 
Synonymous with L. sethiopicus according to Thomas, Ann. Mag. Nat. Hist. 
(8) 6: 223, Aug. 1910. 

Lophiomys imhausii Milne-Edwards 

Lophiomys imhausii Milne-Edwards, L'Institut, Paris, 35: 46, 6 Feb. 1867; 

Compt. Rend. Acad. Sci., Paris, 64: 812, 1867. Probably from African 

coast opposite Aden, where it was purchased. 
Phradomys sethiopicus Peters, Zeitschr. f. Ges. Naturw., Berlin, 29: 195, 1867; 

Sitzb. Ges. Naturf. Freunde, Berlin, p. 19, 1867. Maman, north of Kassala, 

Anglo- Egyptian Sudan. 

Lophiomys ibeanus ibeanus Thomas 

Lophiomys ibeanus Thomas, Ann. Mag. Nat. Hist. (8) 6: 223, Aug. 1910. 

Mile 513 on Uganda Railway, between Londiani and Lumbwa Stations, 

Mau region, Kenya Colony. 

Lophiomys ibeanus hindei Thomas 

Lophiomys ibeanus hindei Thomas, Ann. Mag. Nat. Hist. (8) 6: 223, Aug. 1910. 
Mutaragwa, Aberdare Mts., Kenya Colony. 
Probably a synonym of L. ibeanus. All the recognizable forms are doubtless 
races of L. imhausii. 

Lophiomys smithi Rhoads 

Lophiomys smithi Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 524, text-f., 

pi. 25, 8 Dec. 1896. Sheikh Husein, western Somaliland, about 8° N., 

41° E. 

Lophiomys testudo Thomas 

Lophiomys testudo Thomas, Ann. Mag. Nat. Hist. (7) 15: 80, Jan. 1905. 
Ravine Station, Kenya Colony. 

Lophiomys thomasi Heller 

Lophiomys thomasi Heller, Smithsonian Misc. Coll., 59: no. 16, p. 4, 5 July 
1912. Mt. Gargues, Mathews Range, Kenya Colony, 6000 ft. 



316 bulletin: museum of comparative zoology 

Subfamily MICROTIN^. Voles 
MiCROTUS Schrank. Meadow Mice or Voles 

Microtus Schrank, Fauna Boica, 1: 72, 1798. Genotype Microtus terrestris 
Schrank = Mus arvalis Pallas. 

Microtus mustersi Hinton 

Microtus mustersi Hinton, Ann. Mag. Nat. Hist. (9) 18: 305, Sep. 1926. Merg, 
Cyrenaica, Barka. 

Note: The record of Arvicola arvalis from Abyssinia, by Martoglio (Ann. 
d'Igiene Sperim., Torino, (2) 23: 161-170, 1913) is doubtless based on a mis- 
identification of a species of Lophuromys or of Arvicanthis as suggested by 
Laurent, P., Bull. Soc. d'Hist. Nat. de I'Afrique du Nord, 27: 384-386, Nov. 
1936. 

Subfamily NESOMYIN^E. Madagascan Cricetines 
Brachytarsomys Giinther 

Brachytarsomys Giinther, Proc. Zool. Soc. London, p. 79, text-f. 3 (cranium), 
pi. 16, June 1875. Genotype, by monotypy, Brachytarsomys albicauda 
Giinther. 

Brachytarsomys albicauda Giinther 

Brachytarsomys albicauda Giinther, Proc. Zool. Soc. London, p. 80, text-f.3, 
pi. 16, June 1875. Madagascar, between Tamatave and Murundava. 



Brachyuromys Forsyth Major 

Brachyuromys Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 322, Oct. 1896. 
Genotype Brachyuromys ramirohitra Forsyth Major. 

Brachyuromys betsileonensis (Bartlett) 

Nesomys betsileonensis Bartlett, Proc. Zool. Soc. London, for 1879, p. 770, 
Apl. 1880. Southeastern Betsileo, Madagascar. 

Brachyuromys ramirohitra Forsyth Major 

Brachyuromys ramirohitra Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 
323, Oct. 1896. Ampitambe forest, Betsimisaraka country (border of 
northeastern Betsileo), six hours southeast of Fandriana, Madagascar. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 317 



Eliurus Milne-Edwards 

Eliurus Milne-Edwards, Ann. des Sci. Nat., zool., (6) 20: art. 1 bis, p. 1, 1885. 
Genotype Eliurus myoxinus A. Milne-Edwards. 

Eliurus majori Thomas 

Eliurus majori Thomas, Ann. Mag. Nat. Hist. (6) 16: 164, Aug. 1895. Ambo- 
limitombo forest, central Madagascar. 

Eliurus minor Forsyth Major 

Eliurus minor Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 462, Dec. 1896. 
Ampitambe forest, northeastern Betsileo, Madagascar. 

Eliurus myoxinus Milne-Edwards 

Eliurus myoxinus Milne-Edwards, Ann. des Sci. Nat., zool., (6) 20: art. 1 bis, 
p. 1, 1885. West coast of Madagascar. 

Eliurus tanala Forsyth Major 

Eliurus tanala Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 462, Dec. 1896. 

Forest of the Tanala of Ikongo, near Vinanitelo, 30 miles south of Fiana- 

rantsoa, Madagascar. 

Gymnuromys Forsyth Major 

Gymnuromys Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 324, Oct. 1896; 
Genotype Gymnuromys roberti Forsyth Major. 

Gymnuromys roberti Forsyth Major 

Gymnuromys roberti Forsyth Major, Ann. Mag. Nat. Hist. (6) 18: 324, Oct. 
1896. Ampitambe forest, northeastern Betsileo, Madagascar. 



Hallomys Jentink 

Hallomys Jentink, Notes Leyden Mus., 1: 107, Apl. 1879. Genotype Hallomys 
audeberti Jentink. 

Hallomys audeberti Jentink 

Hallomys audeberti Jentink, Notes Leyden Mus., 1: 107, Apl. 1879. Maisine 
and Savary, northeastern Madagascar. 



Hypogeomys a. Grandidier 

Hypogeomys A. Grandidier, Revue et Mag. de Zool., (2) 21: 338, Sep. 1869. 
Genotype Hypogeomys antimena A. Grandidier. 



318 bulletin: museum of comparative zoology 

Hypogeomys antimena a. Grandidier 

Hypogeomys antimena A. Grandidier, Revue et Mag. de Zool. (2) 21: 338, Sep.. 

1869. Banks of the Tsidsibon and the Andranoumene, Menab6, western 

Madagascar. 

Macrotarsomys Milne-Edwards and G. Grandidier 

Macrotarsomys Milne-Edwards andG. Grandidier, Bull. Mus. d'Hist. Nat. Paris,. 
4: 179, 1898. Genotype Macrotarsomys bastardi Milne-Edwards and G. 
Grandidier. 

Macrotarsomys bastardi Milne-Edwards and G. Grandidier 
Macrotarsomys bastardi Milne-Edwards and G. Grandidier, Bull. Mus. d'Hist. 
Nat. Paris, 4: 179, 180, text-f. 1, 2, 1898. South of Mangoky, southwestern 
Madagascar. 
For Mr. E. Bastard. 

Nesomys Peters 

Nesomys Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 54, 1870. Genotype 
Nesomys rufus Peters. 

Nesomys rufus Peters 

Nesomys rufus Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 55, 1870. Vohima,. 
Madagascar. 



Subfamily GERBILLIN^. Gerbils 
Ammodillus Thomas 

Ammodillus Thomas, Ann. Mag. Nat. Hist. (7) 14: 102, text-f. a, b, Aug. 1904. 
Genotype Gerbillus imbellis De Winton. 

Ammodillus imbellis (De Winton) 

Gerbillus imbellis De Winton, Ann. Mag. Nat. Hist. (7) 1: 249, Mar. 1898. 
Goodar, Somaliland. 

Desmodilliscus Wettstein 

Desmodilliscus Wettstein, Anzeiger K. Akad. Wiss. Wien, math.-nat. Kl., 53: 
153, 1916. Genotype Desmodilliscus braueri Wettstein. 

Desmodilliscus braueri Wettstein 

Desmodilliscus braueri Wettstein, Anzeiger K. Akad. Wiss. Wien, math.-nat. 

KL, 53: 153, 1916. Between Um Ramad and Nubbaka, south of El Obeid, 

Kordof an, Anglo-Egyptian Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 319 

Desmodilliscus BtrcHANANi Thomas and Hinton 

Desmodilliscus buchanani Thomas and Hinton, Novitates Zool., 27: 317, 15 
June 1920. Farniso, near Kano, northern Nigeria. 

Desmodillus Thomas and Schwann 

Desmodillus Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 2, p. 6, 
9 Feb. 1904; Proc. Zool. Soc. London, 1: 177, June 1904. Genotype Ger- 
billas auricularis A. Smith. 

Desmodillus auricularis auricularis (A. Smith) 

Gerbillus auricularis A. Smith, South African Quart. Journ., 2: 160, 1834. 

Mountains of Little Namaqualand, South Africa. 
Gerbillus brevicaudatus F. Cuvier, Trans. Zool. Soc. London, 2: 144, pi. 26, f. 

10-13, 4 May 1838. Cape of Good Hope. 

Desmodillus auricularis pudicus Dollman 

Desmodillus auricularis pudicus Dollman, Ann. Mag. Nat. Hist. (8) 6: 395, 
Oct. 1910. Lehutitung, Kalahari, Bechuanaland. 

Dipodillus Lataste 

Dipodillus Lataste, Le Naturaliste, 3: no. 64, p. 506, 15 Nov. 1881; 4: 12, 15 
Jan. 1882. Subgenus of Gerbillus, including Gerbillus simoni Lataste, and 
G. campestris LevaiUant. 

Dipodillus amcenus De Winton 

Dipodillus amaenus De Winton, Ann. Mag. Nat. Hist. (7) 9: 46, Jan. 1902. 
Giza Province, Egypt. 

Dipodillus bottai (Lataste) 

Gerbillus bottai Lataste, Le Naturaliste, 4: no. 5, p. 36, 1 Mar. 1882. Sennaar, 
Anglo-Egyptain Sudan. 

Dipodillus brockmani Thomas 

Dipodillus brockmani Thomas, Ann. Mag. Nat. Hist. (8) 5: 420, May 1910. 
Burao, 85 miles south of Berbera, British Somaliland. 

Dipodillus campestris campestris (Levaillant) 

Gerbillus campestris Levaillant, Exploration Sci. de I'Algerie, Zool., Mammi- 

feres. Atlas, pi. 5, f. 2 (col.), 2a, 1857; Loche, Cat. Mamm. et Oiseaux 

observes en Algerie, p. 22, 1858. Province of Constantine, Algeria. 
Gerbillus gerbii Loche, Cat. Mamm. et Oiseaux observes en Algerie, p. 23, 1858; 

Exploration Sci. de TAlgerie, Zool., Mammiferes, p. 107, 1867. Country 

of the Beni Shman, Algeria. 



320 bulletin: museum of comparative zoology 

Gerbillus minutus Loche, Cat. Mamm. et Oiseaux observes en Alg^rie, p. 23, 
1858. Douilba, Sahara, Algeria. 

DipODiLLtJS CAMPESTRis ciNNAMOMEus Cabrera 

Dipodillus campestris cinnarnomeus Cabrera, Bol.Real Soc. Espaiiola Hist. Nat., 
Madrid, 16: 385, Oct. 1916. Taguidert, south of Mogador, Morocco. 

Dipodillus campestris riparius Cabrera 

Dipodillus campestris riparius Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., 

Madrid, 22: 112, 27 Feb. 1922. Valley of the Uad Martin, Yebala, 

Morocco. 

Dipodillus campestris rozsik^ Thomas 

Dipodillus campestris roszikse (sic) Thomas, Ann. Mag. Nat. Hist. (8) 2:374, 

Oct. 1908. Biskra, Algeria. 
Dipodillus campestris rozsikse Thomas, Novitates Zool., 20: 589, 21 Oct. 1913 

(correction of typographical error). 

Dipodillus diminutus Dollman 

Dipodillus dimimdus Dollman, Ann. Mag. Nat. Hist. (8) 7: 520, May 1911. 
Nyama Nyango, Northern Guaso Nyiro, Kenya Colony. 

Dipodillus dodsoni dodsoni Thomas 

Dipodillus dodsoni Thomas, Proc. Zool. Soc. London, 2: 7, Oct. 1902. Ain 
Hammam, Tripoli. 

Dipodillus dodsoni patrizii de Beaux 

Dipodillus dodsoni patrizii de Beaux, Ann. Mus. Civ. Stor. Nat., Genova, 55: 
379, 2 June 1932. Oasis of Cufra, Libyan Desert, Libya. 

Dipodillus grobbeni (Klaptocz) 

Gerbillus (Dipodillus) grobbeni Klaptocz, Zool. Jahrbiicher, Syst., 27: 252, 2 

text-f., (1908) 25 Feb. 1909. Dernah, north coast of Barka, Cyrenaica. 

Status uncertain according to H. Heim de Balsac. 

Dipodillus harwoodi harwoodi (Thomas) 

Gerbillus {Dipodillus) harwoodi Thomas, Ann. Mag. Nat. Hist. (7) 8: 275, foot- 
note, Oct. 1901. Lake Naivasha, Kenya Colony. 

Dipodillus harwoodi luteus Dollman 

Dipodillus luteus Dollman, Ann. Mag. Nat. Hist. (8) 14: 489, Dec. 1914. 

Southern Guaso Nyiro, Nyanza Province, Kenya Colony. 
Dipodillus harwoodi luteus Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 27, 

1919. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 321 

DiPODiLLUS HENLEYi De Winton 

Dipodillus henleyi De Winton, Novitates Zool., 10: 284, pi. 8, f. 1, 25 Aug. 1903. 
Zaghig, Wadi Natron, Lower Egypt. 

Dipodillus hilda Thomas 

Dipodillus hilda Thomas, Ann. Mag. Nat. Hist. (9) 2: 62, July 1918. Seacoast 
70 miles southwest of Tangiers, Morocco. 

Dipodillus jordani Thomas 

Dipodillus jordani Thomas, Ann. Mag. Nat. Hist. (9) 2: 60, July 1918. Guelt- 
es-Stel, central plateau of Algeria, 900 m. 

Dipodillus juliani St. Leger 

Dipodillus jidiani St. Leger, Ann. Mag. Nat. Hist. (10) 15: 669, June 1935. 
Bulbar, Somaliland. 

Dipodillus lowei Thomas and Hinton 

Dipodillus lowei Thomas and Hinton, Proc. Zool. Soc. London, p. 261, 6 July 
1923. Jebel Marra, Darfur, Anglo-Egyptian Sudan, 9500 ft. 

Dipodillus mackilligini Thomas 

Dipodillus mackilligini Thomas, Ann. Mag. Nat. Hist. (7) 14: 158, Aug. 1904. 
Wadi Alagi, eastern desert of Nubia, about lat. 22° N., long. 35° E., Anglo- 
Egyptian Sudan. 

Dipodillus marine Bonhote 

Dipodillus marix Bonhote, Proc. Zool. Soc. London, for 1909, p. 792, 18 Apl. 
1910. Mokattam Hills, east of Cairo, Egypt. 

Dipodillus muriculus Thomas and Hinton 

Dipodillus muriculus Thomas and Hinton, Proc. Zool. Soc. London, p. 263, 
6 July 1923. Madu, 80 miles northeast of El Fasher, Darfur, Anglo- 
Egyptian Sudan. 

Dipodillus percivali Dollman 

Dipodillus percivali Dollman, Ann. Mag. Nat. Hist. (8) 14: 488, Dec. 1914. 
Voi, Kenya Colony. 

Dipodillus principulus Thomas and Hinton 

Dipodillus principulus Thomas and Hinton, Proc. Zool. Soc. London, p. 262f 

6 July 1923. El Malha, Jebel Maidob, northern Darfur, Anglo-Egyptian 

Sudan. 



322 bulletin: museum of comparative zoology 

DiPODiLLUs PusiLLUs (Pcters) 

Gerbillus pusillus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 201, 1878. 
Ndi and Kitui, Kenya Colony. 

DiPODiLLUS QUADRiMACULATUs (Lataste) 

Gerbillus quadriniacidatus Lataste, Le Naturaliste, 4: no. 4, p. 27, 15 Feb. 1882. 
Nubia. 

DiPODiLLUs RUBERRiMUS (Rhoads) 

Gerbillus ruberrimus Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 538, 
8 Dec. 1896. Finik, near Webi Shebeli, western British Somaliland. 

DiPODiLLUs siMONi (Latastc) 

G[erbillus\ simoni Lataste, Le Naturaliste, 3: no. 63, p. 499, 1 Nov. 1881. 
Oued Magra, between M'sila and Barika, high plateau of Algeria. 

DiPODiLLUs soMALicus Thomas 

Dipodillus somalicus Thomas, Ann. Mag. Nat. Hist. (8) 5: 197, Feb. 1910. 
Upper Sheikh, British Somaliland. 

Dipodillus stigmonyx stigmonyx (Heuglin) 

Meriones stigmonyx Heuglin, Reise in Nordost-Afrika, 2: 78, 1877. Khartoum, 
Anglo-Egyptian Sudan. 

Dipodillus stigmonyx luteolus (Thomas) 

Gerbillus stigmonyx luteolus Thomas, Ann. Mag. Nat. Hist. (7) 8; 275, Oct. 
1901. Duem, Anglo-Egyptian Sudan. 

Dipodillus venustus (Sundevall) 

Meriones venustus Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 
for 1842, p. 230, 1843. Bahr-el-Abiad, Anglo-Egyptian Sudan. 

Dipodillus vivax Thomas 

Dipodillus vivax Thomas, Proc. Zool. Soc. London, 2: 8, Oct. 1902. Sebha, 
Tripoli. 

Dipodillus watersi (De Winton) 

Gerbillus (Dipodillus) watersi De Winton, Novitates Zool,, 8: 399, pi. 20, 31 
Dec. 1901. Shendi, upper Nile, Anglo-Egyptian Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 323 

Gerbillus Desmarest 
Subgenus Gerbillus Desmarest 

Gerbillus Desmarest, Dictionnaire d'Hist. Nat., ed. 1, 24: Tabl. Methodique d. 
Mamm., p. 22, 1804; Nouv. Dictionnaire d'Hist. Nat., 13: p. 106, 1817. 
Genotype Gerbillus aegyptius Desmarest = Dipus gerbillus Olivier. 

Gerbillus acticola Thomas 

Gerbilbis acticola Thomas, Ann. Mag. Nat. Hist. (9) 2: 147, Aug. 1918. Ber- 
bera, British Somaliland. 

Gerbillus agag Thomas 

Gerbillus agag Thomas, Proc. Zool. Soc. London, 1: 296, 6 Aug. 1903. Agageh 
Wells, western Kordofan, Anglo-Egyptian Sudan. 

Gerbillus andersoni andersoni De Winton 

Gerbillus andersoni De Winton, Ann. Mag. Nat. Hist. (7) 9: 45, Jan. 1902. 
Mandara, eastern environs of Alexandria, Egypt. 
Range: Northwestern and northern Egypt. 

Gerbillus andersoni bonhotei Thomas 

Gerbillus bonhotei Thomas, Ann. Mag. Nat. Hist. (9) 3: 560, June 1919. Khabra 
Abu Guzour, northeastern Sinai. 
Range: Northeastern Sinai to "the Egyptian side of the Palestine frontier" 
(Flower, 1932). 

Gerbillus bilensis Frick 

Gerbillus bilensis Frick, Ann. Carnegie Mus., 9: 12, 6 June 1914. Dry plains 

near Bilen, Abyssinia. 
(?) Mus galanus Heuglin, Reise in Nordost-Afrika, 2: 75, 1877. Stony pastures 

in the Wolo-Gala country, Abyssinia. 

Gerbillus calidus calidus Thomas 

Gerbillus calidus Thomas, Ann. Mag. Nat. Hist. (9) 2: 63, July 1918. Molopo, 
west of Morokwen, Bechuanaland. 

Gerbillus calidus kalaharicus Roberts 

Gerbillus calidus kalaharicus Roberts, Ann. Transvaal Mus., 15: 10, 1 Oct. 1932. 
Gomodimo Pan, central Kalahari, Bechuanaland. 

Gerbillus cosensi Dollman 

Gerbillus cosensi Dollman, Abstr. Proc. Zool. Soc. London, no. 131, p. 25, 14 

Apl. 1914; Proc. Zool. Soc. London, p. 311, 16 June 1914. Kozibiri River, 

Ngamatak, Turkwel River, Uganda. 



324 bulletin: museum of comparative zoology 

Gerbillus dallonii Heim de Balsac 

Gerbillus dallonii Heim de Balsac, Mem. Acad, des Sci. de I'lnstitut de France, 

Paris, (2) 62: 43 (mem. no. 1), text-f. 1, 1936. Tibesti, Chad Military 

Territory. 

Gerbillus dongolanus (Heuglin) 

Meriones dongolanus Heuglin, Reise in Nordost-Afrika, 2: 79, 1877. Dongola, 
Anglo-Egyptian Sudan. 

Gerbillus dunni Thomas 

Gerbillus dunni Thomas, Ann. Mag. Nat. Hist. (7) 14: 101, Aug. 1904. Gerlo- 
gobi, British Somaliland. 
For Major H. N. Dunn, R. A. M. C, the collector. 

Gerbillus eatoni Thomas 

Gerbillus eatoni Thomas, Proc. Zool. Soc. London, 2: 6, Oct. 1902. Elcusher, 
Tripoli. 

Gerbillus foleyi Heim de Balsac 

Gerbillus foleyi Heim de Balsac, Suppl. au Bull. Biol, de France et de Belgique, 

Paris, 21: 317 (brief description), 389, f. 15, no. 12 (skull), 1936; Bull. Soc. 

Zool. de France, 62: 331, 14 Aug. 1937 (formal description). Beni-abb^s, 

western Algeria. 

Gerbillus gerbillus (Olivier) 

Dipus gerbillus Olivier, Bull, des Sci., Soc. Philomath., Paris, 2: no. 40, p. 121, 

1801. Egypt (Giza Province). 
Gerbillus aegypiius Desmarest, Dictionnaire d'Hist. Nat., ed. 1, 24: tabl. 
methodique d. Mamm., p. 22, 1804. 
Range: Algeria and northern Nigeria to Egypt, Nubia and Sinai (Flower). 

Gerbillus hirtipes hirtipes Lataste 

Gerbillus hirtipes Lataste, Le Naturaliste, 3: no. 64, p. 506, 15 Nov. 1881; Actes 

Soc. Linn, de Bordeaux, 40 ( = (4) 10): 377, pi. 10, 1886. Oasis of Nefta, 

in the Djerid, Sahara, Tunis. 

Gerbillus hirtipes hesperinus Cabrera 

Gerbillus hirtipes hesperinus Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., 
Madrid, 6: 365, July 1906. Mogador, Morocco. 

Gerbillus latastei Thomas and Trouessart 

Gerbillus latastei Thomas and Trouessart, Bull. Soc. Zool. de France, p. 172, 
1903. Kebili, southern Timis. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 325 

Gerbillxjs longicaudtjs (Wagner) 

Meriones longicaudus Wagner, Schreber's Saugthiere, Suppl., 3: 477, 1843. 
Egypt. 

Gerbillxjs macroptjs (Heuglin) 

Meriones macro-pus Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 31: 
no. 7, p. 9, 1864. Between Djur and Kosanga Rivers, Bongo, Sudan. 

Gerbillus nancillus Thomas and Hinton 

Gerbillus nancillus Thomas and Hinton, Proc. Zool. Soc. London, p. 260, 6 July 

1923. Plains of Darfur, 45 miles north of El Fasher, Anglo-Egyptian 

Sudan. 

Gerbillus Nigeria Thomas and Hinton 

Gerbillus nigerix Thomas and Hinton, Novitates Zool., 27: 317, 15 June 1920. 
Farniso, near Kano, northern Nigeria. 

Gerbillus nigrotibialis Monard 

Gerbillus nigrotibialis Monard, Bull. Soc. Neuchateloise d. Sci. Nat., Neu- 
chatel, 57: 54, text-f. 5, 6 (p. 65), 1932. Kubango, Angola. 

Gerbillus p^ba p^ba A. Smith 

Gerbillus pseba A. Smith, Rept. Exped. for Exploring Central Africa from Cape 

of Good Hope, June 23, 1834, under Superintendence of A. Smith, p. 43, 

1836. "Country beyond Latakoo," South Africa. 
Gerbillus tenuis A. Smith, Illustr. Zool. South Africa, Mamm., pi. 36, f. 2 and 

text, Feb. 1842. North of Latakoo. 
A synonym of Gerbillus pseba according to Thomas (Ann. Mag. Nat. Hist. 
(9) 2: 64, July 1918). 

Gerbillus p^ba broomi Thomas 

Gerbillus pseba broomi Thomas, Ann. Mag. Nat. Hist. (9) 2: 64, July 1918. 
Port NoUoth, Namaqualand, Cape of Good Hope. 

Gerbillus p^ba coombsi Roberts- 

Gerbillus pa^ba coombsi Roberts, Ann. Transvaal Mus., 13: 98, 6 July 1929. 
Zwarthoek, Zoutpansberg, Transvaal. 

Gerbillus pulvinatus Rhoads 

Gerbillus pulvinatus Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 537, S 
Dec. 1896. Rusia, Lake Rudolf, Abyssinia. 



326 bulletin: museum of comparative zoology 

Gerbillus pygargus F. Cuvier 

G[erbillus] pygargus F. Cuvier, Trans. Zool. Soc. London, 2: 142, pi. 25, f. 10-14, 
4 May 1838. Upper Egypt. 

Gerbillus pyramid tjm pyramidum I. Geoff roy. Greater Egyptian Gerbil 
Gerbillus pyramidum I. Geoffroy, Dictionnaire Classique d'Hist. Nat., 7: 321, 

Feb. 1825. Giza Province, Egypt. 
G[erbillus\ burioni F. Cuvier, Trans. Zool. Soc. London, 2: 145, pis. 22, 23, 4 

May 1838. "Dahrfur"; "the type specimen was an animal which James 

Burton took alive to Paris in 1833" (Flower). 

Gerbillus pyramidum tarabuli Thomas 

Gerbillus pyramidum tarabuli Thomas, Proc. Zool. Soc. London, 2: 5, Oct. 1902. 
Sebha, Tripoli. 

Gerbillus riggenbachi Thomas 

Gerbillus riggenbachi Thomas, Novitates Zool., 10: 301, 25 Aug. 1903. Rio de 
Oro. 

Gerbillus Rosalinda St. Leger 

Gerbillus rosalinda St. Leger, Ann. Mag. Nat. Hist. (10) 4: 295, Sep. 1929. 

Abu Zabad, 145 km. southwest of El Obeid, Kordofan, Anglo-Egyptian 

Sudan. 

Gerbillus sw alius sw alius Thomas and HLnton 

Gerbillus swalius Thomas and Hinton, Proc. Zool. Soc. London, p. 235, 3 Apl. 
1925. Karibib, Southwest Africa. 



'} 



Gerbillus swalius leucanthus Thomas 

Gerbillus swalius leucanthus Thomas, Proc. Zool. Soc. London, p. 382, 12 July 
1927. Ondongwa, northwestern Ovamboland, Southwest Africa. 

Gerbillus swalius oralis Thomas and Hinton 

Gerbillus swalius oralis Thomas and Hinton, Proc. Zool. Soc. London, p. 236, 
3 Apl. 1925. Rooibank, coast region near Walvis Bay, Southwest Africa. 

Gerbillus vallinus Thomas 

Gerbillus vallinus Thomas, Ann. Mag. Nat. Hist. (9) 2: 148, Aug. 1918. Tuin, 

near Kenhart, Hartebeest River, near 29° S. and 21° E., Bushmanland, 

Southwest Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 327 

Subgenus Endecapleura Lataste 

Endecapleura Lataste, Le Naturaliste, Paris, 4: 127, 15 Aug. 1882. Subgenus 

of Gerbillus, type Gerbillus garamantis Lataste. 
H endecapleura Lataste, Ann. Mus. Civ. Stor. Nat., Geneva, 20: 258, footnote, 

8 Mar. 1884 (emendation). 

Gerbillus garamantis Lataste 

Gerbillus garamentis Lataste, Le Naturaliste, 3: no. 64, p. 506, 507 (spelled 

garamantis), 15 Nov. 1881. Sidi-Roueld, Ouargla, Algeria. 
Dipodillus garamantis Thomas, Ann. Mag. Nat. Hist. (8) 2: 374, Oct. 1908. 

Meriones Illiger. Jirds 

Meriones Illiger, Prodromus Syst. Mamm. et Avium, p. 82, 1811. Genotype 
not specified; included Dipus tamaricinus (Pallas) and D. meridianus 
(Gmelin). 

Idomeneus Schulze, Zeitschr. f. Naturw., Stuttgart, 73: 201, 19 Dec. 1900. 
Subgenus of Meriones; type Mus tamaricinus Pallas. 

Meriones calurus (Thomas) 

Gerbillus calurus Thomas, Ann. Mag. Nat. Hist. (6) 9: 76, Jan. 1892. No 
locality. 
Range: Sinai and eastern Egypt. 

Meriones erythrourus legeri Aharoni 

Meriones erythrourus legeri Aharoni, Zeitschr. f. Saugetierkunde, 7: 202, 15' Dec. 
1932. Wadi-el-Abjad, southern Palestine. 

Meriones grandis Cabrera 

Meriones grandis Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 7: 175, 
Apl. 1907. Marrakesh, central Morocco. 

Meriones lacernatus Riippell 

Meriones lacernatus Riippell, Mus. Senckenburgianum, 3: 96, 115, pi. 6, f. 1, 
1842. Dembea Lake, Abyssinia. 

Meriones libycus libycus Lichtenstein 

Meriones libycus Lichtenstein, Verzeichn. d. Doubletten Zool. Mus. Berlin, 

p. 5, 1823. "E deserto libyco." Egypt, "no doubt Alexandria" (Thomas). 
Meriones shawi Rozet, Voy. Reg. Alger., 1: 243, 1833 (ex Shaw, Voy., 1: 321, 

1743). Algeria. 
Gerbillus shawii Duvernoy, LTnstitut, p. 400, Nov. 1841. 
Gerbillus sellysii (sic) Pomel, Compt. Rend. Acad. Sci. Paris, 42: 652, 1856. 

Algeria, Oran province. 



328 bulletin: museum of comparative zoology 

Gerbillus richardii Loche, Exploration Sci. de I'Algerie, ZooL, Mammiferes, 
p. 104, 1867. Environs of Boghar and country of the Beni Sliman, Al- 
geria. Status uncertain. 

Gerbillus savvii Loche, Exploration Sci. de I'Algerie, Zool., Mammiferes, pi. 6, 
1867 (lapsus for shawii). 

Meriones isis Thomas, Ann. Mag. Nat. Hist. (9) 3: 271, Mar. 1919. Ramleh, 
near Alexandria, Egj^pt. 

Meriones libycus albipes Lataste 

Meriones albipes Lataste, Le Naturaliste, 4: no. 13, p. 101, 1 July 1882. M'sila, 
high plateau of Algeria. 

Meriones libyctjs auziensis Lataste 

Meriones auziensis Lataste, Le Naturaliste, 4: no. 10, p. 77, 15 May 1882. 
Borders of Oued Akarit, between Aumale and Oued Okris, Algeria. 

Meriones libycus caudattts Thomas 

Meriones libycus caudatus Thomas, Ann. Mag. Nat. Hist. (9) 3: 267, Mar. 
1919. Ferdjan, Tripoli, Libya. 

Meriones libycus confalonierii de Beaux 

Merione[s] libycus confalonierii de Beaux, Ann. Mus. Civ. Stor. Nat., Genova, 

55: 384, 2 June 1932. El Agheila, Libyan Desert, near Oasis of Cufra, 

Libya. 

Meriones libycus crassibulla Lataste 

Meriones shawi var. crassibulla Lataste, Explor. Sci. de Tunisie, Cat. Critique 
d. Mamm., p. 27, 1887. Tebessa, Tamesmida, and Taferma, Tunis. 

Meriones libycus guyonii (Loche) 

Gerbillus guyonii Loche, Exploration Sci. de I'Algerie, Zool., Mammiferes, p. 
103, 1867. Region of Ain-el-Atrech, Algeria. 
A valid race according to Thomas, O., Ann. Mag. Nat. Hist. (9) 3: 271, 
Mar. 1919. 

Meriones libycus laticeps Lataste 

Meriones shawi var. laticeps Lataste, Explor. Sci. de Tunisie, Cat. Critique d. 
Mamm., p. 27, 1887. Tunis. 
See also Lataste, F., Proc. Zool. Soc. London, p. 94, text-f. 1 (skull), June 
1884. 

Meriones libycus longiceps Lataste 

Meriones shawi var. longiceps Lataste, Exploration Sci. de Tunisie, Cat. 
Critique d. Mamm., p. 27, 1887. Tunis. Status uncertain. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 329 

Meriones libycus pallidus Bonhote 

Meriones crassus pallidus Bonhote, Abstr. Proc. Zool. Soc. London, no. 103, 

p. 3, 13 Feb. 1912; Proc. Zool. Soc. London, p. 226, 13 Mar. 1912. Atbara, 

Anglo-Egyptian Sudan. 

Meriones libycus tripolius Thomas 

Meriones pallidus tripolius Thomas, Ann. Mag. Nat. Hist. (9) 3: 265, Mar. 
1919. Gebel Limhersuk, northwestern Tripoli. 

Meriones mari.e Cabrera 

Meriones marise Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 7: 177, 
Apl. 1907. Tarfaya, Sahara region, Cabo Juby, Rio de Oro. 

Meriones schousbceii schousbceii (Loche) 

Gerbillus schoushoeii Loche, Cat. des Mamm. et Oiseaux de I'Alg^rie, p. 23, 

1858; Exploration Sci. de I'Algerie, Zool., Mammiferea, p. 105, 1867. 

Ras-nili, Sahara, Algeria. 
Gerbillus renaultii Loche, Exploration Sci. de I'Algerie, Zool., Mammif^res, p. 

106, 1867. Environs of Messad, Sahara, Algeria. 
Meriones gMulus Lataste, Le Naturaliste, 4: no. 11, p. 83, 1 June 1882. Algeria; 

specimens mentioned from Tilremt, between Laghouat and Berrian, 

Sahara, and from M'sila, high plateaux. 

Meriones schousbceii tuareg Thomas 

Meriones schousboei tuareg Thomas, Ann. Mag. Nat. Hist. (9) 16: 193, July 
1925. Teguidda, west of Asben, French West Africa (Sahara). 

Meriones trouessarti Lataste 

Meriones trouessarti Lataste, Le Naturaliste, 4: no. 9, p. 69, 1 May 1882. Bou- 

Sdada, and Oued Magra, near M'sila, Algeria. 
Meriones melanurus Riippell, Mus. Senckenburgianum, 3: 95, 115, pi. 7, f. 3, 
1842. Alexandria, Egypt, and Tor, Arabia. 
See Thomas, O., (Ann. Mag. Nat. Hist. (9) 3: 264, Mar. 1919) as to status 
and lectotype. 

MiCRODiLLUs Thomas 

Microdillus Thomas, Ann. Mag. Nat. Hist. (8) 5: 197, Feb. 1910. Genotype 
Gerbillus peeli De Winton. 

Microdillus peeli (De Winton) 

Gerbillus peeli De Winton, Ann. Mag. Nat. Hist. (7) 1 : 250, Mar. 1898 (skull 
but not skin as described). Eyk, Somaliland. 



330 bulletin: museum of comparative zoology 



Pachyuromys Lataste 

Pachyuromys Lataste, Le Naturaliste, 2: no. 40, p. 313, 15 Nov. 1880. Geno- 
type Pachyuromys duprasi Lataste. 

Pachyuromys duprasi duprasi Lataste. The "Boubieda" 
Pachyuromys duprasi Lataste, Le Naturaliste, 2: no. 40, p. 313, 15 Nov. 1880. 
Algerian Sahara. 

Pachyuromys duprasi faroulti Thomas 

Pachyuromys duprasi faroulti Thomas, Novitates Zool., 27: 313, 15 June 1920. 
Mecheria, 100 km. northwest of Ain Sefra, plateau of western Algeria. 

Pachyuromys duprasi natronensis De Winton 

Pachyuromys dupresi (sic) natronensis De Winton, Novitates Zool., 10: 285, 

pi. 8, f. 3, 25 Aug. 1903. Bir Victoria, on the way to Wadi Natron from 

the Nile, Egypt. 



PsAMMOMYS Cretzschmar. Sand Rats 

Psaynmomys Cretzschmar, in RiippeU's Atlas zu d. Reise im nordliche Afrika, 
Saugeth., 1 : 56, 1828. Genotype Psammomys obesus Cretzschmar. 

PSAMMOMYS ELEGANS Hcuglin 

Psammomys elegans Heuglin, Reise in Nordost-Afrika, 2: 80, 1877. "Sauakin" 
or Suakin, Red Sea, Anglo-Egyptian Sudan. 

Psammomys algiricus Thomas 

Ps[ammomys] algiricus Thomas, Ann. Mag. Nat. Hist. (7) 9: 363, 364, May 
1902. Biskra, southern Algeria. 

Psammomys obesus obesus Cretzschmar 

Psammomys obesus Cretzschmar, in RiippeU's Atlas zu d. Reise im nordliche 

Afrika, Saugeth., p. 58, pi. 22, 23, 1826. Alexandria, Egypt. 
Psammomys roudairei Lataste, Le Naturaliste, 3: no. 62, p. 492, 15 Oct. 1881. 

Thomas, Proc. Zool. Soc. London, 2: 10, Oct. 1902. Algeria. 
Range: Algeria, Tunis and Libya to northern Lower Egypt and southern 

Palestine. 



Psammomys obesus nicolli Thomas 

Psammomys obesus nicolli Thomas, Ann. Mag. Nat. Hist. (8) 2: 92, July 1908. 
Damietta, northern Egypt. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 331 

PSAMMOMYS TRIPOLITANUS ThomaS 

Psammomys tripolitanus Thomas, Proc. Zool. Soc. London, 2: 9, Oct. 1902. 
Bou Cheifa, coast of Tripoli. 

Psammomys vexillaris vexillaris Thomas 

Psammomys vexillaris Thomas, Ann. Mag. Nat. Hist. (9) 16: 198, July 1925. 
Bondjem, Tripoli. 

Psammomys vexillaris edusa Thomas 

Psammomys vexillaris edusa Thomas, Ann. Mag. Nat. Hist. (9) 16: 199, July 
1925. Mil Mahases, Chegga, just south of Biskra, Algeria. 



Tatera Lataste 

Tatera Lataste, Le Naturaliste, 4: no. 16, p. 126, 15 Aug. 1882. Subgenus of 

Gerbillus; type Gerbillus indicus Hardwicke. 
Taterona Wroughton, Journ. Bombay Nat. Hist. Soc, 25: 40, 20 Mar. 1917. 
Genotype Gerbillus afer Gray. A synonym of Tatera, see Hollister; also 

Roberts, Ann. Transvaal Mus., 11: 251, 11 Sep. 1926. 

Review: Wroughton, R. C. Notes on the genus Tatera, with descriptions of 
new species. Ann. Mag. Nat. Hist. (7) 17: 474-499, May 1906; with key. 



Subgenus Tatera Lataste 

Tatera afra afra (Gray) 

Gerbillus afra Gray, Spicilegia Zool., 2: 10, 1 Aug. 1830. Cape of Good Hope. 

Meriones schlegelii Smuts, Enumeratio Mamm. Capens., p. 41, pi. 1; pi. 3, 

fig. 1-5, 1832. Cape of Good Hope, Port Elizabeth. 
Gerbillus africanus F. Cuvier, Trans. Zool. Soc. London, 2: pi. 26, f. 5-9, 1838. 

No locality, but presumably Cape of Good Hope. 

Tatera afra gilli Roberts 

Tatera afra gilli Roberts, Ann. Transvaal Mus., 13: 100, 6 July 1929. Lambert's 
Bay, Cape Colony. 

Tatera angola Wroughton 

Tatera ajigolse Wroughton, Ann. Mag. Nat. Hist. (7) 17: 488, May 1906. Fort 
Quilenges, Angola. 

Tatera benvenuta benvenuta (Hinton and Kershaw) 
Taterona benvenuta Hinton and Kershaw, Ann. Mag. Nat. Hist. (9) 6: 97, 
July 1920. Bahr-el-Jebel, Mongalla, Anglo-Egyptian Sudan. 



332 bulletin: museum of comparative zoology 

Tatera benvenuta lxjcia (Hinton and Kershaw) 

Taterona benvenuta lucia Hinton and Kershaw, Ann. Mag. Nat. Hist. (9) 6: 99,, 
July 1920. Musisi River, Lake Albert, Kenya Colony. 

Tatera bodessana Frick 

Tatera vicina bodessana Frick, Ann. Carnegie Mus., 9: 15, Sep. 1914. Bodessa, 
southern Abyssinia, 5000 ft. 

Tatera brantsii brantsii (A. Smith) 

Gerbillus brantsii A. Smith, Rept. Exped. for E.xploring Central Africa, from 

Cape of Good Hope, June 23, 1834, under Superintendence of Dr. A. 

Smith, p. 43, 1836. "Tops of hills towards sources of Caledon River," 

Orange Free State. 
Meriones montanus A. Smith, Illustr. Zool. South Africa, Mammals, pi. 36, f. 

1 and text, Feb. 1842. "Bashartoo Country" (?=Basutoland). 
Meriones (Rhombomys) maccalinus Sundevall, Ofversigt af Kongl. Vet.-Akad. 

Forhandl., Stockholm, for 1846, 3: 120, 1847. "In Caffraria interiore, 

prope montes nomine Maccali cognitos," i.e., Magaliesberg, Transvaal. 
Not a valid form according to Roberts, A., Ann. Transvaal Mus., 11: 250, 
14 Sep. 1926. 

Tatera brantsii perpallida Dollman 

Tatera brantsi perpallida Dollman, Ann. Mag. Nat. Hist. (8) 6: 394, Oct. 1910. 
East bank of the Tamalakan River, Ngamiland, Bechuanaland. 

Tatera breyeri Roberts 

Tatera breyeri Roberts, Ann. Transvaal Mus., 11: 250, 14 Sep. 1926. Nyls- 
troom, Transvaal. 

Tatera caffer (Wagner) 

Aferiones ca^er Wagner, Arch. f.Naturgesch., 8: sec. l,p. 18, 1842. South Africa. 

Tatera cosensi (Kershaw) 

Taterona cosensi Kershaw, Ann. Mag. Nat. Hist. (9) 8: 567, Nov. 1921. Vi- 
hingo, near Ruvu Station, 40 miles inland from Dar-es-Salaam, Tangan- 
yika Territory. 

Tatera dichrura Thomas 

Tatera dichrura Thomas, Ann. Mag. Nat. Hist. (8) 16: 147, Aug. 1915. Poko, 
Upper Uele River, Congo Beige. 

Tatera draco Wroughton 

Tatera draco Wroughton, Ann. Mag. Nat. Hist, (7) 17: 479, May 1906. Wak- 
kerstroom, Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 333 

Tatera fallax Thomas and Schwann 

Taterafallax Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 6, p. 22, 

26 Apl. 1904; Proc. Zool. Soc. London, 1: 461, 2 Aug. 1904. Burumba, 

Ankole, southwestern Uganda. 

Tatera flavipes G. M. Allen 

Tatera flavipes G. M. Allen, Bull. Mus. Comp. Zool., 58: 331, text-f. 2, July 
1914. Aradeiba, above Roseires, Blue Nile, Anglo-Egyptian Sudan. 

Tatera qambiana Thomas 

Tatera gambiana Thomas, Ann. Mag. Nat. Hist. (8) 6: 428, Oct. 1910. Mara- 
kissa, Gambia. 

Tatera giffardi Wroughton 

Tatera giffardi Wroughton, Ann. Mag. Nat. Hist. (7) 17: 489, May 1906. 
Gambaga, Gold Coast. 

Tatera guinea guinea Thomas 

Tatera guinex Thomas, Ann. Mag. Nat. Hist. (8) 5: 353, Apl. 1910. Gunnal, 
Portuguese Guinea. 

Tatera guinea picta (Hayman) 

Taterona guinex picta Hayman, Proc. Zool. Soc. London, for 1935, p. 930, 10 
Jan. 1936. Pong, Tamale, Northern Territories of Gold Coast. 

Tatera hopkinsoni Thomas 

Tatera hopkinsoni Thomas, Ann. Mag. Nat. Hist. (8) 8: 375, Sep. 1911. Ku- 
dang, Gambia. 

Tatera humpatensis (Hill and Carter) 

Taterona humpatensis Hill and Carter, Amer. Mus. Novitates, no. 913, p. 5, 
f. 3, 8 Mar. 1937. Humpata, Angola, 6300 ft. 

Tatera inclusa Thomas and Wroughton 

Tatera inclusa Thomas and Wroughton, Proc. Zool. Soc. London, p. 169, 
7 Sep. 1908. Tambarara, Gorongoza district, Mozambique. 

Tatera joan^ (Thomas) 

Taterona joanse Thomas, Proc. Zool. Soc. London, p. 299, 29 Apl. 1926. Uku- 
ambi, Ovamboland, Southwest Africa. 

Tatera kempii Wroughton 

Tatera kempii Wroughton, Ann. Mag. Nat. Hist. (7) 17: 375, Apl. 1906. Agou- 
lerie, Anambra Creek, southern Nigeria. 



334 bulletin: museum of comparative zoology 

Tatera leucogaster (Peters) 

Meriones leucogaster Peters, Reise nach Mossambique, Saugeth., p. 145, pi. 33,, 
f. 1; pi. 35, f. 4, 1852. Mesuril and Boror, Mozambique. 

Tatera liodon liodon Thomas 

Tatera liodon Thomas, Ann. Mag. Nat. Hist. (7) 9; 441, June 1902. Lake 
Mweru, northern Nyasaland. 

Tatera liodon dtjndasi Wroughton 

Tatera dundasi Wroughton, Ann. Mag. Nat. Hist. (8) 4: 539, Dec. 1909. 
Kirui, Mt. Elgon, Kenya Colony, 6000 ft. 

Tatera liodon smithi Wroughton 

Tatera smithi Wroughton, Ann. Mag. Nat. Hist. (8) 3: 249, Mar. 1909. Mu- 
bende, Unyoro, Uganda. 

Tatera lobengul.e lobengul.e (De Winton) 

Gerbillus {Tatera) lobengulae De Winton, Ann. Mag. Nat. Hist. (7) 2: 4, July 
1898. Essex Vale, Matabeleland, Southern Rhodesia. 

Tatera lobengul^ bechuan.e Wroughton 

Tatera lohengulse bechuanse Wroughton, Ann. Mag. Nat. Hist. (7) 17: 482, May 
1906. Molopo, Bechuanaland. 

Tatera lobengul.e beirensis Roberts 

Tatera lobengulie beirensis Roberts, Ann. Transvaal Mus., 13: 106, 6 July 1929. 
Near Beira, Mozambique. 

Tatera lobengul^ griqu^ Wroughton 

Tatera lobengulse griquse Wroughton, Ann. Mag, Nat. Hist. (7) 17: 483, May 

1906. Kuruman, Bechuanaland. 
Tatera miliaria stellae Wroughton, Ann. Mag. Nat. Hist. (7) 17: 485, May 1906. 

Kuruman, Bechuanaland, 4000 ft. 
Identical with griquse (Roberts, A., Ann. Transvaal Mus., 13: 102, 6 July 

1929). 

Tatera lobengul.e limpopoensis Roberts 

Tatera lobengulx limpopoensis Roberts, Ann. Transvaal Mus., 13: 104, 6 July 
1929. Njellele River, Zoutpansberg district, Transvaal. 

Tatera lobenqul^ littoralis Roberts 

Tatera lobengulse littoralis Roberts, Ann. Transvaal Mus., 13: 105, 6 July 1929. 
Masiene, near mouth of Limpopo River, Mozambique. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 335 

Tatera lobengtjl^ MASHON.E Wroughton 

Tatera lobengulx mashonae. Wroughton, Ann. Mag. Nat. Hist. (7) 17: 483, 
May 1906. Mazoe, Mashonaland, Rhodesia. 

Tatera lobenqxjl^ mitchelli Roberts 

Tatera lobenguls mitchelli Roberts, Ann. Transvaal Mus., 13: 103, 6 July 1929. 
Wonder fontein, Transvaal. 

Tatera lobengtjl^ ndol^ (Kershaw) 

Taterona lobengulse ndolx Kershaw, Ann. Mag. Nat. Hist. (9) 10: 105, July 
1922. N'Dola, Northern Rhodesia, 12° 50' S., 28° 40' E. 

Tatera lobengulse panja Wroughton 

Meriones tenuis Peters (not A. Smith), Reise nach Mossambique, Saugeth., 

p. 149, 1852. Tette, Mozambique. 
Tatera panja Wroughton, Ann. Mag. Nat. Hist. (7) 17: 486, May 1906. 

Chicosta, 60 miles above Tette, Zambesi River, Mozambique. 
As to subspecific status, see Roberts, A., Ann. Transvaal Mus., 13: 103, 
6 July 1929. 

Tatera lobengulse pestis Roberts 

Tatera lobengulse pestis Roberts, Ann. Transvaal Mus., 13: 103, 6 July 1929. 
Bothaville, Orange Free State. 

Tatera lobengulse Pretoria Roberts 

Tatera lobengulse pretorise Roberts, Ann. Transvaal Mus., 13: 104, 6 July 1929. 
Pretoria North, Transvaal. 

Tatera lobengul^ salsa Wroughton 

Tatera miliaria salsa Wroughton, Ann. Mag. Nat. Hist. (7) 17: 485, May 1906. 
Klein Letaba, Zoutpansberg, Transvaal. 
As to status, see Roberts, A., Ann. Transvaal Mus., 13: 102, 6 July 1929. 

Tatera LOBENGULiE tzaneenensis Roberts 

Tatera lobengulse tzaneenensis Roberts, Ann. Transvaal Mus., 13: 105, 6 July 
1929. Tzaneen Estates, northeastern Transvaal. 

Tatera lobengul^ zuluensis Roberts 

Tatera lobengulse zuluensis Roberts, Ann. Transvaal Mus., 14: 230, 28 July 
1931. Manaba, 30 miles from Kosi Bay, northern Zululand, Natal. 

Tatera macropus (Heuglin) 

Meriones macropus Heuglin, Reise in Nordost-Afrika, 2: 79, 1877. Bongo, 
Bahr-el-Ghazal region, Anglo-Egyptian Sudan. 



336 bulletin: museltm of compakative zoology 

Tatera maputa Roberts 

Tatera maputa Roberts, Ann. Transvaal Mus., 18: 238, 21 Feb. 1936. Maputa, 
northern Zululand, Natal. 

Tatera miliaria Wroughton 

Tatera miliaria Wroughton, Ann. Mag. Nat. Hist. (7) 17: 484, May 1906. 
Deelfontein, Cape Colony. 

Tatera minuscula Osgood 

Tatera minuscula Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 230, 28 Dec. 
1936. Sheik Hussein, near Webi Shebeli, Bale, Ethiopia. 

Tatera natalensis Roberts 

Tatera natalensis Roberts, Ann. Transvaal Mus., 13: 101, 6 July 1929. Kil- 
gobbin, Lidgetton, Natal. 

Tatera neavei Wroughton 

Tatera neavei Wroughton, Mem. and Proc. Manchester Lit. and Phil. Soc, 51: 
noj 5, p. 18, 13 Mar. 1907. 'Ndola, Rhodesia. 

Tatera niqricatjda nigricauda (Peters) 

Gerbillus nigricaudus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 200, 
1878. Ndi, Taita, Kenya Colony. 

Tatera nigricauda bayeri Lonnberg 

Tatera nigricauda bayeri Lonnberg, Revue Zool. Africaine, 5: 179, 30 Apl. 
1918. Maroon River, near Mt. Elgon, Kenya Colony. 

Tatera nigricauda bodess^ Frick 

Tatera nigricauda hodessx Frick, Ann. Carnegie Mus., 9: 14, 6 June 1914. 
Sagan River, Bodessa, Abyssinia, 5000 ft. 

Tatera nigricauda nyama DoUman 

Tatera nigricauda nyama Dollman, Ann. Mag. Nat. Hist. (8) 7: 522, May 1911. 
Nyama Nyango, Northern Guaso Nyiro, Kenya Colony. 

Tatera nigricauda percivali Heller 

Tatera nigracauda (sic) percivali Heller, Smithsonian Misc. Coll., 63: no. 7, p. 
8, 24 June 1914. Lorian Swamp, Kenya Colony. 

Tatera nigrita nigrita Wroughton 

Tatera nigrita Wroughton, Ann. Mag. Nat. Hist. (7) 17: 491, May 1906j 
Masindi, Unyoro, Uganda. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 337 

Tatera nigrita beniensis Hatt 

Tatera nigrita beniensis Hatt, Amer. Mus. Novitates, no. 791, p. 2, 11 Apl. 
1935. New post of Beni (Bungulu), eastern Congo Beige, 3900 ft. 

Tatera nyas^ nyas^ Wroughton 

Tatera nyasse Wroughton, Ann. Mag. Nat. Hist. (7) 17: 490, May 1906. 
Deep Bay, northern Nyasaland. 

Tatera nyas^ loveridgei Hatt 

Tatera nyassx loveridgei Hatt, Amer. Mus. Novitates, no. 791, p. 2, 11 Apl. 
1935. Kilosa, Tanganyika Territory. 

Tatera nyas^ shirensis Wroughton 

Tatera nyasse shirensis Wroughton, Ann. Mag. Nat. Hist. (7) 17: 490, May 
1906. Mount Malosa, Upper Shu-e, Nyasaland. 

Tatera phillipsi phillipsi (De Winton) 

Gerbillus (Tatera) phillipsi De Winton, Ann. Mag. Nat. Hist. (7) 1 : 253, Mar. 
1898. Hanka Dadi, Somaliland. 
Named for E. Lort-Phillips 

Tatera phillipsi ttmbrosa DoUman 

Tatera phillipsi umbrosa Dollman, Ann. Mag. Nat. Hist. (8) 9: 219, Feb. 1912. 
Baringo, Kenya Colony. 

Tatera robusta robusta (Cretzschmar) 

Meriones robustus Cretzschmar, in Riippell's Atlas zu d. Reise im nordl. Afrika, 

Saugeth., p. 75, pi. 29, f. b, 1826. Kordofan. 
Meriones murinus Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 

for 1842, p. 231, 1843. Bahr-el-Abiad, Anglo-Egyptian Sudan. 

Tatera robusta taylori Hatt 

Tatera robusta taylori Hatt, Amer. Mus. Novitates, no. 791, p. 1, 11 Apl. 1935. 
Khor Birum, Red Sea hills, Anglo-Egyptian Sudan. 

Tatera rxjddi ruddi Wroughton 

Tatera ruddi Wroughton, Ann. Mag. Nat. Hist. (7) 17: 478, May 1906. Um- 
volosi, Zululand, Natal. 

Tatera ruddi tongensis Roberts 

Tatera ruddi tongensis Roberts, Ann. Transvaal Mus., 14: 230, 28 July 1931. 
Maputa, northern Zululand, Natal. 



338 bulletin: museum of comparative zoology 

Tatera RtJWENzoRii Thomas and Wroughton 

Tatera ruwenzorii Thomas and Wroughton, Trans. Zool. Soc. London, 19: 500, 
Mar. 1910. Mokia, southeastern Ruwenzori, Uganda. 

Tatera schinzi (Noack) 

Gerbillus tenuis, var. schinzi Noack, Zool. Jahrb., Syst., 4: 134, pi. 3, f. 13-16, 
28 Feb. 1889. Kalahari Desert, Southwest Africa. 

Tatera shoana Wroughton 

Tatera shoana Wroughton, Ann. Mag. Nat. Hist. (7) 17: 492, May 1906. Jefir 
Medir, British Somaliland. 

Tatera soror G. M. Allen 

Tatera soror G. M. Allen, Bull. Mus. Comp. Zool,, 58: 333, text-f. 3, July 1914. 
Fazogli, Blue Nile, Anglo-Egyptian Sudan. 

Tatera swaythlingi (Kershaw) 

Taterona swaythlingi Kershaw, Ann. Mag. Nat. Hist. (9) 8: 565, Nov. 1921. 
Morogoro, 150 miles west of Dar-es-Salaam, Tanganyika Territory. 

Tatera tabors (Kershaw) 

Taterona tahorse Kershaw, Ann. Mag. Nat. Hist. (9) 8: 566, Nov. 1921. Tabora, 
Tanganyika Territory. 

Tatera validtjs (Bocage) 

Gerbillus validus Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 2: no. 5, 

p. 6, pi. 1, f. 1, la, Sep. 1890. Ambaca, Quissange, Caconda, Rio Cuando, 

Angola; 

Tatera vicina vicina (Peters) 

Gerbillus vicinus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 200, 1878. 

Kitui, Ukambani, Kenya Colony. 
Tatera mombasse Wroughton, Ann. Mag. Nat. Hist. (7) 17: 493, May 1906. 

Takangu, Kenya Colony. 

Tatera vicina bodessana Frick 

Tatera vicina bodessana Frick, Ann. Carnegie Mus., 9: 15, 6 June 1914. Bo- 

dessa, southern Abyssinia, 5000 ft. 

Tatera vicina iconica Dollman 

Tatera iconica Dollman, Ann. Mag. Nat. Hist. (8) 7: 521, May 1911. Nyama 
Nyango, Northern Guaso Nyiro, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 339 

Tatera viciNA MUANS.E (Matschie) 

Gerbillus (Tatera) vicinus muansse Matschie, Sitzb. Ges. Naturf. Freunde, 
Berlin, p. 333, Oct. 1911. Mwansa, south coast of Victoria Nyanza, Tan- 
ganyika Territory. 

Tatera vicina poth^ Heller 

Tatera pothas Heller, Smithsonian Misc. Coll., 56: no. 9, p. 2, pi. 1, 4 figs., 22 
July 1910. Potha, Kapiti Plains, Kenya Colony. 

Tatera welmanni (St. Leger) 

Taterona welmanni St. Leger, Ann. Mag. Nat. Hist. (10) 3: 387, Apl. 1929. 
Maiduguri, northeastern Nigeria. 



Subgenus Gerbilliscus Thomas 

Gerbilliscus Thomas, Proc. Zool. Soc. London, p. 433, 1 Oct. 1897. Genotype 
Tatera bohmi (Noack). 

Tatera bohmi bohmi (Noack) 

Gerbillus bohmi Noack, Zool. Jahrb., Syst., 2: 241, pi. 9, f. 11-13 (skull), 7 May 
1887. Qua Mpala, Marunga, Northern Rhodesia. 

Tatera bohmi varia Heller 

Tatera varia Heller, Smithsonian Misc. Coll., 56: no. 9, p. 1, pi. 1, 4 figs., 22 July 
1910. Loletai plains, Southern Guaso Nyiro, Kenya Colony. 

Tatera fraterculus (Thomas) 

Gerbillus (Gerbilliscus) fraterculus Thomas, Proc. Zool. Soc. London, p. 392, 
1 Oct. 1898. Songwe, northern Nyasaland. 



Taterillus Thomas 

Taterillus Thomas, Ann. Mag. Nat. Hist. (8) 6: 222, Aug. 1910. Genotype 

Gerbillus emini Thomas. 
Taterina Wettstein, Anzeiger K. Akad. Wiss. Wien, math.-nat. KL, 53: 152, 

1916. Subgenus of Gerbillus, type Gerbillus lorenzi Wettstein. 

See Wroughton, R. C. African geVbils of the genera Tatera and Taterillus. 
Ann. Mag. Nat. Hist. (8) 6: 292-294, Sep. 1910 (with key to species of 
Taterillus). 

Taterillus butleri Wroughton 

Taterillus butleri Wroughton, Ann. Mag. Nat. Hist. (8) 6: 293, 294, Sep. 1910. 
Dugdug, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 



340 bulletin: museum of comparative zoology 

Gerbillus (Taterillus) kadugliensis Wettstein, Anzeiger K. Akad. Wiss. Wien, 
math.-nat. Kl., 53: 151, 1916. Kadugli, southern Kordofan, Anglo- 
Egyptian Sudan. 
Regarded as a synonym of Taterillus butleri by Thomas and Hinton (Proc. 
Zool. Soc. London, p. 257, 6 July 1923). 

Gerbillus (Taterina) lorenzi Wettstein, Anzeiger K. Akad. Wiss. Wien, math.- 
nat. KL, 53: 152, 1916. El Obeid, Kordofan, Anglo-Egyptian Sudan. 
As to status see Thomas, O., and Hinton, M. A. C, Proc. Zool. Soc. London, 
p. 257, 6 July 1923. 

Taterillus clivosus Thomas and Hinton 

Taterillus clivosus Thomas and Hinton, Proc. Zool. Soc. London, p. 258, 6 July 
1923. Jebel Marra, Darfur, Anglo-Egyptian Sudan. 

Taterillus congicus Thomas 

Taterillus congicus Thomas, Ann. Mag. Nat. Hist. (8) 16: 147, Aug. 1915. 
Poko, Upper Uele River, Congo Beige. 

Taterillus emini emini (Thomas) 

Gerbillus emini Thomas, Ann. Mag. Nat. Hist. (6) 9: 78, Jan. 1892. Wadelai, 
Uganda. 

Taterillus emini anthonyi Hatt 

Taterillus emini anthonyi Hatt, Amer. Mus. Novitates, no. 708, p. 2, text-f., 
4 Apl. 1934. White Nile, west bank, 20 miles south of Jebel Ain, Anglo- 
Egyptian Sudan. 

Taterillus emini zammarani de Beaux 

[Taterillus emini] zammarani de Beaux, Atti Soc. Ital. Sci. Nat. e Mug. Civ, 
Stor. Nat. Milano, 61: 27, Feb. 1922. Dolo, southern Somaliland. 

Taterillus gracilis gracilis (Thomas) 

Gerbillus gracilis Thomas, Ann. Mag. Nat. Hist. (6) 9: 77, Jan. 1892. Gambia. 

Taterillus gracilis angelus Thomas and Hinton 

Taterillus gracilis angelus Thomas and Hinton, Novitates Zool., 27: 317, 15 
June 1920. Farniso, near Kano, northern Nigeria. 

Taterillus gyas Thomas 

Taterillus gyas Thomas, Ann. Mag. Nat. Hist. (9) 2: 150, Aug. 1918. Kamisa, 
Binder River, Anglo-Egyptian Sudan. 

Taterillus harringtoni (Thomas) 

Tatera harringtoni Thomas, Ann. Mag. Nat. Hist. (7) 18: 303, Oct. 1906. 
Mutti Caleb, east of Lake Rudolf, Abyssinia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 341 

Taterillus lacustris (Thomas and Wroughton) 

Tatera lacustris Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 378^ 
May 1907. Lake Chad. 

Taterillus lowei DoHman 

Taterillus lowei Dollman, Abstr. Proc. Zool. Soc. London, no. 131, p. 25, 14 Apl. 

1914; Proc. Zool. Soc. London, p. 312, 16 June 1914. Ten miles west of 

Ngamatak Hills, Turkwel River, Uganda. 

Taterillus melanops G. M. Allen 

Taterillus inelanops G. M. Allen, Bull. Mus. Comp. Zool., 54: 446, Apl. 1912. 
Meru River, Northern Guaso Nyiro, Kenya Colony. 

Taterillus Nigeria Thomas 

Taterillus nigerise Thomas, Ann. Mag. Nat. Hist. (8) 7: 459, May 1911. Kabir, 
northern Nigeria. 

Taterillus nubilus nubilus Dollman 

Taterillus nubilus Dollman, Ann. Mag. Nat. Hist. (8) 8: 656, Nov. 1911. Orr 
Valley, Mt. Nyiro, Kenya Colony. 

Taterillus nubilus illustris Dollman 

Taterillus nubilus illustris Dollman, Ann. Mag. Nat. Hist. (8) 8: 656, Nov. 
1911. Twelve miles north of the Northern Guaso Nyiro, Kenya Colony. 

Taterillus osgoodi Wroughton 

Taterillus osgoodi Wroughton, Ann. Mag. Nat. Hist. (8) 6: 293, Sep. 1910. Voi, 
Kenya Colony. 

Taterillus perluteus Thomas and Hinton 

Taterillus perluteus Thomas and Hinton, Proc. Zool. Soc. London, p. 259, 

6 July 1923. Um Kedada, about 100 miles east of the capital, Darfur, 

Anglo-Egyptian Sudan. 

Taterillus rufus (Wettstein) 

Gerbillus (Tatera) rufa Wettstein, Anzeiger K. Akad. Wiss. Wien, math.-nat. 
Ivl., 53: 151, 1916. El Obeid, northern Kordofan, Anglo-Egyptian Sudan. 
As to status, see Thomas and Hinton, Proc. Zool. Soc. London, p. 259, 
6 July 1923. 

Taterillus tenebricus Dollman 

Taterillus tenebricus Dollman, Ann. Mag. Nat. Hist. (8) 7: 520, May 1911. 
Nyama Nyango, Northern Guaso Nyiro, Kenya Colony. 



342 bulletin: museum of comparative zoology 



Subfamily OTOMYIN^ Groove-toothed Rats 

See Thomas, Oldfield. A revised classification of the Otomyinae, with de- 
scriptions of new genera and species. Ann. Mag. Nat. Hist. (9) 2: 203-211, 
Sep. 1918. 

Myotomys Thomas 

Myotomys Thomas, Ann. Mag. Nat. Hist. (9) 2: 204, 206, Sep. 1918. Genotype 
Otomys unisulcatus Brants. 

Myotomys sloggetti slogqetti (Thomas) 

Otomys sloggetti Thomas, Ann. Mag. Nat. Hist. (7) 10 : 31 1, Oct. 1902. Deelfon- 
tein, Cape of Good Hope. 

Myotomys sloggetti basuticus Roberts 

Myotomys sloggetti basuticus Roberts, Ann. Transvaal Mus., 13: 110, 6 July 
1929. Bolepeletsa, Basutoland, Cape of Good Hope. 

Myotomys sloggetti jeppei Roberts 

Myotomys sloggetti jeppei Roberts, Ann. Transvaal Mus., 13: 109, 6 July 1929. 
Jamestown, Cape of Good Hope. 

Myotomys turneri (Wroughton) 

Otomys turneri Wroughton, Ann. Mag. Nat. Hist. (7) 20: 31, July 1907. Aber- 
feldy, northeastern Orange Free State. 

Myotomys unisulcatus unisulcatus (F. Cuvier) 

Otomys unisulcatus F. Cuvier, Hist. Nat. des Mammiferes, pt. 60, 1829. South 
Africa. 

Myotomys unisulcatus bergensis Roberts 

Myotomys imisulcatus bergensis Roberts, Ann. Transvaal Mus., 13: 108, 6 July 
1929. Lambert's Bay, Cape of Good Hope. 

Myotomys unisulcatus broomi (Thomas) 

Otomys broomi Thomas, Ann. Mag. Nat. Hist. (7) 10: 313, Oct. 1902. Port 
Nolloth, Little Namaqualand, Cape of Good Hope. 

Myotomys unisulcatus grantii (Thomas) 

Otomijs unisulcatus grantii Thomas, Ann. Mag. Nat. Hist. (7) 10: 312, Oct. 1902. 
Deelfontein, Cape of Good Hope. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 343 

Otomys F. Cuvier 

Otomys F. Cuvier, Dents des Mammiferes, p. 168, 255, pi. 60, 1823. Genotype 

Euryotis irroratus Brants. 
Euryotis Brants, Het Geslacht der Muizen, p. 93, 1827. Genotype Euryotis 

irroratus Brants. 
Oreomys Heuglin, Reise in Nordost-Afrika, 2: 76, 1877. Genotype Oreomys 

typus Heuglin. 
Oreinomys Trouessart, Bull. Sec. d'Etudes Sci., Angers, 10: 111, 1880, and 

footnote. Substitute for Oreomys Heuglin, 1877, preoccupied. 

See Wroughton, R. C. Notes on the genus Otomys. Ann. Mag. Nat. Hist. (7) 
18:264-278, Oct. 1906. 

Thomas, O. On some new forms of Otomys. Ann. Mag. Nat. Hist. (7) 10: 
311-314, 1902. 

Dollman, Guy. On the swamp-rats (Otomys) of East Africa. Ann. Mag. 
Nat. Hist. (8) 15: 149-170, Jan. 1915. 



Subgenus Anchotomys Thomas 

Anchotomys Thomas, Ann. Mag. Nat. Hist. (9) 2: 204, 208, Sep. 1918. As a 
subgenus of Otomys; type and only species Otomys anchietas Bocage. 

Otomys anchiet^e anchiet^ (Bocage) 

Euryotis anchietas Bocage, Jorn. Sci. Phys., Math, e Nat., Lisboa, (1)9: 26, July 
1882. Caconda, Angola. 

Otomys anchiet^ lacustris G. M. Allen and Loveridge. 

Otomys (Anchotomys) anchietse lacustris G. M. Allen and Loveridge, Bull. Mus. 

Comp. Zool., 75: 120, 7 Feb. 1933. Madehani, Ukinga Mts., north end of 

Lake Nyasa, Tanganyika Territory. 



Subgenus Otomys F. Cuvier 

Otomys angoniensis angoniensis Wroughton 

Otomys irroratus angoniensis Wroughton, Ann. Mag. Nat. Hist. (7) 18: 274, 

Oct. 1906. M'Kombhuie, Angoniland, Nyasaland. 
Otomys angoniensis Dollman, Ann. Mag. Nat. Hist. (8) 15: 163, Jan. 1915. 

Otomys angoniensis elassodon Osgood 

Otomys angoniensis elassodon Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: 10, 16 Feb. 1910. Naivasha, Kenya Colony. 



344 bulletin: museum of comparative zoology 

Otomys burtoni Thomas 

Otomys burtoni Thomas, Ann. Mag. Nat. Hist. (9) 2: 210, Sep. 1918. Camer- 
oon Mts., French West Africa, 7000 ft. 
For Sir Richard Burton. 

Otomys cuanzensis Hill and Carter 

Otomys cuanzensis Hill and Carter, Amer. Mus. Novitates, no. 913, p. 7, f. 4, 
8 Mar. 1937. Chitau, Angola, 4930 ft. 

Otomys dartmouthi Thomas 

Otomys dartmouthi Thomas, Ann. Mag. Nat. Hist. (7) 18: 141, Aug. 1906. 
Ruwenzori East, Uganda, 12,500 ft. 

Otomys denti Thomas 

Otomys denti Thomas, Ann. Mag. Nat. Hist. (7) 18: 142, Aug. 1906. Ruwen- 
zori East, Uganda, 6000 ft. 

Otomys divinorum Thomas 

Otomys divinorumThomas, Ann. Mag. Nat. Hist. (8) 6: 311, Sep. 1910. Rombo, 
Mt. KiUmanjaro, Tanganyika Territory, 5300 ft. 

Otomys irroratus irroratus (Brants) 

E[uTyotis\ irroratus Brants, Het Geslacht der Muizen, p. 94, pi., 1827. Uiten- 

hage. Cape of Good Hope (see Roberts, Ann. Transvaal Mus., 13: 111, 

6 July 1929). 
Otomys bisulcatus F. Cuvier, Hist. Nat. des Mammif^res, pt. 61, 1829. 
Euryotis typicus A. Smith, South African Quart. Journ., 2: 149, 1834. Near 

Constantia, Orange Free State. 
Euryotis obscura Lichtenstein, Verzeichn. Samml. d. Saugeth. u. Vogeln Kaf- 

ernlande, p. 10, 1842. South Africa, "Kaifirland." 

Otomys irroratus atjratus Wroughton 

Otomys irroratus auratus Wroughton, Ann. Mag. Nat. Hist. (7) 18: 272, Oct. 
1906. Vredefort, Orange Free State. 

Otomys irroratus ccenosus Thomas 

Otomys irroratus ccenosus Thomas, Ann. Mag. Nat. Hist. (9) 2: 208, Sep. 1918. 
Kuruman, Bechuanaland. 

Otomys irroratus cupreus Wroughton 

Otomys irroratus cupreus Wroughton, Ann. Mag. Nat. Hist. (7) 18: 273, Oct. 
1906. Zoutpansberg, Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 345 

Otomys irroratus maximus Roberts 

Olomys irroratus maximus Roberts, Ann. Transvaal Mus., 10: 70, 31 Jan. 1924. 
Machile River, northwestern Rhodesia. 

Otomys irroratus natalensis Roberts 

Otomys irroratus natalensis Roberts, Ann. Transvaal Mus., 13: 111, 6 July 1929. 
Bilgobbin, Dargle district, Natal. 

Otomys irroratus randensis Roberts 

Otomys irroratus randensis Roberts, Ann. Transvaal Mus., 13: 112, 6 July 1929. 
Fontainebleau, Johannesburg, Transvaal. 

Otomys jacksoni jacksoni Thomas 

Otomys jacksoni Thomas, Ann. Mag. Nat. Hist. (6) 7: 304, Mar. 1891. [Crater 
of] Mt. Elgon, Uganda side, 13,000 ft. 

Otomys jacksoni helleri Frick 

Otomys jacksoni helleri Frick, Ann. Carnegie Mus., 9: 10, pi. 3, f. 1-5, 6 June 

1914. Chilalo Divide Camp, Abyssinia, 9000 ft. 
Otomys jacksoni mearnsi Frick, Ann. Carnegie Mus., 9: pi. 3, f. 1-5, 6 June 

1914 (accidental renaming of 0. jacksoni helleri Frick). 

Otomys jacksoni malkensis Frick 

Otomys jacksoni malkensis Frick, Ann. Carnegie Mus., 9: 11, text-f. 1, pi. 3, 
f. 6-10, 6 June 1914. Malka, Sidamo, Abyssinia (Ethiopia), 7000 ft. 

Otomys karoensis Roberts 

Oto7nys karoensis Roberts, Ann. Transvaal Mus., 14: 231, pi. 6, 7, 8, fig. b, 28 
July 1931. Wolseley, Cape of Good Hope. 

Otomys kempi Dollman 

Otomys kempi Dollman, Ann. Mag. Nat. Hist. (8) 15: 152, Jan. 1915. Burunga, 
Mt. Mikeno, Congo Beige, 6000 ft. 

Otomys mashona Thomas 

Otomys mashona Thomas, Ann. Mag. Nat. Hist. (9) 2: 210, Sep. 1918. Mazoe, 
Mashonaland, Southern Rhodesia. 

Otomys nyik^ nyik^ Wroughton 

Otomys irroratus nyikx Wroughton, Ann. Mag. Nat. Hist. (7) 18: 276, Oct. 

1906. Nyika Plateau, Nyasaland. 
Otomys nyikse Dollman, Ann. Mag. Nat. Hist. (8) 15: 164, Jan. 1915 (distinct 

species) . 



346 bulletin: museum of comparative zoology 

Otomys nyik^ canesckns Osgood 

Otomys nyikae canescens Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 
10, 16 Feb. 1910. Kijabe, Kenya Colony. 

Otomys orestes orestes Thomas 

Otomys irroratus orestes Thomas, Proc. Zool. Soc. London, p. 175, 1 June 1900. 

Teliki Valley, west slope of Mt. Kenya, Kenya Colony, 13,000 ft. 
Otomys orestes Dollman, Ann. Mag. Nat. Hist. (8) 15: 156, Jan. 1915 (distinct 

species). 

Otomys orestes dollmani Heller 

Otomys orestes dollmani Heller, Smithsonian Misc. Coll., 59: no. 16, p. 5, 5 July 
1912. Mt. Gargues, Mathews Range, Kenya Colony, 7000 ft. 

Otomys percivali Dollman 

Otomys percivali Dollman, Ann. Mag. Nat. Hist. (8) 15: 168, Jan. 1915. 

Naivasha district, 12 miles south of Lake OlboUossat, Kenya Colony, 

8700 ft. 

Otomys robertsi Hewitt 

Otomys robertsi Hewitt, Records Albany Mus., 3: 430, pi. 26, f. 3-4, 31 July 
1927. Mont-aux-Sources, Orange Free State-Basuto Land border. 

Otomys rowleyi Thomas 

Otomys rowleyi Thomas, Ann. Mag. Nat. Hist. (9) 2: 209, Sep. 1918. Coguno, 
Inhambane, Mozambique. 

Otomys rubecultjs Dollman 

Otomys ruheculus Dollman, Ann. Mag. Nat. Hist. (8) 15: 161, Jan. 1915. 
Kagambah, Uganda, 4800 ft. 

Otomys silberbaueri Roberts 

Otomys silberbaueri Roberts, Ann. Transvaal Mus., 6: 114, Apl. 1919. Lor- 
marins. Pearl district, Cape of Good Hope. 

Otomys thomasi thomasi Osgood 

Otomys thomasi Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 9, 16 Feb. 
1910. Molo, Kenya Colony. 

Otomys thomasi malleus Dollman 

Otomys thomasi malleus Dollman, Ann. Mag. Nat. Hist. (8) 15: 154, Jan. 1915. 
Lake OlboUossat, Naivasha district, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 347 



Otomys thomasi squalus Dollman 

Otomys thomasi squalus Dollman, Ann. Mag. Nat. Hist. (8) 15: 155, Jan. 1915. 
Mt. Kinangop, Aberdare Range, Kenya Colony, 12,000 ft. 

Otomys tropicalis tropicalis Thomas 

Otomys irroratus tropicalis Thomas, Ann. Mag. Nat. Hist. (7) 10: 314, Oct. 

1902. West slope of Mt. Kenya, Kenya Colony, 10,000 ft. 
Otomys tropicalis Dollman, Ann. Mag. Nat. Hist. (8) 15: 157, Jan. 1915 

(distinct species). 

Otomys tropicalis elgonis Wroughton 

Otomys irroratus elgonis Wroughton, Ann. Mag. Nat. Hist. (8) 5: 207, Feb. 
1910. Elgonyi, Mt. Elgon, Kenya Colony, 7000 ft. 

Otomys tropicalis faradjius Hatt 

Otomys tropicalis faradjius Hatt, Amer. Mus. Novitates, no. 708, p. 1, 4 Apl. 
1934. Faradje, Upper Uele district, Congo Beige. 

Otomys tropicalis ghigii de Beaux 

Otomys tropicalis ghigii de Beaux, Ann. Mus. Civ. Stor. Nat., Genova, 51: 213, 
7 Mar. 1924. Bugala, Uganda. 

Otomys tropicalis nubilits Dollman 

Otomys tropicalis nubilus Dollman, Ann. Mag. Nat. Hist. (8) 15: 160, Jan. 

1915. Jombeni (Igembi) Range, northeast of Mt. Kenya, Kenya Colony, 

6000 ft. 

Otomys tropicalis vivax Dollman 

Otomys tropicalis vivax Dollman, Ann. Mag. Nat. Hist. (8) 15: 159, Jan. 1915, 
Mt. Nyiro, south of Lake Rudolf, Kenya Colony, 8000 ft. 

Otomys tropicalis vulcanicus Lonnberg and Gyldenstolpe 
Otomys tropicalis vulcanicus Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17 B: 
no. 5, p. 2, 2 May 1925. Mt. Sabinio, Birimga Volcanoes, Congo Beige. 



Otomys tugelensis tugelensis Roberts 

Otomys tugelensis Roberts, Ann. Transvaal Mus., 13: 113, 6 July 1929. Klip- 
snruit. Utrecht. Natal. 



Otomys tugelensis pretorle Roberts 

Otomys tugelensis pretorise Roberts, Ann. Transvaal Mus., 13: 114, 6 July 1929. 
Fountains Valley, Pretoria, Transvaal. 



348 bulletin: museum of comparative zoology 

Otomys tugelensis sabiensis Roberts 

Otomys tugelensis sabiensis Roberts, Ann. Transvaal Mus., 13: 114, 6 July 
1929. Mariepskop, Lyndenburg district, Transvaal. 

Otomys tugelensis saundersi^ Roberts 

Otomys tugelensis saundersise Roberts, Ann. Transvaal Mus., 13: 115, 6 July 
1929. Grahamstown, Cape Colony. 

Otomys typus typus (Heuglin) 

Oreomys typus Heuglin, Reise in Nordost-Afrika, 2: 77, 1877. Highlands of 

Simen, Shoa, Abyssinia. 
Otomys degeni Thomas, Proc. Zool. Soc. London, for 1902, 2: 311, Apl. 1903; 

Ann. Mag. Nat. Hist. (7) 18: 302, Oct. 1906 (status). Gombitchu, Shoa, 

Abyssinia, 8000 ft. 

Otomys typus fortior Thomas 

Otomys typus fortior Thomas, Ann. Mag. Nat. Hist. (7) 18: 302, Oct. 1906. 

Charada, Kaffa, Abyssinia, 6000 ft. 
Otomys fortior Dollman, Ann. Mag. Nat. Hist. (8) 15: 170, Jan. 1915 (species). 

Subgenus Lamotomys Thomas 

Lamotomys Thomas, Ann. Mag. Nat. Hist. (9) 2: 208, Sep. 1918. As a subgenus 
of Otomys; type and only species Otomys laminatus Thomas and Schwann. 

Otomys laminatus laminatus Thomas and Schwann 

Otomys laminatus Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 

18, p. 23, 25 Apl. 1905; Proc. Zool. Soc. London, 1: 267, 10 Aug. 1905. 

Subudeni, Zululand, Natal. 

Otomys laminatus mariepsi Roberts 

Otomys (Lamotomys) laminatus mariepsi Roberts, Ann. Transvaal Mus., 13: 
110, 6 July 1929. Mariepskop, Lydenburg district, Transvaal. 

Otomys laminatus pondoensis Roberts 

Otomys (Lamotomys) laminatus pondoensis Roberts, Ann. Transvaal Mus., 10: 
71, 31 Jan. 1924. Ngqueleni, western Pondoland, Cape of Good Hope. 

Parotomys Thomas 
Subgenus Parotomys Thomas 

Parotomys Thomas, Ann. Mag. Nat. Hist. (9) 2: 204, Sep. 1918. Genotype 
Eur y Otis brantsii A. Smith. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 349 

Parotomys brantsii brantsii (A. Smith) 

Euryotis brantsii A. Smith, Illustr. Zool. South Africa, Mamm., pi. 24 and 

text, Mar. 1840. Interior districts of South Africa. 
Euryotis pallida Wagner, Arch. f. Naturgesch., 7: sect. 1, p. 134, 1841. South 

Africa. 
Otomys rufifrons Riippell, Mus. Senckenbergianum, 1 : 28, 1842 (nomen nudum); 

Wagner, Schreber's Saugthiere, Suppl., 3: 507, 1843. Cape of Good Hope. 

Parotomys brantsii deserti Roberts 

Parotomys brantsi deserti Roberts, Ann. Transvaal Mus., 15: 267, 1 Aug. 1933. 
Bushman Pits, Kuruman River, long. 22° E., Bechuanaland. 

Parotomys brantsii luteoltjs (Thomas and Schwann) 
Otomys brantsii luteolus Thomas and Schwann, Proc. Zool. Soc. London, 1: 
178, 9 June 1904. Deelfontein, Cape Colony. 



Subgenus Liotomys Thomas 

Liotomys Thomas, Ann. Mag. Nat. Hist. (9) 2: 204, 205, Sep. 1918. Subgenus 
of Parotomys; type Parotomys littledalei Thomas. 

Parotomys littledalei littledalei Thomas 

Parotomys (Liotomys) littledalei Thomas, Ann. Mag. Nat. Hist. (9) 2: 205, 
Sep. 1918. Tuin, Kenhart, Bushmanland, Cape of Good Hope. 

Parotomys littledalei molopensis Roberts 

Parotomys (Liotomys) littledalei molopensis Roberts, Ann. Transvaal Mus., 15: 

267, 1 Aug. 1933. Hakscheen Pan, near Rietfontein PoUce Post, west of 
Molopo River, Bechuanaland. 

Parotomys littledalei namibensis Roberts 

Parotomys (Liotomys) littledalei namibensis Roberts, Ann. Transvaal Mus., 15: 

268, 1 Aug. 1933. Swakopmund, Southwest Africa. 

Subfamily DENDROMURIN^E. Tree Mice 
Beamys Thomas 

Beamys Thomas, Ann. Mag. Nat. Hist. (8) 4: 107, Aug. 1909. Genotype 
Beamys hindei Thomas. 

Beamys hindei Thomas 

Beamys hindei Thomas, Ann. Mag. Nat. Hist. (8) 4: 108, Aug. 1909. Taveta, 
coast region, Kenya Colony, 2000 ft. 



350 bulletin: museum of compakative zoology 

Beamys major DoUman 

Beamys major Dollman, Ann. Mag. Nat. Hist. (8) 14: 428, Nov. 1914. Mlanje, 
Nyasaland. 

Dendromus a. Smith. Tree Mice 

Dendromus A. Smith, Zool. Journ., 4: 438, May 1829. Genotype Dendromus 

typus A. Smith = AfMS mesomelas Brants. 
Dendromys Smuts, Enumeratio Mamm. Capens., p. 39, 1832; substitute for 

Dendromus. 

For key to pumilio and mesomelas groups, see Wroughton, R. C, Ann. Mag. 
Nat. Hist. (8) 3: 248, 249, Mar. 1909. 

Subgenus Dendromus A. Smith 

Dendromys Thomas, Ann. Mag. Nat. Hist. (8) 18: 237, Aug. 1916. As a sub- 
genus; type Dendromus mesomelas Brants. List of species given. 

Dendromus ansorgei Thomas and Wroughton 

Dendromus ansorgei Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 16: 
173, Aug. 1905. Caconda, Benguella, Angola. 

Dendromus insignis insignis Thomas 

Dendronujs insignis Thomas, Ann. Mag. Nat. Hist. (7) 12: 341, Sep. 1903. 
Nandi, Kenya Colony. 

Dendromus insignis abyssinicus Osgood 

Dendromus insignis abyssinicus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 
235, 28 Dec. 1936. East slope of Mt. Albasso, Chilalo Mts., Arussi, Ethio- 
pia. 

Dendromus insignis kivu Thomas 

Dendromus insignis kivu Thomas, Ann. Mag. Nat. Hist. (8) 18: 242, Aug. 1916. 
Buhamba, Kivu region, Congo Beige. 

Dendromus insignis percivali Heller 

Dendromus mesamelas percivali Heller, Smithsonian Misc. Coll., 59: no. 16, p. 5, 
5 July 1912. Mt. Gargues, Mathews Range, Kenya Colony. 
A synonym of D. insignis according to Thomas (Ann. Mag. Nat. Hist. (8) 
18: 242, Aug. 1916). 

Dendromus jamesoni jamesoni Wroughton 

Dendromus jamesoni Wroughton, Ann. Mag. Nat. Hist. (8) 3: 247, Mar. 1909. 
Zoutpansberg, Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 351 

Dendromus jamesoni pongolensis Roberts 

Dendromus jamesoni pongolensis Roberts, Ann. Transvaal Mus., 14: 232, 28 

July 1931. Pongola River, 15 miles west of Manaba, northern Zululand, 

Natal. 

Dendromus leucostomus Monard 

Dendromus leucostomus Monard, Bull. Soc. Neuchateloise des Sci. Nat., 
Neuchatel, 57: 55, text-f. 7, 8 (p. 65), 1932. Caluquemb^, Angola. 

Dendromus lutstaris Osgood 

Dendromus lunaris Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 235, 28 Dec. 

1936. Kalongi, Butagu valley, west slope of Mt. Ruwenzori, central 

Africa. 

Dendromus mesomelas mesomelas (Brants) 

Mus mesomelas Brants, Het Geslacht der Muizen, p. 123, 1827. Cape of Good 
Hope, near Zondags River. 

Dendromus typus A. Smith, Zool. Journ., 4: 439, May 1829. South Africa. 

Dendromys typicus A. Smith, South African Quart. Journ., 2: 158, 1834 (emen- 
dation). 

Dendromus ayresi Roberts, Ann. Transvaal Mus., 4: 83, 20 Oct. 1913. Port St, 
John's, Cape of Good Hope. Synonym of D. mesomelas fide O. Thomas. 

Dendromus mesomelas major St. Leger 

Dendromus mesomelas major St. Leger, Ann. Mag. Nat. Hist. (10) 6: 622, Nov, 
1930. Ssanukanu Village, Grootfontein district. Southwest Africa. 

Dendromus mesomelas vernayi Hill and Carter 

Dendromus mesomelas vernayi Hill and Carter, Amer. Mus. Novitates, no. 913, 
p. 4, 8 Mar. 1937. Chitau, Angola, 4930 ft. 

Dendromus messorius Thomas 

Dendromys messorius Thomas, Ann. Mag. Nat. Hist. (7) 12: 340, Sep. 1903, 
Efulen, Cameroons. 
Range: Cameroons to northeastern Congo Beige. 

Dendromus nyas^e Thomas 

Dendromus nyasse Thomas, Ann. Mag. Nat. Hist. (8) 18: 241, Aug. 1916. 
Nyika Plateau, northern Nyasaland. 

Dendromus oreas Osgood 

Dendromus oreas Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 236, 28 Dec. 

1936. Southwest side of Mt. Cameroon, Cameroon mandate, BritisL 

Nigeria. 



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Subgenus Poemys Thomas 

Poemys Thomas, Ann. Mag. Nat. Hist. (8) 18: 238, Aug. 1916. As a subgenus 
of Dendromus; type Dendromus melanotis A. Smith. 
Regarded as a genus by G. C. Shortridge (Mamm. South West Africa, 1 : 250, 
footnote, 1934). 

Dendromus acr^us Wroughton 

Dendromus acrseus Wroughton, Ann, Mag. Nat. Hist. (8) 4: 541, Dec. 1909. 
Kirui, Mt. Elgon, Kenya Colony, 6000 ft. 

Dendromus angolensis Roberts 

Dendromus {Poemys) angolensis Roberts, Ann. Transvaal Mus., 13: 115, 6 July 
1929. Mombola, Angola. 

Dendromus arenarius Roberts 

Dendromus {Poemys) arenarius Roberts, Ann. Transvaal Mus., 10: 71, 31 Jan. 
1924. South Africa, no locality given. 

Dendromus concinnus Thomas 

Dendrovius {Poemys) concinnus Thomas, Proc. Zool. Soc. London, p. 299, 29 
Apl. 1926. Otjumbumbi, Cunene River, Southwest Africa. 

Dendromus exoneratus Thomas 

Dendromus {Poemys) exoneratus Thomas, Ann. Mag. Nat. Hist. (9) 2: 59, July 
1918. Panyam, Bauchi Province, northern Nigeria. 

Dendromus haymani Hatt 

Dendromus haymani Hatt, Amer. Mus. Novitates, no. 708, p. 13, 4 Apl. 1934. 
Mbogo, Congo Beige, 1° N., 30° E. 

Dendromus lineatus Heller 

Dendromus lineatus Heller, Smithsonian Misc. Coll., 56: no. 17, p. 4, 28 Feb. 
1911. Rhino Camp, Lado Enclave. 

Dendromus longicaudatus Roberts 

Dendromus longicaudatus Roberts, Ann. Transvaal Mus., 4: 83, 20 Oct. 1913. 
Tzaneen Estate, Transvaal. Synonym of D. melanotis fide Thomas. 

Dendromus melanotis melanotis A. Smith. 

Dendromys melanotis A. Smith, South African Quart. Journ., 2: 158, 1834. 

Port Natal, Natal. 
Dendromys subtilis Sundevall, Ofvers. Kongl. Svenska Vet.-Akad. Forhandl., 

Stockholm, 3: no. 5, pi. 120, 1846. No locality beyond South Africa. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 353 

Dendromus melanotis basxjticus Roberts 

Dendromus melanotis basuticus Roberts, Records Albany Mus., 3: 484, 31 July 
1927. Thaba Putsua Mt., Basutoland, Cape of Good Hope. 

Dendromus melanotis capensis Roberts 

Dendromus melanotis capensis Roberts, Ann. Transvaal Mus., 14: 232, 28 July 
1931. Wolseley, Cape of Good Hope. 

Dendromus melanotis chiversi Roberts 

Dendromus (Poemys) melanotis chiversi Roberts, Ann. Transvaal Mus., 13: 116, 
6 July 1929. Vlakfontein, Parys district, Orange Free State. 

Dendromus melanotis Pretoria Roberts 

Dendromus melanotis pretorige Roberts, Ann. Transvaal Mus., 14: 232, 28 July 
1931. Rietondale, Pretoria, Transvaal. 

Dendromus melanotis thorntoni Roberts 

Dendromus melanotis thorntoni Roberts, Ann. Transvaal Mus., 14: 231, 28 July 
1931. Port Elizabeth, Cape of Good Hope. 

Dendromus mystacalis Heuglin 

Dendromys mystacalis Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Dresden, 
30: art. 2, suppl., p. 5, 1863. Baschlo region, Abyssinia. 

Dendromus Nairobi Osgood 

Dendromus nairobse Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 7, 16 
Feb. 1910. Nairobi, Kenya Colony. 

Dendromus nigrifrons nigrifrons True 

Dendromys nigrifrons True, Proc. U. S. Nat. Mus., 15: 462, text-f. 2, 26 Oct. 
1892. Mt. Kilimanjaro, Tanganyika Territory. 

Dendromus nigrifrons shortridgei St. Leger 

Dendromus (Poemys) nigrifrons shortridgei St. Leger, Ann. Mag. Nat. Hist. (10) 

6: 622, Nov. 1930. Ssanukanu Village, Grootfontein district. Southwest 

Africa. 

Dendromus nigrifrons spectabilis Heller 

Dendromus spectabilis Heller, Smithsonian Misc. Coll., 56: no. 17, p. 3, 28 Feb. 
1911. Rhino Camp, Lado Enclave. 

Dendromus nigrifrons vulturnus Thomas 

Dendromus (Poemys) nigrifrons vidturnus Thomas, Ann. Mag. Nat. Hist. (8) 
18: 242, Aug. 1916. Chirinda Forest, Melsetter, Rhodesia. 



3.54 bulletin: museum of comparative zoology 

Dendromus nyik^ Wroughton 

Dendromus nyikas Wroughton, Ann. Mag. Nat. Hist. (8) 3: 248, Mar. 1909. 
Nyika Plateau, northern Nyasaland. 

Dendromus ochropus Osgood 

Dendromus ochropus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 6, 16 
Feb. 1910. Lake Elmenteita, Kenya Colony. 

Dendromus pallidus Heuglin 

Dendromys pallidus Heuglin, Reise in Nordost-Afrika, 2: 74, 1877. Middle 
Tigre River, Abyssinia (in weaver-bird nests). 

Dendromus pecilei Milne-Edwards 

Dendromys pecilei Milne-Edwards, Revue Scientifique, 38 =(3) 12: 16, 3 July 

1886. Pousargues, Ann. des Sci. Nat., zool. (8) 3: 385-391, 1896. Lower 

Congo to Ogowe River. 

Dendromus pumilio pumilio (Wagner) 

Dendromys pumilio Wagner, Gelehrte Anzeigen, Miinchen, 12: no. 54, col. 437, 
17 Mar. 1841. Cape of Good Hope. 

Dendromus ruddi Wroughton 

Dendromus ruddi Wroughton, Ann. Mag. Nat. Hist. (8) 5: 275, Mar. 1910. 
Malikisi, Mt. Elgon, Kenya Colony, 5000 ft. 

Dendromus whytei whytei Wroughton 

Dendromus whytei Wroughton, Ann. Mag. Nat. Hist. (8) 3: 247, Mar. 1909. 
Fort Hill, Nyasaland. 

Dendromus whytei capitis Heller 

Dendromus whytei capitis Heller, Smithsonian Misc. Coll., 59: no. 16, p. 6, 5 
July 1912. Mt. Lololokui, Mathews Range, Kenya Colony, 6000 ft. 

Dendromus whytei pallescens Osgood 

Dendromus whytei pallescens Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: no. 2, p. 7, 16 Feb. 1910. Lukenya Mt., Ulukenia Hills, Kenya Colony. 

Subgenus Chortomys Thomas 

Chortomys Thomas, Ann. Mag. Nat. Hist. (8) 18: 238, Aug. 1916. Subgenus of 
Dendromus, type Dendromus lovati De Winton. 

Dendromus lovati De Winton 

Dendromys lovati De Winton, Proc. Zool. Soc. London, p. 82, pi. 10, 1 June 
1900. Managasha, southern Abyssinia, 7000 ft. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 355 

Deomys Thomas 

Deomys Thomas, Proc. Zool. Soc. London, p. 130, pi. 5, 1 June 1888. Genotype 
Deomys Jerrugineus Thomas. 

Deomys ferrugineus ferrugineus Thomas 

Deotnys ferrugineus Thomas, Proc. Zool. Soc. London, p. 130, pi. 5, 1 June 1888. 
Lower Congo. 

Deomys ferrtjginetjs christyi Thomas 

Deomys christyi Thomas, Ann. Mag. Nat. Hist. (8) 16: 150, Aug. 1915. Poke, 
Uele River, Congo Beige. 

Leimacomys Matschie 

Leimacomys Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 107, 1893. Gen- 
otype Leimacomys buttneri Matschie. 
Limacomys Lydekker, Zool. Record, 30: Mamm., p. 31, 1894 (emendation). 

Leimacomys BtixxNERi Matschie 

Leimacomys biittneri Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 109, 
1893. Bismarckburg, Togo, Dahomey. 



Malacothrix Wagner 

Malacothrix Wagner, Schreber's Saugthiere, SuppL, 3: 496, 1843. Genotype 
Otomys typicus A. Smith. (See W. L. Sclater, Mammals South Africa, 2: 
34, 1901). 

Malacothrix pentonyx W. L. Sclater 

Malacothrix pentonyx W. L. Sclater, Ann. South African Mus., 1: 202, Mar. 
1899. Cape Flats near Cape Town, Cape of Good Hope. 

Malacothrix typicus typicus (A. Smith) 

Otomys typicus A. Smith, South African Quart. Journ., 2: 148, 1834. District 
east of Graaff-Remet, South Africa. 

Malacothrix typicus damarensis Roberts 

Malacothrix typicus damarensis Roberts, Ann. Transvaal Mus., 15: 10, 1 Oct. 
1932. Gobabis, Southwest Africa. * 

Malacothrix typicus egeria Thomas 

Malacothrix egeria Thomas, Proc. Zool. Soc. London, p. 301, 29 Apl. 1926. 
Ondongwa, central Ovamboland, Southwest Africa. 
Regarded as a subspecies of typicus by Roberts. 



356 bulletin: museum of comparative zoology 

Malacothrix typicus fryi Roberts 

Malacothrix typicus fryi Roberts, Ann. Transvaal Mus., 5: 268, 16 May 1917. 
Klipriviersoog, Krugersdorp district, Transvaal. 

Malacothrix typicus kalaharicus Roberts 

Malacothrix typicus kalaharicus Roberts, Ann. Transvaal Mus., 15: 10, 1 Oct. 

1932. Kuke Pan, central Kalahari, Bechuanaland. 

Malacothrix typicus molopensis Roberts 

Malacothrix typicus molopensis Roberts, Ann. Transvaal Mus., 15: 266, 1 Aug. 

1933. Eight miles west of Pitsani, Molopo River, Bechuanaland. 



Petromyscus Thomas. Rock Mice 

Petromyscus Thomas, Ann. Mag. Nat. Hist. (9) 17: 179, Jan. 1926. Genotype 
Praomys collinus Thomas and Hinton; aflBnities with Dendromyinse 
(ibid., p. 175). 

Petromyscus collinus collinus (Thomas and Hinton) 

Praomys collinus Thomas and Hinton, Proc. Zool. Soc. London, p. 237, 3 Apl. 

1925. Karibib, Southwest Africa. 

Petromyscus collinus bruchus (Thomas and Hinton) 

Praomys collinus bruchus Thomas and Hinton, p. 238, 3 Apl. 1925. Great 
Brukaros Mt., Southwest Africa. 

Petromyscus monticularis (Thomas and Hinton) 

Praom,ys monticularis Thomas and Hinton, Proc. Zool. Soc. London, p. 238, 
3 Apl. 1925. Great Brukaros Mt., near Berseba, Southwest Africa. 

Petromyscus shortridgei Thomas 

Petromyscus shortridgei Thomas, Proc. Zool. Soc. London, p. 302, 29 Apl. 

1926. Cunene Falls, northwestern Ovamboland, Southwest Africa. 



Prionomys Dollman 

Prionomys Dollman, Ann. Mag. Nat. Hist. (8) 6: 226, Aug. 1910. Genotype 
Prionomys batesi Dollman. 

Prionomys batesi Dollman 

Prionomys batesi Dollman, Ann. Mag. Nat. Hist. (8) 6: 228, Aug. 1910. 
Bitye, Ja River, Cameroons, 2000 ft. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 357 



Saccostomus Peters 

Saccostomus Peters, Ber. iiber Verb. K. Preuss. Akad. Wiss., Berlin, p. 258, 

1846. Genotype Saccostomus campestris Peters. 
Eosaccomys Palmer, Science, new ser., 17: 873, 29 May 1903, to replace 

Saccostomus Peters, supposed to be preoccupied by Saccostoma Fitzinger, 

1843, in reptiles. 

Saccostomus anderssoni anderssoni De Winton 

Saccostomus anderssoni De Winton, Ann. Mag. Nat. Hist. (7) 2: 6, footnote, 
July 1898. Damaraland. 

Saccostomus anderssoni hild^e Schwann 

Saccostomus hildx Schwann, Proc. Zool. Soc. London, p. 110, 7 June 1906i 
Kuruman, southern Bechuanaland. 

Saccostomus anderssoni pagei Thomas and Hinton 

Saccostomus pagei Thomas and Hinton, Proc. Zool. Soc. London, p. 495, 3 
Sep. 1923. Lehutitung, northern Bechuanaland. 

Saccostomus campestris Peters 

Saccostomus campestris Peters, Ber. iiber Verhandl. K. Preuss. Akad. Wiss., 

Berlin, p. 258, Aug. 1846. Tette, Mozambique. 
Saccostomus lapidarius Peters, Reise nach Mossambique, Saugeth., p. 167, pi. 

34, f. 3; pi. 35, f. 12, 1852. Sena, Tette, Macanga and Boror, Mozambique. 

Saccostomus cricetulus G. M. Allen and Lawrence 

Saccostomus cricetulus G. M. Allen and Lawrence, Bull. Mus. Comp. Zool., 79: 

100, 30 Jan. 1936. South bank of Greeki River, Sabei district, north of 

Mt. Elgon, Uganda. 

Saccostomus elegans Thomas 

Saccostomus elegans Thomas, Proc. Zool. Soc. London, p. 431, 1 Oct. 1897. 
Karonga, Lake Nyasa, northern Nyasaland. 

Saccostomus fuscus Peters 

Saccostomus fuscus Peters, Reise nach Mossambique, Saugeth., p. 168, pi. 36, 
f. 4; pi. 35, f. 13, 1852. Inhambane, Mozambique. 

Saccostomus isiol^ Heller 

Saccostomus isiolas Heller, Smithsonian Misc. Coll., 59: no. 16, p. 14, 5 July 
1912. Isiola River, Northern Guaso Nyiro, Kenya Colony. 



358 bulletin: museum of comparative zoology 

Saccostomus limpopoensis Roberts 

Saccostomus limpopoensis Roberts, Ann. Transvaal Mus., 4: 183, 22 Aug. 1914. 
Sand River, northeastern Transvaal. 

Saccostomus mashon^ De Winton 

Saccostomus mashonse De Winton, Proc. Zool. Soc. London, for 1896, p. 804, 
Apl. 1897. Mazoe, Mashonaland, Rhodesia. 

Saccostomus mearnsi Heller 

Saccostomus mearnsi Heller, Smithsonian Misc. Coll., 54: no. 1924, p. 3, pi. 2, 
f. b, 28 Feb. 1910. Changamwe, Kenya Colony. 

Saccostomus streeteri Roberts 

Saccostomus streeteri Roberts, Ann. Transvaal Mus., 4: 183, 22 Aug. 1914. 
Hector Spruit, Transvaal. 

Saccostomus umbriventer Miller 

Saccostomus umbriventer Miller, Smithsonian Misc. Coll., 54: no. 1925, p. 1, 
pi. 1, f. a, 28 Feb. 1910. Njori Osolali, Sotik, Kenya Colony. 



Steatomys Peters. Fat Mice 

Steatomys Peters, Ber. iiber Verh. K. Preuss. Akad. Wiss., Berlin, p. 258, 
Aug. 1846. Genotype Steatomys pratensis Peters. 

Steatomys angolensis angolensis Hill and Carter 

Steatomys angolensis Hill and Carter, Amer. Mus. Novitates, no. 913, p. 5, 
8 Mar. 1937. Chitau, Angola, 4930 ft. 

Steatomys angolensis bradleyi Hill and Carter 

Steatomys angolensis bradleyi Hill and Carter, Amer. Mus. Novitates, no. 913, 
p. 5, 8 Mar. 1937. Humpata, Angola, 6300 ft. 

Steatomys aquilo Thomas and Hinton 

Steatomys aquilo Thomas and Hinton, Proc. Zool. Soc. London, p. 264, 6 July 
1923. Niurmeya, Jebel Marra, Darfur, Anglo-Egyptian Sudan. 

Steatomys athi Heller 

Steatomys athi Heller, Smithsonian Misc. Coll., 54: no. 1924, p. 3, pi. 2, f.c, 
28 Feb. 1910. Lukenia Hills, Athi Plains, Kenya Colony. 

Steatomys bocagei Thomas 

Steatomys bocagei Thomas, Ann. Mag. Nat. Hist. (6) 10: 264, Sep. 1892. 
Caconda, Angola. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 359 

Steatomys cacrinus Thomas 

Steatomys caurinus Thomas, Ann. Mag. Nat. Hist. (8) 9: 271, Feb. 1912. 
Panyam Plateau, northern Nigeria. 

Steatomys chiversi chiversi Roberts 

Steatomys chiversi Roberts, Ann. Transvaal Mus., 14: 233, 28 July 1931. Blood 
River, Natal. 

Steatomys chiversi tongensis Roberts 

Steatomys chiversi tongensis Roberts, Ann. Transvaal Mus., 14: 233, 28 July 

1931. Manaba, northern Zululand, Natal. 

Steatomys cuppedius Thomas and Hinton 

Steatomys cuppedius Thomas and Hinton, Novitates Zool., 27: 318, 15 June 
1920. Farniso near Kano, northern Nigeria. 

Steatomys gazell.« Thomas and Hinton 

Steatomys gazellse Thomas and Hinton, Proc. Zool. Soc. London, p. 265, 6 July 
1923. Tamburas, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 

Steatomys jacksoni Hayman 

Steatomys jacksoni Hayman, Proc. Zool. Soc. London, for 1935, p. 930, 10 Jan. 
1936. Wenchi, Ashanti, Gold Coast. 

Steatomys krebsii krebsii Peters 

Steatomys krebsii Peters, Reise nach Mossambique, Saugeth., p. 165, pi; 36, 
f. 3, 1852. Interior of Caffraria, South Africa. 

Steatomys krebsii kalaharicus Roberts 

Steatomys krebsi kalaharicus Roberts, Ann. Transvaal Mus., 15: 11, 1 Oct. 

1932. Central Kalahari, 25 miles west of Damara Pan, Bechuanaland. 

Steatomys krebsii orangi.e Roberts 

Steatomys krebsi orangise Roberts, Ann. Transvaal Mus., 13: 116,-6 July 1929. 
"Angra Pequina," Bothaville district. Orange Free State. 

Steatomys krebsii transvaalensis Roberts 

Steatomys krebsi transvaalensis Roberts, Ann. Transvaal Mus., 13: 117, 6 
July 1929. Witfontein, Randfontein, Transvaal. 

Steatomys loveridgei Thomas ' 

Steatomys loveridgei Thomas, Ann. Mag. Nat. Hist. (9) 4: 33, July 1919. 
Lumbo, Mozambique. 



360 bulletin: museum of comparative zoology 

Steatomys minutus minutus Thomas and Wroughton 

Steatomys minutus Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 16: 
174, Aug. 1905. Fort Quillenges, Benguella, Angola. 

Steatomys minutus leucorhynchus Hill and Carter 

Steatomys minutus leucorhynchus Hill and Carter, Amer. Mus. Novitates, no. 
913, p. 4, 8 Mar. 1937. Capelongo, Huilla district, Angola. 

Steatomys muanz.e Kershaw 

Steatomys muanzse Kershaw, Ann. Mag. Nat. Hist. (9) 12: 535, Oct. 1923. 
Nyambita, Mwanza, Tanganyika Territory. 

Steatomys natalensis Roberts 

Steatomys natalensis Roberts, Ann. Transvaal Mus., 13: 117, 6 July 1929. 
Bergville, Natal. 

Steatomys opimus Pousargues 

Steatomys opimus Pousargues, Bull. Soc. Zool. de France, 19: 131, Oct. 1894. 
Near Balao, Dakoas district, 5° 26' N., 17° 40' E., Congo. 

Steatomys parvus Rhoads 

Steatomys parvus Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 529, 8 Dec. 
1896. Rusia, Lake Rudolf, Abyssinia. 

Steatomys pratensis pratensis Peters 

Steatomys pratensis Peters, Ber. liber Verb. K. Preuss. Akad. Wiss., Berlin, 

p. 258, Aug. 1846; Reise nach Mossambique, Saugeth., p. 162, pi. 34, f. 2; 

pi. 35, f. 11; pi. 36, f. 3, 1852. Tette, Mozambique. 
Steatomys edulis Peters, Reise nach Mossambique, Saugeth., p. 163, pi. 34, f. 2; 

pi. 35, f . 1 1, 1852 (substitute for S. pratensis Peters). Sena, Tette, Macanga, 

Mozambique. 

Steatomys pratensis kasaicus Hatt 

Steatomys pratensis kasaicus Hatt, Amer. Mus. Novitates, no. 708, p. 15, 4 Apl. 
1934. Luluabourg, Kasai district, Congo Beige. 

Steatomys pratensis maunensis Roberts 

Steatomys pratensis maunensis Roberts, Ann. Transvaal Mus., 15: 11, 1 Oct. 
1932. Shorobe, Maun district, Ngamiland, Bechuanaland. 

Steatomys swalius Thomas 

Steatomys swalius Thomas, Proc. Zool. Soc. London, p. 300, 29 Apl. 1926. 

Ondongwa, Ovamboland, Southwest Africa. 
Steatomys swalius umbratus Thomas, Proc. Zool. Soc. London, p. 301, 29 Apl. 
1926. Cunene Falls, northwestern Ovamboland, Southwest Africa. 
Synonym of S. stvalius fide Shortridge, G. C, Mamm. South West Africa, 
1:259, footnote, 1934. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 361 

I 

Family RHIZOMYID^E. Mole-rats 

Subfamily TACHYORYCTIN.E 

Tachyoryctes Riippell. African Mole-rats 

Tachyorydes Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 

Siiugeth., p. 35, footnote, 1835. Genotype Bathyergus splendens Riippell. 

Chrysomys Gray, List Spec. Mamm. Brit. Mus., pp. xxvi, 150, 13 May 1843. 

Genotype Bathyergus splendens Riippell. 

"When collections have been made over all parts of British East Africa and 

Abyssinia' numerous forms will doubtless be connected by complete chains of 

intergrades and the final monographer of the genus will be obliged to reduce 

many of the named forms to the rank of subspecies" (Hollister, N., Bull. U.S. 

Nat. Mus., no. 99, pt. 2, p. 40, 1919). 

Tachyoryctes ankolIvE Thomas 

Tachyoryctes ankolix Thomas, Ann. Mag. Nat. Hist. (8) 4: 545, Dec. 1909. 
Burumba, Ankole, southern Uganda. 

Tachyoryctes annectens (Thomas) 

Rhizomys annectens Thomas, Ann. Mag. Nat. Hist. (6) 7: 304, Mar. 1891. 
"Either Massai-land or inland," Kenya Colony. 

Tachyoryctes audax Thomas 

Tachyoryctes audax Thomas, Ann. Mag. Nat. Hist. (8) 5: 421, May 1910. Top 
of Aberdare Range, Kenya Colony, 10,000 ft. 
(See footnote for corrected condylobasal lengths of various forms.) 

Tachyoryctes cheesmani cheesmani Thomas 

Tachyoryctes cheesmani Thomas, Ann. Mag. Nat. Hist. (10) 1: 302, Feb. 1928. 
Dangila, Abyssinia. 

Tachyoryctes cheesmani qallarum Osgood 

Tachyoryctes cheesemani gallarum Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 
233, 28 Dec. 1936. Mt. Albasso, northeastern Chilalo Mts., Arussi, Ethio- 
pia. 

Tachyoryctes d^mon Thomas 

Tachyoryctes dasmon Thomas, Ann. Mag. Nat. Hist. (8) 4: 545, Dec. 1909. 
Mt. Kilimanjaro, Tanganyika Territory, 5000 ft. 

Tachyoryctes macrocephalus macrocephalus (Riippell) 
Rhizomys macrocephalus Riippell, Mus. Senckenbergianum, 3: 97, 115, pi. 8,^ 
f. 2; pi. 10, f. 2, 1842. Shoa, Abyssmia. 



362 bulletin: museum of compakative zoology 

Tachyoryctes macrocephalus hecki Neumann and Riimmler 
Tachyorydes macrocephalus hecki Neumann and Riimmler, Zeitschr. f. Sauge- 
tierk., 3: 302, text-f. 7-8, 11 June 1928. Abakkara, about 150 km. west 
of Lake Abassi, between Djamdjam and Arussi Galla, Abyssinia. 

Tachyoryctes naivash^ Thomas 

Tachyoryctes naivashas Thomas, Ann. Mag. Nat. Hist. (8) 4: 547, Dec. 1909. 
Lake Naivasha, Kenya Colony. 

Tachyoryctes pontifex Neumann and Riimmler 

Tachyoryctes pontifex Neumann and Riimmler, Zeitschr. f. Saugethierk., 3: 300, 
text-f. 5-6, 11 June 1928. Buka, Kaffa, Abyssinia. 

Tachyoryctes rex Heller 

Tachyoryctes rex Heller, Smithsonian Misc. Coll., 56: no. 9, p. 4, pi. 3, 4 figs., 
22 July 1910. Western slope of Mt. Kenya, Kenya Colony, 10,000 ft. 

Tachyoryctes ruand^ Lonnberg and Gyldenstolpe 

Tachyoryctes ruandse Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17 B: no. 5, 
p. 6, 2 May 1925. Mt. Muhavura, British Ruanda. 

Tachyoryctes ruddi ruddi Thomas 

Tachyoryctes ruddi Thomas, Ann. Mag. Nat. Hist. (8) 4: 546, Dec. 1909. Kirui, 
Mt. Elgon, Kenya Colony, 6000 ft. 

Tachyoryctes ruddi badius Thomas 

Tachyoryctes ruddi badius Thomas, Ann. Mag. Nat. Hist. (8) 4: 546, Dec. 1909. 
Eldoma Ravine, Kenya Colony, 7200 ft. 

Tachyoryctes spalacinus Thomas 

Tachyoryctes spalacinus Thomas, Ann. Mag. Nat. Hist. (8) 4: 547, Dec. 1909. 
Embi=Embu, near Mt. Kenya, Kenya Colony, 5400 ft. 

Tachyoryctes splendens splendens (Riippell) 

Bathyergus splendens Riippell, Neue Wirbelth. zu d. Fauna Abyssinien gehorig, 
1: Saugeth., p. 36, pi. 12, 1835. Gondar, Dembea Province, Abyssinia. 

Tachyoryctes splendens canicaudus Osgood 

Tachyoryctes splendens canicaudus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 
232, 28 Dec. 1936. Sirre, Awash valley, northern Arussi, Ethiopia. 

Tachyoryctes splendens ibeanus Thomas 

Tachyoryctes splendens ibeanus Thomas, Proc. Zool. Soc. London, p. 179, text-f. 
(skull), 1 June 1900. Machakos, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 363 

Tachyoryctes splendens omoensis Neumann and Riimmler 
Tachyorydes splendens omoensis Neumann and Rtimmler, Zeitschr. f. Siiuge- 
tierk., 3: 297, text- f. 3-4, 11 June 1928. Bolagoschana, Doko, Abyssinia. 

Tachyoryctes splendens somalicus Osgood 

Tachyoryctes splendens somalicus Osgood, Ann. Mag. Nat. Hist. (8) 5: 276, 
Mar. 1910. Ujawaj, about 100 miles southwest of Berbera, British Somali- 
land. 

Tachyoryctes storeyi Thomas 

Tachyoryctes storeyi Thomas, Ann. Mag. Nat. Hist. (8) 4: 547, Dec. 1909. Lake 
Elmenteita, Kenya Colony. 



Family SPALACID^E. Blind Mole-rats 

Subfamily SPALACIN^ 
Spalax Giildenstadt. Blind Mole-rats 

Spalax Giildenstadt, Novi Comment. Acad. Sci. Petropolitanse, 14: pt. 1, p. 

409, pis. 8, 9, 1770. Genotype Spalax microphthalmus Giildenstadt = Mus 

typhlus PaUas. 
Ommatostergus Nordmann, in Keyserling and Blasius's Wirbelth. Europas, 

pp. vii, 31, 1840. Genotype Ommatostergus pallasii Nordmann. 
Microspalax Mehely, Species Generis Spalax, p. 22, 1909. 

No genotype specified, but included four forms, one of which is S. 

xgyptiacus. Preoccupied by Microspalax Megnin and Trouessart, 1884, 

in Arachnida. 
Ujhelyiana Strand, Arch. f. Naturgesch., 88: sect, A, pt. 4, p. 142, June 1922. 

To replace Microspalax Megnin and Trouessart; ? misprint for Mehely- 

iana. 

Spalax ^qyptiacus Nehring. Egyptian Blind-rat 

Spalax segyptiacus Nehring, Sitzb. Ges. Naturf. Freunde, Berlin, p. 180, 1897. 
Ramleh, near Alexandria, Lower Egypt. 
Range: Northwestern Lower Egypt and Cyrenaica in Libya. 

Family MURID^. Typical Rats 
Subfamily MURIN.E 

For a key to the subgenera of African rats then included in Rattus, see 
Thomas, O., Ann. Mag. Nat. Hist. (8) 16: 477, Dec. 1915. 

See Thomas, O. The generic position of certain African Muridse, hitherto 
referred to Aethomys and Praomys. Ann. Mag. Nat. Hist. (9) 17: 174-179, 
Jan. 1926. 



364 bulletin: museum of comparative zoology 



AcoMYS I. Geoffrey. Spiny Mice 

Acomys I. Geoffrey, Ann. des Sci. Nat., Zool., (2) 10: 126, Aug. 1838. Geno- 
type Mus cahirinus Desmarest. 

Acanthomys Lesson, Nouv. Tabl. Regne Animal, Mamm., p. 135, 1842. Geno- 
type not specified; included five species, one of them an Acomys, of which 
it is a part synonym. 

See Dollman, Guy. On the Spiny Mice of British East Africa, with a des- 
cription of a new species from Magadi. Ann. Mag. Nat. Hist. (8) 14: 485-489, 
Dec. 1914 (key). 

Acomys airensis Thomas and Hinton 

Acomys airensis Thomas and Hinton, Novitates Zool., 28: 8, 14 May 1921; 
Mt. Baguezan, Asben, northern Nigeria. 

Acomys albigena Heuglin 

Acomys albigena Heuglin, Reise in Nordost-Afrika, 2: 69, 1877. Bogos country, 
Abyssinia. 

Acomys bovonei de Beaux 

Acomys bovonei de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. Nat., 

Milano, 73: 280, pi. 8, f. A.b; pi. 9, f. A.b, Dec. 1934. El Bur, southern 

Italian Somaliland. 

Acomys brockmani Dollman 

Acomys brockmani Dollman, Ann. Mag. Nat. Hist. (8) 8: 259, Aug. 1911. 
Bulbar, British Somaliland. 

Acomys cahirinus (Desmarest). Cairo Spiny Mouse 

Mus cahirinus Desmarest, Nouv. Dictionnaire d'Hist. Nat., 29: 70, 1819. 

Cairo, Egypt. 
Acomys sabryi Kershaw, Ann. Mag. Nat. Hist. (9) 10: 99, 107, July 1922. 

Helwan, 10 miles south of Cairo, Egypt. Probably a synonym of A. 

cahirinus (see Flower, S. S., Proc. Zool. Soc. London, p. 414, 8 July 1932). 

Acomys chudeaui Kollman 

Acomys chudeaui Kollman, Bull. Mus. d'Hist. Nat., Paris, 17: 402, 1911. Atar, 
southwest of Biskra, Mauretania. 

Acomys cineraceus Fitzinger and Heuglin 

Acomys cineraceus Fitzinger and Heuglin, Sitzb. K. Akad. Wiss. Wien, 54: 

sect. 1, p. 573, 1866. Doka, eastern Sennaar, between the Atbara and 

Rahad Rivers, Anglo-Egyptian Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 365 

Acomys cinerascens Heuglin, Reise in Nordost-Afrika, 2: 70, 1877. Eastern 

Sennaar and about Kalabat, Anglo-Egyptian Sudan. 
Acomys witherbyi De Winton, Novitates Zool., 8: 400, 31 Dec. 1901. El Kowa, 

south of Khartoum, Anglo-Egyptian Sudan. 

Acomys dimidiatus (Cretzschmar) 

Mus dimidiatus Cretzschmar, in Riippell's Atlas zu d. Reise im Nordliche 

Afrika, p. 37, pi. 13, f.a, 1826. Sinai region and Nubia. 
Mus megalotis Lichtenstein, Darstellung. neuer oder wenig bekannt. Saugeth., 

pi. 37, f.2, and text (unnumbered pp.), 1829. Arabia. 

Acomys hawashensis Frick 

Acomys hawashensis Frick, Ann. Carnegie Mus., 9: 26, 6 June 1914. Sadi 
Malka, Abyssinia, 3500 ft. 

Acomys hunteri De Winton 

Acomys hunteri De Winton, Novitates Zool., 8: 401, footnote, 31 Dec. 1900. 
Plain of Tokar, Suakin, Anglo-Egyptian Sudan. 

Acomys hystrella Heller 

Acomys hystrella Heller, Smithsonian Misc. Coll., 56: no. 17, p. 13, 28 Feb. 
1911. Nimule, Uganda. 

Acomys ignitus ignitus Dollman 

Acomys ignitus Dollman, Ann. Mag. Nat. Hist. (8) 6: 229, Aug. 1910. Voi, 
Kenya Colony. 

Acomys ignitus kempi Dollman 

Acomys kempi Dollman, Ann. Mag. Nat. Hist. (8) 8: 125, July 1911. Chanler 
Falls, Northern Guaso Nyiro, Kenya Colony. 

Acomys pulchellus Dollman, Ann. Mag. Nat. Hist. (8) 8: 127, July 1911. 
Chanler Falls, Northern Guaso Nyiro, Kenya Colony. Based on immature 
example (see HoUister, N., Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 117, 1919). 

Acomys ignitus montanus Heller . 

Acomys ignitis (sic) montanus Heller, Smithsonian Misc. Coll., 63: no. 7, p. 
12, 24 June 1914. North slope of Mt. Marsabit, Kenya Colony, 4600 ft. 

Acomys intermedius Wettstein 

Acomys intermedius Wettstein, Anzeiger K. Akad. Wiss. Wien, math.-nat. Kl., 
53: 161, 1916. Dilling, southern Kordofan, Anglo-Egyptian Sudan. 



366 bulletin: museum of comparative zoology 

AcoMYS JOHANNis Thomas 

Acomys johannis Thomas, Ann. Mag. Nat. Hist. (8) 9: 272, Feb. 1912. Kabwir, 
northern Nigeria. 

Acomys louis^ LOUisiE Thomas 

Acomys louisse Thomas, Ann. Mag. Nat. Hist. (6) 18: 269, Sep. 1896. Hen- 
waina plain, 40 miles south of Berbera, Somaliland. 

Acomys louis^ tjmbratus Thomas 

Acomys louisae umbratus Thomas, Ann. Mag. Nat. Hist. (9) 12: 174, July 1923. 
Wagar Mountain, Golis Mts., northern Somaliland. 

Acomys mullah Thomas 

Acomys mullah Thomas, Ann. Mag. Nat. Hist. (7) 14: 103, Aug. 1904. Harar, 
British Somaliland. 

Acomys nubicus Heuglin 

Acomys nubicus Heuglin, Reise in Nordost-Afrika, 2: 70, 1877. De Winton, 
Novitates Zool., 8: 401, 31 Dec. 1901. Nubia, along the Nile. 

Acomys percivali Dollman 

Acomys percivali Dollman, Ann. Mag. Nat. Hist. (8) 8: 126, July 1911. Chanler 
Falls, Northern Guaso Nyiro, Kenya Colony. 

Acomys rtjssatus ^gyptiacus Bonhote 

Acomys russatus segyptiacus Bonhote, Abstr. Proc. Zool. Soc. London, no. 103, 

p. 3, 13 Feb. 1912; Proc. Zool. Soc. London, p. 230, 13 Mar. 1912. Wadi 

Hof, near Helwan, Egypt. 
Mus affinis Gray, List Spec. Mamm. Brit. Mus., p. 108, 13 May 1843 (nomen 

nudum). Egypt. 
As to the history of the specimen on which this name is based, see Flower, 
S. S., Proc. Zool. Soc. London, p. 412, 8 July 1932. 

Acomys selousi De Winton 

Acomys selousi De Winton, Proc. Zoo). Soc. London, for 1896, p. 807, pi. 40, 

f. 2, Apl. 1897. Essex Farm, Matabeleland, near Bulawayo, Southern 

Rhodesia. 

Acomys seurati Heim de Balsac 

Acomys seurati Heim de Balsac, Suppl. au Bull. Biol, de France et de Belgique, 

Paris, 21: 356, f. 6, no. 4 (foot); 389, f. 15, no. 15 (skuU), 1936; Bull. Soc. 

Zool. de France, 62: no. 5, 332, 14 Aug. 1937 (formal description). Iniker, 

A'haggar, southern Algeria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 3G7 

AcoMYS spiNosissiMus Peters 

Acomys spinosissimus Peters, Reise nach Mossambique, Saugeth., p. 160, pi. 
34, f. 1; pi. 35, f. 10, 1852. Tette and Buio, Mozambique. 

Acomys subspinosxjs (Waterhouse) 

Mus subspinosus Waterhouse, Proc. Zool. Soc. London, for 1837, p. 104, 25 
May 1838. Cape of Good Hope. 

Acomys transvaalensis Roberts 

Acomys transvaalensis Roberts, Ann. Transvaal Mus., 11: 252, 14 Sep. 1926. 
Newgate, ^outpansberg, Transvaal. 

Acomys viator Thomas 

Acomys viator Thomas, Proc. Zool. Soc. London, 2: 10, Oct. 1902. Wadi 
Sultan, near Sokna, Tripoli. 

Acomys wilsoni wilsoni Thomas 

Acomys wilsoni Thomas, Ann. Mag. Nat. Hist. (6) 10: 22, Julj' 1892. Mombasa, 
Kenya Colony. 

Acomys wilsoni ablutus Dollman 

Acomys ablutus Dollman, Ann. Mag. Nat. Hist. (8) 8: 127, July 1911. Nyama 
Nyango, Northern Guaso Nyiro, Kenya Colon}-. 

Acomys wilsoni argillaceus Hinton and Kershaw 

Acomys wilsoni argillaceus Hinton and Kershaw, Ann. Mag. Nat. Hist. (9) 6: 
101, July 1920. Mongalla, Bahr-el-Djebel, Anglo-Egyptian Sudan. 

Acomys wilsoni enid St. Leger 

Acomys wilsoni enid St. Leger, Ann. Mag. Nat. Hist. (10) 9: 241, Mar. 1932. 
Koliokwell River, Lake Rudolf, Uganda. 

Acomys wilsoni nubilus Dollman 

Acomys nubilus Dollman, Ann. Mag. Nat. Hist. (8) 14: 486, Dec. 1914. Mag- 
adi, southern Masailand district, Kenya Colony. 



.Ethomys Thomas 

Mthomys Thomas, Ann. Mag. Nat. Hist. (8) 16: 477, Dec. 1915. As a sub- 
genus of Epimys { = Rattus), type Epiniys hindei Thomas. Raised to gen- 
eric rank; list of included species given, see Thomas, O., Ann. Mag. Nat. 
Hist. (9) 17: 177, Jan. 1926. 



368 bulletin: museum of comparative zoology 

iExHOMYS AVARILLUS (Thomas and Wroughton) 

Mus avarillus Thomas and Wroughton, Proc. Zool. Soc. London, p. 547, 30 
Oct. 1908. Tette, Mozambique. 

iExHOMYS BOCAGEi (Thomas) 

Mus bocagei Thomas, Ann. Mag. Nat. Hist. (7) 13: 416, June 1904. Pungo 

Andongo, Angola. 
jEthomys bocagei Thomas, Ann. Mag. Nat. Hist. (9) 17: 177, Jan. 1926. 

iETHOMYS CHRYSOPHiLus CHRYSOPHiLus (De Winton) 

Mus chrysophilus De Winton, Proc. Zool. Soc. London, for 1896, p. 801, Apl. 
1897. Mazoe, Mashonaland, Southern Rhodesia. 

J^^THOMYS CHRYSOPHILUS ACTicoLA (Thomas and Wroughton) 

Mus chrysophilus acticola Thomas and Wroughton, Proc. Zool. Soc. London, p. 

547, 30 Oct. 1908. Beira, Mozambique. 
Mus chrysophilus alticola Lydekker, Intern. Cat. Sci. Literature, 8 (1908): 

Mamm., p. 74, Jan. 1910. Accidental renaming of M. c. acticola Thomas 

and Wroughton. 

JiItHOMYS CHRYSOPHILUS ALBIVENTER (Jeutiuk) 

Mus albiventer Jentink, Zool. Jahrb., Syst., 28: 246, text-f. H, 15 Oct. 1909. 

Mossel Bay, Cape of Good Hope. 
Mus albiventris Jentink, Zool. Jahrb., Syst., 28: 247, 15 Oct. 1909. Used inad- 
vertently in place of M. albiventer Jentink. 
M[us] muscardinus Wagner, Schreber's Saugthiere, Suppl., 3: 430, 1843. Kaffir- 
land, = South Africa. 
Possibly this name should supersede chrysophilus, but is at present undeter- 
minable. 

J^ItHOMYS CHRYSOPHILUS CAPRICORNIS RobcrtS 

jEthortiys chrysophilus capricornis Roberts, Ann. Transvaal Mus., 11: 254, 14 
Sep. 1926. Newgate, Zoutpansberg, Transvaal. 

iETHOMYS CHRYSOPHILUS DOLLMANI Hatt 

jEthomys chrysophilus dollmani Hatt, Amer. Mus. Novitates, no. 708, p. 8, 
4 Apl. 1934. Upper Lufira River, Katanga, Congo Beige. 

^THOMYS CHRYSOPHILUS IMAGO ThomaS 

^thomys chrysophilus imago Thomas, Proc. Zool. Soc. London, p. 387, 12 July 
1927. Stampriet, Southwest Africa. 

^Ethomys CHRYSOPHILUS MAGALAKuiNi Roberts 

^Ihomys chrysophilus magalakuini Roberts, Ann. Transvaal Mus., 11: 254, 14 
Sep. 1926. WUhanshohe, Magalakuin, Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 369 

^THOMYS CHRYsoPHiLUS INEPTUS (Thomas and Wroughton) 
Mus chrysophilus inept us Thomas and Wroughton, Proc. Zool. See. London, p. 
546, 30 Oct. 1908. Tette, Mozambique. 

iETHOMYS CHRYSOPHILUS PRETORI.E (Roberts) 

Mus chrysophilus pretorise Roberts, Ann. Transvaal Mus., 4: 85, 20 Oct. 1913. 
Fountains, Pretoria, Transvaal. 

jEthomys chrysophilus singid^ (Kershaw) 

Rattus {/Ethomys) chrysophilus singidse Kershaw, Ann. Mag. Nat. Hist. (9) 12: 

535, Oct. 1923. Gwao's, near Itigi, Singida, Tanganyika Territory, about 

30° 40' E., 4° 25' S. 

.Ethomys chrysophilus tongensis Roberts 

Ethomys chrysophilus tongensis Roberts, Ann. Transvaal Mus., 14: 235, 28 

July 1931. Mangusi Forest, northern Zululand, Natal. 
JEthomys chrysophilus tongensis Roberts, Ann. Transvaal Mus., 18: 235, 21 

Feb. 1936. Otobotini, northern Zululand, Natal. 
Not M thorny s chrysophilus tongensis Roberts, 1931? 

^Ethomys chrysophilus tzaneenensis (Jameson) 

Mus chrysophilus tzaneenensis Jameson, Ann. Mag. Nat. Hist. (8) 4 : 460, Nov. 
1909. Tzaneen, Zoutpansberg district, Transvaal. 

-Ethomys chrysophilus voi (Osgood) 

Mus voi Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 11, 16 Feb. 1910. 
Voi, Kenya Colony. 

.(Ethomys kaiseri kaiseri (Noack) 

Epivnjs kaiseri Noack, Zool. Jahrb., Syst., 2: 228, pi. 9, f. 1-3, 7 May 1887. 
Marungu, (Qua Mpala), Tanganyika Territory. 

.(Ethomys kaiseri alghazal (Wroughton) 

Mus alghazal Wroughton, Ann. Mag. Nat. Hist. (7) 20: 501, Dec. 1907. Chak 
Chak, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 

.(Ethomys kaiseri helleri (Hollister) 

Epimys kaiseri centralis Heller, Smithsonian Misc. Coll., 63: no. 7, p. 10, 24 
June 1914. Rhino Camp, Lado Enclave. Doubtfully distinct (Thomas). 

Rattus helleri Hollister, Proc. Biol. Soc. Washington, 31 : 97, 29 June 1918 (new 
name to replace Epimys kaiseri centralis Heller, not Mus auricomis cen- 
tralis Schwann, 1906). 



370 bulletin: museum of comparative zoology 

^THOMYS KAISERI HINDEI (Thomas) 

Mus hindei Thomas, Ann. Mag. Nat. Hist. (7) 9: 219, Mar. 1902. Machakos, 
Kenya Colony. 

^THOMYS KAISERI MANTEUFELI (Matschie) 

Mus (Epimys?) manteufeli Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 
341, Oct. 1911. Mwanza, Lake Victoria, Tanganyika Territory. 

.^THOMYS KAISERI MEDICATtJS (WrOUghton) 

Mus medicatus Wroughton, Ann. Mag. Nat. Hist. (8) 4: 540, Dec. 1909. 
Mumia's, Kenya Colony. 

jEthomys KAISERI NOR^ (Wroughton) 

Mus norse Wroughton, Ann. Mag. Nat. Hist. (8) 4: 541, Dec. 1909. Guaso 
Narok, Northern Guaso Nyiro, Kenya Colony. 

iExHOMYS KAISERI TURNERI (Heller) 

Epimys kaiseri turneri Heller, Smithsonian Misc. Coll., 63: no. 7, p. 8, 24 June 
1914. Kisumu, Kenya Colony. 
Range: probably confined to shore area of Lake Victoria. 

iExHOMYS NAMAQUENSIS NAMAQUENSIS (A. Smith) 

Gerbillus namaquensis A. Smith, South African Quart. Journ., 2: 160, 1834. 

Namaqualand, Cape of Good Hope. 
Otoniys namaquensis y ar. fusca F. Cuvier, Hist. Nat. des Mammifdres, pt. 61, 
1829. 
For key to various forms of ^. namaquensis see Jameson, H. L., Ann. Mag. 
Nat. Hist. (8) 4: 462, Nov. 1909. 

^THOMYS NAMAQUENSIS ARBORARItJS (Peters) 

Mus arborarius Peters, Reise nach Mossambique, Saugeth., p. 152, pi. 35, f. 7; 
pi. 36, f. 2, 1852. Tette, Mozambique. 
Probably a distinct race rather than a synonym of ^. namaquensis nam,a- 
quensis (see Thomas and Wroughton, Proc. Zool. Soc. London, p. 548, 30 Oct. 
1908). 

.^THOMYS NAMAQUENSIS AURICOMIS (De Winton) 

Mus auricomis De Winton, Proc. Zool. Soc. London, for 1896, pp. 802, 807, 
Apl. 1897. Mazoe, Mashonaland, Southern Rhodesia. 

Mus namaquensis auricomis Thomas and Wroughton, Proc. Zool. Soc. London, 
p. 548, 30 Oct. 1908. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 371 

^THOMYS NAMAQUENSIS CALARIUS ThomaS 

Mthoimjs namaquensis calarius Thomas, Ann. Mag. Nat. Hist. (9) 17: 184, 
Jan. 1926. Lehutitung, Ngamiland, Bechuanaland. 

iETHOMYS NAMAQUENSIS CAPENSIS (Robei'ts) 

Praomys namaquensis capensis Roberts, Ann. Transvaal Mus., 11: 254, 14 Sep. 
1926. Lormarins, Paarl, Cape of Good Hope. 

.^THOMYS NAMAQUENSIS CENTRALIS (Schwann) 

Mus auricotnis centralis Schwann, Proc. Zool. Soc. London, 1 : 107, 7 June 1906. 

Deelfontein, Cape of Good Hope. 
Mus na7naquensis centralis Thomas and Wroughton, Proc. Zool. Soc. London, 

p. 548, 30 Oct. 1908. 

.^THOMYS NAMAQUENSIS DRAKENSBERGI (Roberts) 

Praomys namaquensis drakensbergi Roberts, Ann. Transvaal Mus., 11: 255, 
14 Sep. 1926. Klipspruit, Utrecht, Natal. 

uEthomys NAMAQUENSIS GRAHAMi (Roberts) 

Mus namaquensis grahami Roberts, Ann. Transvaal Mus., 5: 118, 28 Aug. 1915. 
Godwin's Kloof, Grahamstown, Cape of Good Hope. 

.^THOMYS NAMAQUENSIS KLAVERENSIS (Roberts) 

Praomys namaquensis klaverensis Roberts, Ann. Transvaal Mus., 11: 254, 14 
Sep. 1926. Klaver, Cape of Good Hope. 

iExHOMYS NAMAQUENSIS LEHOCLA (A. Smith) 

Mus lehocla A. Smith, Rept. Exped. for Exploring South Africa, App., p. 43, 

1836. Latakoo, South Africa. 
Mus namaquensis lehocla Thomas and Wroughton, Proc. Zool. Soc. London, 

p. 548, 30 Oct. 1908. 

^THOMYS NAMAQUENSIS LEHOCHLOIDES (Roberts) 

Praomys namaquensis lehochloides Roberts, Ann. Transvaal Mus., 11: 255, 
14 Sep. 1926. Wilhanshohe, Magalakuin, Transvaal. 

^THOMYS NAMAQUENSIS MONTICULARIS (JameSOn) 

Mus namaquensis monticularis Jameson, Ann. Mag. Nat. Hist. (8) 4: 461, 
Nov. 1909. Johannesburg, Transvaal. 

^THOMYS NAMAQUENSIS SICCATUS ThomaS 

Mthomys namaquensis siccatus Thomas, Proc. Zool. Soc. London, p. 304, 29 Apl. 
1926. Cunene Falls, Ovamboland, Southwest Africa. 



372 bulletin: museum of comparative zoology 

iExHOMYS NYiK^ (Thomas) 

Mus nyikx Thomas, Proc. Zool. Soc. London, p. 431, 1 Oct. 1897. Nyika plat- 
eau, northern Nyasaland. 

^THOMYS P^DULCUS (Sundevall) 

Mus -psedulcus Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. Forhandl., 

Stockholm, for 1846, 3: no. 4, p. 120, 1847. "In Caffraria interiore, prope 

tropicum." 

.^THOMYS STANNARIUS (Thomas) 

E-pimys stannarius Thomas, Ann. Mag. Nat. Hist. (8) 11: 482, May 1913. 

Kabwir, Bauchi Province, northern Nigeria. 
Mthomys stannarius Thomas, Ann. Mag. Nat. Hist. (9) 17: 177, Jan. 1926. 

^THOMYS VERNAYi Hill and Carter 

Mthomys vernayi Hill and Carter, Amer. Mus. Novitates, no. 913, p. 1, f. 1, 
8 Mar. 1937. Chissonque, 20 km. east of Dande, Angola. 

.^THOMYS WALAMB.E WALAMB^ (WrOUghtou) 

Mus walamhse Wroughton, Mem. and Proc. Manchester Lit. and Phil. Soc, 51 : 
no. 5, p. 21, 13 Mar. 1907. Msofu River, Rhodesia. 

^Ethomys walamb^ amal^ (Dollman) 

Epimys walamhx amalas Dollman, Abstr. Proc. Zool. Soc. London, no. 131, 
p. 25, 14 Apl. 1914; Proc. Zool. Soc. London, p. 313, 16 June 1914. Lemek 
Valley, between the Amala River and Southern Guaso Nyiro, Kenya 
Colony. 

.^Ethomys walamb^ hintoni Hatt 

JEthomys walamhse hintoni Hatt, Amer. Mus. Novitates, no. 708, p. 7, 4 Apl. 
1934. Kambove, Katanga, Congo Beige, 4400 ft. 

iExHOMYS WALAMB^ PEDESTER (Thomas) 

Epimys ivalambse pedester Thomas, Ann. Mag. Nat. Hist. (8) 8: 376, Sep. 1911. 
Kigezi, Mfumbiro region, southwestern Uganda. 



Apodemus Kaup. Wood Mice 

Apodemus Kaup, Skizzirte Entw.-gesch. u. Naturl. Syst. Europ. Thierwelt, 1: 
154, 1829. Genotype Mus agrarius Pallas. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 373 



Subgenus Sylv^mus Ognev 

Sylvsemus Ognev, Fauna Zemn. Pozvonocn. Woronesch, 1923. Type Mus sylva- 

ticus Linnaeus. 
Nemomys Thomas, Journ. Bombay Nat. Hist. Soc, 29: 889, 15 May 1924. 

Subgenus of Apodemus, type Mus sylvaticus Linnaeus. 

Apodemus sylvaticus ALGIRT7S (Pomel). Algerian Wood Mouse 

Mus algirus Pomel, Compt. Rend. Acad. Sci. Paris, 42: 654, 1856. Algeria, 

Or an Province. 
Probably a valid subspecies (Cabrera, A., Trab. d. Mus. Nac. de Cienc. Nat., 
Madrid, zool. ser., no. 57, p. 253, 30 Dec. 1932). 
Mtis chamseropsis Levaillant, Exploration Sci. de I'Algerie, Zool., Mammiferes, 

Atlas, pi. 5, f. 1 (col.), la, 1857; Loche, Cat. Mamm. et Oiseaux observes 

en Algerie, p. 25, 1858. Constantine Province, Algeria. 
Although identified as a Gerbillus by Lataste (Actes Soc. Linn, de Bordeaux, 
37: 14, 1883), Thomas points out that the ungrooved incisors indicate Mus. 
Mus reboudia Loche, Exploration Sci. de I'Algerie, Zool., Mammiferes, p. 117, 

1867. Oasis of Messad, Algeria. 

Apodemus sylvaticus hayi (Waterhouse) 

Mus hayi Waterhouse, Proc. Zool. Soc. London, for 1837, p. 76, July 1838. 

(Tangiers), Morocco. 
Apodemus sylvaticus dichrurus Cabrera, Mem. Real Soc. Espafiola Hist. 
Nat., Madrid, 50th anniv. vol., p. 44, 1921 (not of Rafinesque). 
Range: Northern Morocco. 

Arvicanthis Lesson. Kusu 

Arvicanthis Lesson, Nouv. Tableau Regne Anim., Mamm., p. 147, 1842. 

Thomas, Proc. Zool. Soc. London, p. 553, footnote, Oct. 1895. Genotype 

Lemmus niloticus E. Geoffrey. 
Isomys Sundevall, K. Svenska Vet.-Akad. Handl., Stockholm, for 1842, p. 219, 

1843. Genotype Mxis variegatus Brants. 
For diagnostic characters, see Thomas O., Ann. Mag. Nat. Hist. (8) 18: 68, 
July 1916. 

See Dollman, Guy. On Arvicanthis abyssinicus and alUed East-African 
species, with descriptions of four new forms. Ann. Mag. Nat. Hist. (8) 8: 
334-353, Sep. 1911. 

Arvicanthis abyssinicus abyssinicus (Riippell) 

Mus abyssinicus Riippell, Mus. Senckenbergianum, 3: 104, pi. 7, f. 1, 1842. 
Entschetqab, Simen province, Abyssinia. 



374 bulletin: museum of comparative zoology 

Mus (?) rufidorsalis Heuglin, Reise in Nordost-Afrika, 2: 70, 1877. Grassy 
plains in Simen and Wogara, Abyssinia. 
"Probably ... a synonym of A. abyssinicus" (Thomas, O., Ann. Mag. Nat. 
Hist. (10) 1: 304, Feb. 1928). 

Ahvicanthis abyssinicus blicki Flick 

Arvicanthis abyssinicus blicki Frick, Ann. Carnegie Mus., 9: 20, pi. 4, f. 1-5, 
6 June 1914. Hora Mt. base camp, 9000 ft.. South Chilalo Mts., Abys- 
sinia. 

Arvicanthis abyssinicus centrosus HolUster 

Arvicarithis abyssinicus centrosus Hollister, Smithsonian Misc. Coll., 66: no. 
10, p. 1, 26 Oct. 1916. Rhino Camp, Lado Enclave. 

Arvicanthis abyssinicus fluvicinctus Osgood 

Arvicanthis abyssinicus fluvicinctus Osgood, Zool. Ser. Field Mus. Nat. Hist., 
20: 251, 28 Dec. 1936. Bichana, Gojjam, Ethiopia. 

Arvicanthis abyssinicus mearnsi Frick 

Arvicanthis abyssinicus mearnsi Frick, Ann. Carnegie Mus., 9: 22, pi. 4, f. 
6-10, 6 June 1914. Sadi Malka, Hawash River, Abyssinia, 2800 ft. 

Arvicanthis abyssinicus muans^ Matschie 

Arvicanthis abyssinicus muansx Matschie, Sitzb. - Ges. Naturf. Freunde, 
Berlin, p. 339, Oct. 1911. Mwanza, south coast of Lake Victoria, Tangan- 
yika Territory. 

Arvicanthis abyssinicus nairob.e J. A. Allen 

Arvicanthis nairobse J. A. Allen, Bull. Amer. Mus. Nat. Hist., 26: 168, 19 Mar. 
1909. Nairobi, Kenya Colony. 

Arvicanthis abyssinicus nubilans Wroughton 

Arvicanthus (sic) abyssinicus nubilans Wroughton, Ann. Mag. Nat. Hist. (8) 
4: 539, Dec. 1909. Kisumu, Kenya Colony. 

Arvicanthis abyssinicus pallescens Dollman 

Arvicanthis rumruti •pallescens Dollman, Abstr. Proc. Zool. Soc. London, no. 

131, p. 25, 14 Apl. 1914; Proc. Zool. Soc. London, p. 316, 16 June 1914. 

Loita Plains, Kenya Colony. 
Arvicanthis abyssinicus pallescens Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 

2, p. 130, 1919. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 375 

Arvicanthis abyssinicus pr^ceps Wroughton 

Arvicanthis abyssinicus praeceps Wroughton, Ann. Mag. Nat. Hist. (8) 4: 
538, Dec. 1909. Naivasha, Kenya Colony. 

Arvicanthis abyssinicus raffertyi Frick 

Arvicanthis abyssinicus raffertyi Frick, Ann. Carnegie Mus., 9: 23, pi. 5, 6 June 
1914. Gardula, southern Abyssinia. 

Arvicanthis abyssinicus rossii de Beaux 

Arvicanthis abyssinicus rossii de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. 
Stor. Nat., Milano, 64: 90, Sep. 1925. AtaUa, Congo Beige. 

Arvicanthis abyssinicus rubescens Wroughton 

Arvicanthis abyssinicus rubescens Wroughton, Ann. Mag. Nat. Hist. (8) 4: 
538, Dec. 1909. Kibero, Unyoro, Uganda. 

Arvicanthis abyssinicus rumruti Dollman 

Arvicanthis rumruti Dollman, Ann. Mag. Nat. Hist. (8) 8: 128, July 1911. 
Rumruti, Laikipia Plateau, Kenya Colony. 

Arvicanthis abyssinicus saturatus Dollman 

Arvicanthis abyssinicus saturatus Dollman, Ann. Mag. Nat. Hist. (8) 8: 343, 
Sep. 1911. Didessa River, near Guma, Abyssinia. 

Arvicanthis abyssinicus virescens Heller 

Arvicanthis abyssinicus virescens Heller, Smithsonian Misc. Coll., 63: no. 7, 
p. 11, 24 June 1914. Vol, Kenya Colony. 

Arvicanthis abyssinicus zaphiri Dollman 

Arvicanthis zaphiri Dollman, Ann. Mag. Nat. Hist. (8) 8: 349, Sep. 1911. 
Guma, southern Abyssinia. 

Arvicanthis lacernatus (Riippell) 

Meriones lacernatus Riippell, Mus. Senckenbergianum, 3: 96, 115, pi. 6, f. 1, 

1842. Grassy plains about Lake Dembea (=Lake Tana), Abyssinia 

(see Thomas, Proc. Zool. Soc. London, for 1902, 2: 312, Apl. 1903; and 

Ann. Mag. Nat. Hist. (7) 8: 274, Oct. 1901). 
Mus dembeensis Riippell, Mus. Senckenbergianum, 3: 109, pi. 6, f. 3, 1842. 

Deraske, Lake Tana, Abyssinia? 
"I do not now think that Rtippell's Mus dembeensis is a Desmomys at all, 
as was formerly supposed to be the case. The 'straffen haaren', 'schmutzig- 
aschgrau' under surface and large claws ... do not agree by any means with 
Desmomys, but apply well to an Arvicanthis, to which genus I have little 
doubt dcmbeeyisis is referable" (Thomas, O., Ann. Mag. Nat. Hist. (10) 1: 
302, Feb. 1928). 



376 bulletin: museum of comparative zoology 

Aridcanthis abyssinicus pelliceus Thomas, Ann. Mag. Nat. Hist. (10) 1 : 303, 
Feb. 1928. Zauday Grar, south shore of Lake Tana, Abyssinia. 
Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 253, 28 Dec. 1936) shows that 
this is a synonym of A . lacer?iatus. 

Arvicanthis niloticus niloticus (Desmarest). Nile Kusu 

Arvicola niloticus Desmarest, Encyclop. Methodique, Mammalogie, pt. 2, 

p. 281, 1822. Egypt. 
Hypiidseus variegatus Lichtenstein, Verzeichn. d. Doubletten d. Zool. Mus. 

Berlin, p. 2, 1823. "E provincia Fajum Aegypt." 
Echimys niloticus E. Geoflfroy and Audouin, Descr. de I'Egypte, 2: 734, pi. 5, 

f.2, 1829. Egypt, banks of the Nile. 
Mus discolor Wagner, Arch. f. Naturg., 8: sect. 1, p. 9, 1842. Lower Egypt. 
Isomys variegatus, vars. major et minor Sundevall, Kongl. Svenska Vet.- 

Akad. Handl., Stockholm, for 1842, p. 220, 1843. Syrkut, upper Nubia. 
Relationship to A. abyssinicus and A. lacernatus still uncertain (Thomas, 
O., Ann. Mag. Nat. Hist. (7) 8: 274, Sep. 1901). 

Range : Upper Egypt, the Fayum, Giza area, and parts of the Nile delta. 

Arvicanthis niloticus ansorgei Thomas 

Arvicanthis ansorgei Thomas, Ann. Mag. Nat. Hist. (8) 5: 353, Apl. 1910. 
Gunnal, Portuguese Guinea. 

Arvicanthis niloticus centralis Dollman 

Arvicanthis testicularis centralis Dollman, Ann. Mag. Nat. Hist. (8) 8: 338, 

Sep. 1911. Between Chak Chak and Dem Zubeir, western Bahr-el- 

Ghazal, Anglo-Egyptian Sudan. 
According to Brsestrup (Vidensk. Medd. fra Dansk Naturh. Foren., Kjoben- 

havn, 99: 119, 11 Sep. 1935) this is "apparently not distinct from rufimis." 

Arvicanthis niloticus jebel^ Heller 

Arvicanthis jebelse Heller, Smithsonian Misc. Coll., 56: no. 17, p. 9, 28 Feb. 

1911. Rhino Camp, Lado Enclave. 
Arvicanthis testicularis jebelae Dollman, Ann. Mag. Nat. Hist. (8) 8: 338, Sep. 

1911. 
Probably a synonym of A.n. testicularis (Brsstrup, F. W., Vidensk. Medd. 
fra Dansk Naturh. Foren., Kjobenhavn, 99: 119, 11 Sep. 1935). 

Arvicanthis niloticus kordofanensis Wettstein 

Arvicanthis testicularis kordofanensis Wettstein, Anzeiger K. Akad. Wiss. Wien, 
math.-nat. KL, 53: 161, 1916. Kadugli, southern Kordofan, Anglo- 
Egyptian Sudan. 
A sjTionym of ^. w. testicularis (Brajstrup, in Vidensk. Medd. fra Dansk 

Naturh. Foren., Kjobenhavn, 99: 119, 11 Sep. 1935). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 377 

Arvicanthis niloticus ltjctuosus Dollman 

Arvicanthis luctuosus Dollman, Ann. Mag. Nat. Hist. (8) 8: 339, Sep. 1911. 
Kaka, north of Fashoda, Anglo-Egyptian Sudan. 
Probably a synonym oi A. n. testicularis (Brajstrup, F. W., Vidensk. Medd. 
fra Dansk Naturh. Foren., Kjobenhavn, 99: 119, 11 Sep. 1935). 

Arvicanthis niloticus mordax Thomas 

Arvicanthis mordax Thomas, Ann. Mag. Nat. Hist. (8) 7: 460, May 1911. 
Panyam, northern Nigeria. 

Arvicanthis niloticus occidentalis Wroughton 

Arvicanthis occidentalis Wroughton, Ann. Mag. Nat. Hist. (7) 17: 377, Apl. 
1906. Bo, interior Sierra Leone. 
A synonym of A. n. rufinus according to Brsestrup (Vidensk. Medd. fra 
Dansk Naturh. Foren., Kjobenhavn, 99: 120, 11 Sep. 1935). 

Arvicanthis niloticus reichardi (Noack) 

Pelomys reichardi Noack, Zool. Jahrb., Syst., 2: 235, pi. 9, f. 4-7, 7 May 1887. 
Near Karema, Tanganyika Territory. 

Arvicanthis niloticus rufinus (Temminck) 

Mus rufinus Temminck, Esquisses Zool. sur la Cote de Guine, p. 163, 1853. 

"Toutes les parties de la Guin^." Gold Coast. 
Arvicanthis niloticus rufinus Brsestrup, Vidensk. Medd. fra Dansk Naturh. 

Foren., Kjobenhavn, 99: 120, 11 Sep. 1935. 

Arvicanthis niloticus setosus Thomas 

Arvicanthis setosus Thomas, Ann. Mag. Nat. Hist. (7) 15: 79, Jan. 1905. Fra- 
Fra Country, Gold Coast hinterland. 

Arvicanthis niloticus solatus Thomas 

Arvicanthis testicularis solatus Thomas, Ann. Mag. Nat. Hist. (9) 16: 194, July 
1925. Aouderas, Asben, French West Africa. 

Arvicanthis niloticus testicularis (Sundevall) 

Isomys testicularis Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, 
for 1842, p. 221, 1843. Bahr-el-Abiad, Anglo-Egyptian Sudan. 

Arvicanthis ochropus (Heuglin) 

Mus ochropus Heuglin, Reise in Nordost-Afrika, 2: 68, 1877. Bogos country, 
Abyssinia. 
An Arvicanthis according to Trouessart (Cat. Mamm. Viv. Foss., suppl., 
p. 375, 1904). 



378 bulletin: museum of comparative zoology 

Arvicanthis somalicus somalicus Thomas 

Arvicanthis somalicus Thomas, Proc. Zool. Soc. London, for 1902, 2: 312, foot- 
note, Apl. 1903. Shuk, northern Somaliland, 4000 ft. 

Arvicanthis somalicus chanleri Dollman 

Arvicanthis chanleri Dollman, Ann. Mag. Nat. Hist. (8) 8: 130, July 1911. 
Chanler Falls, Northern Guaso Nyiro, Kenya Colony. 

Arvicanthis somalicus reptans Dollman 

Arvicanthis reptans Dollman, Ann. Mag. Nat. Hist. (8) 8: 129, July 1911. 

Nyama Nyango, Northern Guaso Nyiro, Kenya Colony. 
Arvicanthis somalicus reptans Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, 48: no. 5, p. 99, 26 June 1912. 

Arvicanthis tenebrosus tenebrosus Kershaw 

Arvica7ithis abyssinicus tenebrosus Kershaw, Ann. Mag. Nat. Hist. (9) 11: 595, 
May 1923. Tabora, Tanganyika Territory. 

Arvicanthis tenebrosus rhodesi^ St. Leger 

Arvicanthis abyssinicus rhodesix St. Leger, Ann. Mag. Nat. Hist. (10) 10: 85, 
July 1932. SichiU River, Sesheke district. Northern Rhodesia. 

CoLOMYS Thomas and Wroughton 

Colomys Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 379, May 1907. 
Genotype Colomys goslingi Thomas and Wroughton. 

Colomys goslingi goslingi Thomas and Wroughton 

Colomys goslingi Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 380, 
May 1907. Gambi, Uele River, Congo Beige. 

Colomys goslingi bicolor Thomas 

Colomys bicolor Thomas, Ann. Mag. Nat. Hist. (8) 10: 42, July 1912. Bitye, 
southwestern Cameroons, 2000 ft. 

Colomys goslingi denti St. Leger 

Colomys goslingi denti St. Leger, Ann. Mag. Nat. Hist. (10) 6: 527, Oct. 1930. 
Elburgon, Kenya Colony, 8000 ft. 

Cricetomys Waterhouse. Giant Rats 

Cricetomys Waterhouse, Proc. Zool. Soc. London, no. 85, p. 2, July 1840. Sub- 
genus of Mus, type Cricetomys gambianus Waterhouse. 

See Hinton, M. A. C. Notes on the genus Cricetomys, with descriptions of 
four new forms. Ann. Mag. Nat. Hist. (9) 4: 282-289, Oct. 1919. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 379 

Cricetomys buchanani Thomas and Hinton 

Cricetomys buchanani Thomas and Hinton, Novitates Zool., 28; 7, 14 May 1921. 
Farniso, near Kano, northern Nigeria. 

Cricetomys cosensi Hinton 

Cricetomys cosensi Hinton, Ann. Mag. Nat. Hist. (9) 4: 286, Oct. 1919. Zanzi- 
bar Island. 

Cricetomys emini emini Wroughton 

Cricetomys gambianus emini Wroughton, Ann. Mag. Nat. Hist. (8) 5; 106, Jan. 
1910. Gaddu, Monbottu, Congo Beige. 

Cricetomys emini ansorgei Thomas 

Cricetomys ansorgei Thomas, Ann. Mag. Nat. Hist. (7) 13: 412, June 1904. 
Pungo Andongo, Angola. 

Cricetomys emini sanctus Hinton 

Cricetomys emini sanctus Hinton, Ann. Mag. Nat. Hist. (9) 4: 286, Oct. 1919. 
Inkongo, Sankuru, Congo Beige. 

Cricetomys emini vaughan-jonesi St. Leger 

Cricetomys emini vaughan-jonesi St. Leger, Ann. Mag. Nat. Hist. (10) 20: 
148, July 1937. Balovale, upper Zambesi, northwestern Rhodesia. 

Cricetomys gambianus gambianus Waterhouse 

Cricetomys gambianus Waterhouse, Proc. Zool. Soc. London, p. 2, July 1840. 

River Gambia, West Africa. 
Mus goliath Rtippell, Mus. Senckenbergianum, 3: 114, pi. 9; pi. 10, f.l, a-c, 

1842. Senegambia. 
Cricetomys gambiensis Temminck, Esquisses Zool. sur la Cote de Guine, p. 168, 

1853 (lapsus cal.). Guinea. 

Cricetomys gambianus adventor Thomas and Wroughton 
Cricetomys gambianus adventor Thomas and Wroughton, Proc. Zool. Soc. 
London, p. 295, 1 Aug. 1907. Coguno, Inhambane, Mozambique. 

Cricetomys gambianus cunctator Thomas and Wroughton 
Cricetomys gambianus cunctator Thomas and Wroughton, Proc. Zool. Soc. 
London, p. 171, Sep_. 1908. Tambarara, Gorongoza district, Mozambique. 

Cricetomys gambianus dichrurus Osgood 

Cricetomys gambianus dichrurus Osgood, Ann. Mag. Nat. Hist. (8) 5: 280, 
Mar. 1910. Agoulene, Anambara River, southern Nigeria. 



380 bulletin: museum of comparative zoology 

Cricetomys gambianus dissimilis Rochebrune 

Cricetomys dissimilis Rochebrune, Bull. Soc. Philomathique, Paris, (7) 9: 

86, 1885. Landana, Angola. 
Mus tephrus Rochebrune, Bull. Soc. Philomathique, Paris, (7) 9: 87, 1885. 

Landana, Angola. 

Cricetomys gambianus dolichops Osgood 

Cricetomys gambianus dolichops Osgood, Ann. Mag. Nat. Hist. (8) 5: 280, 
Mar. 1910. Como River, 70 miles from Gaboon, French Congo. 

Cricetomys gambianus elgonis Thomas 

Cricetomys gambianus elgonis Thomas, Ann. Mag. Nat. Hist. (8) 5: 198, Feb. 
1910. South face of Mt. Elgon, Kenya Colony, 10,000 ft. 

Cricetomys gambianus enguvi Heller 

Cricetomys gambianus enguvi Heller, Smithsonian Misc. Coll., 59: no. 16, p. 16, 
5 July 1912. Mt. Umengo, Taita Mts., Kenya Colony. 

Cricetomys gambianus grahami Hinton 

Cricetomys gambianus grahami Hinton, Ann. Mag. Nat. Hist. (9) 4: 283, 
Oct. 1919. Nuba Mts., southeastern Anglo-Egyptian Sudan. 

Cricetomys gambianus haagneri Roberts 

Cricetomys gambianus haagneri Roberts, Ann. Transvaal Mus., 11: 252, 

text-f. 1, 14 Sep. 1926. Zoutpansberg, 10 miles north of Louis Trichardt, 

Transvaal. 

Cricetomys gambianus kenyensis Osgood 

Cricetomys gambianus kenyensis Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: no. 2, p. 9, 16 Feb. 1910. South side of Mt. Kenya, Kenya Colony. 

Cricetomys gambianus kivuensis Lonnberg 

Cricetomys gambianus kivuensis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 
Stockholm, 58: art. 2, p. 75, 1 Sep. 1917. Masisi, Congo Beige. 

Cricetomys gambianus langi Hatt 

Cricetomys gambianus langi Hatt, Amer. Mus. Novitates, no. 708, p. 5, 4 Apl. 
1934. Faradje, upper Uele district, Congo Beige. 

Cricetomys gambianus liberie Osgood 

Cricetomys gambianus liberies Osgood, Ann. Mag. Nat. Hist. (8) 5: 281, Mar. 
1910. Fifty miles inland from Monrovia, Liberia. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 381 

Cricetomys gambianus luteus DoUman 

Cricetomys gambianus luteus Dollman, Ann. Mag. Nat. Hist. (8) 8: 124, July 
1911. Igembi Hills, northeast of Mt. Kenya, Kenya Colony. 

Cricetomys gambianus olivx^ Dollman 

Cricetomys gambianus olivix Dollman, Ann. Mag. Nat. Hist. (8) 8: 258, Aug. 
1911. Fika, Bornu, northern Nigeria. 

Cricetomys gambianus osgoodi Heller 

Cricetomys gambianus osgoodi Heller, Smithsonian Misc. Coll., 59: no. 16, p. 16, 
5 July 1912. Mazeras, Kenya Colony. 

Cricetomys gambianus poensis Osgood 

Cricetomys gambianus poensis Osgood, Ann. Mag. Nat. Hist. (8) 5: 279, Mar. 
1910. Bubi Town, Bantaberi, Island of Fernando Po. 

Cricetomys gambianus proparator Wroughton 

Cricetomys gambianus proparator Wroughton, Ann. Mag. Nat. Hist. (8) 5: 
107, Jan. 1910. Ruwenzori East, Uganda, 6000 ft. 

Cricetomys gambianus raineyi Heller 

Cricetomys gambianus raineyi Heller, Smithsonian Misc. Coll., 59: no. 16, 
p. 15, 5 July 1912. Mt. Gargues, Mathews Range, Kenya Colony, 6000 ft. 

Cricetomys gambianus viator Thomas 

Cricetomys gambianus viator Thomas, Ann. Mag. Nat. Hist. (7) 13: 413, 
June 1904. Likangala, Nyasaland. 

Cricetomys microtis Lonnberg 

Cricetomys microtis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., Stockholm, 
(2) 58: art. 2, p. 77, 1 Sep. 1917. Masisi, Congo Beige. 

Cricetomys servorum Hinton 

Cricetomys servorum Hinton, Ann. Mag. Nat. Hist. (9) 4: 288, Oct. 1919. 
Lagos, southern Nigeria. 

Dasymys Peters.- Shaggy-haired Rats 

Dasymys Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 12, 1875. Geno- 
type Dasymys gueinzii Peters = M us incomtus Sundevall. 

Dasymys bentley^ bentley^ (Thomas) 

Mus {Dasymys) bentleyse Thomas, Ann. Mag. Nat. Hist. (6) 10: 179, Aug. 
1892. Ngombi, Lower Congo, Congo Beige. 



382 bulletin: museum of comparative zoology 

DaSYMYS BENTLEY.E MEDIUS ThomaS 

Dasymys medius Thomas, Ann. Mag. Nat. Hist. (7) 18: 143, Aug. 1906. 

Mubuku Valley, Ruwenzori East, Uganda, 6000 ft. 
Dasymys bentleyae medius Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 123, 

1919. 

Dasymys foxi Thomas 

Dasymys foxi Thomas, Ann. Mag. Nat. Hist. (8) 9: 685, June 1912. Panyam, 
northern Nigeria. 

Dasymys helukus helukus Heller 

Dasymys helukus Heller, Smithsonian Misc. Coll., 54: no. 1924, p. 2, pi. 1, b 
(skull), 28 Feb. 1910. Sirgoit, Guas Ngishu Plateau, Kenya Colony. 

Dasymys helukus nigridius Hollister 

Dasymys helukus nigridius Hollister, Smithsonian Misc. Coll., 66: no. 10, p. 2, 
26 Oct. 1916. Naivasha Station, Kenya Colony. 

Dasymys helukus sav annus Heller 

Dasymys savannus Heller, Smithsonian Misc. Coll., 56: no. 17, p. 14, 28 Feb. 
1911. Fort Hall, Kenya Colony. 

Dasymys incomtus incomtus (Sundevall) 

Mus incomtus Sundevall, Ofversigt af Kongl. Svenska Vet.-Akad. Forhandl., 

Stockholm, for 1846,3: no. 4, p. 120, 1847. "Caffraria propePortum Natal." 
Dasymys gueinzii Peters, Monatsb. K. Preuss. Akad. Wiss., Berhn, p. 12, pi. 1 

(col.); pi. 2 (skull), 1875. Interior of Port Natal, Natal. 

Dasymys incomtus capensis Roberts 

Dasymys incomtus capensis Roberts, Ann. Transvaal Mus., 18: 254, 21 Feb. 
1936. La Plisante, Wolseley, Cape Province. 

Dasymys incomtus puscus De Winton 

Dasymys incomtus fuscus De Winton, Proc. Zool. Soc. London, for 1896, p. 
804, Apl. 1897. Mazoe, Mashonaland, Southern Rhodesia. 

Dasymys incomtus griseifrons Osgood 

Dasymys incomptus (sic) griseifrons Osgood, Zool. Ser. Field Mus. Nat. Hist., 

20: 255, 28 Dec. 1936. Southwest side of Lake Tana, near Dungulbar, 

Gojjam, Ethiopia. 

Dasymys montanus Thomas 

Dasymys montanus Thomas, Ann. Mag. Nat. Hist. (7) 18: 143, Aug. 1906; 

Thomas and Wroughton, Trans. Zool. Soc. London, 19: 513, Mar. 1910. 

Mubuku Valley, Ruwenzori East, Uganda, 12,500 ft. 



^ ALLEN: CHECKLIST OF AFRICAN MAMMALS 383 

Dasymys ntjdipes nudipes (Peters) 

Mus (Isomys) nvdipes Peters, Jorn. Sci. Math., Phys. e Nat., Lisboa, (1) 3: 
126, Dec. 1870. Huilla, Angola. 

Dasymys nudipes edsoni Hatt 

Dasymys nudipes edsoni Hatt, Amer. Mus. Novitates, no. 708, p. 6, 4 Apl, 
1934. Lukolela, middle Congo, Congo Beige. 

Dasymys orthos Heller 

Dasymys orthos Heller, Smithsonian Misc. Coll., 56: no. 17, p. 13, 28 Feb. 1911. 
Butiaba, Albert Nyanza, Uganda. 

Dasymys rufulus Miller 

Dasymys rufulus Miller, Proc. Washington Acad. Sci., 2: 639, text-f. 40, 28 
Dec. 1900. Mount Coffee, Liberia. 

Dasymys shawi Kershaw 

Dasymys shawi Kershaw, Ann. Mag. Nat. Hist. (9) 13: 25, Jan. 1924. Mount 
Baginzi, southern Bahr-el-Ghazal, near Congo border, 40 miles south- 
southeast of Yambio, Anglo-Egyptian Sudan. 

Dasymys thomasi (De Winton) 

MusthomasiBeWinton, Ann. Mag. Nat. Hist. (6) 20: 321, Sep. 1897. Galanga, 
Angola. 



Dephomys Thomas 

Dephomys Thomas, Ann. Mag. Nat. Hist. (9) 17: 177, Jan. 1926. Genotype 
Mus defua Miller. 

Dephomys defua (Miller). Bristly Rat 

Mus defua Miller, Proc. Washington Acad. Sci., 2: 635, 28 Dec. 1900. Mount 
Coffee, Liberia. 

Desmomys Thomas 

Desmomys Thomas, Ann. Mag. Nat. Hist. (8) 5: 284, Mar. 1910. Genotype 
Pelomys harringtoni Thomas. 

Desmomys harringtoni (Thomas) 

Pelomys harringtoni Thomas, Proc. Zool. Soc. London, for 1902, 2: 313, Apl. 
1903. Katchisa, Kutai, western Shoa, Abyssinia. 



384 bulletin: museum of comparative zoology 

Desmomys rex (Thomas) 

Arvicanthis rex Thomas, Ann. Mag. Nat. Hist. (7) 18: 304, Oct. 1906. Charada 
Forest, Kaffa, central Abyssinia. 
"It may be provisionally considered as a giant member of Desmomys" 
(Thomas, O., Ann. Mag. Nat. Hist. (8) 18: 68, July 1916). 

Note: Des?nomys dembeensis (Riippell) proves to be an Arvicanthis; see 
Thomas, O., Ann. Mag. Nat. Hist. (10) 1: 302, footnote, Feb. 1928. 



Hybomys Thomas. Back-striped Mice 

Hybomys Thomas, Ann. Mag. Nat. Hist. (8) 5: 85, Jan. 1910. Genotype Mus 

univittatus Peters. 
Ty-pomys Thomas, Ann. Mag. Nat. Hist. (8) 7: 382, Apl. 1911. Genotype Mus 

trivirgatus Temminck. Synonym of Hybomys (see Ingoldby, C. M., Ann. 

Mag. Nat. Hist. (10) 3: 522, May 1929). 

Hybomys univittatus univittatus (Peters) 

Mus univittatus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 479, pi. 2, 

f. 3 (col.), 1876. Dongila, Gaboon. 
Mus (Isomys) rufocanus Tullberg, Nova Acta Reg. Soc. Sci., Upsala, (3) 16: 

no. 12, p. 23, pi. 2, f. 9, 10; pi. 3, f. 41, 42, 1893. Vevaka, Cameroons. 

Hybomys univittatus badius Osgood 

Hybotnys univittatus badius Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 254, 

28 Dec. 1936. Southwest slope of Mt. Cameroon, Cameroon mandate, 

British Nigeria. 

Hybomys univittatus lunaris (Thomas) 

Mus univittatus /imaris Thomas, Ann. Mag. Nat. Hist. (7) 18: 145, Aug. 1906. 
Mubuku Valley, Ruwenzori East, Uganda, 6000 ft. 

Hybomys univittatus planifrons (Miller) 

Arvicanthis planifrons Miller, Proc. Washington Acad. Sci., 2: 641, text-f. 41, 
a, 28 Dec. 1900. Mount Coffee, Liberia. 

Hybomy's trivirgatus trivirgatus (Temminck) 

Mus trivirgatus Temminck, Esquisses Zool. sur la Cote de Guin6, p. 159, 1853. 
Dabacrom, Gold Coast. 

Hybomys trivirgatus pearcei Ingoldby 

Hybomys trivirgatus pearcei Ingoldby, Ann. Mag. Nat. Hist. (10) 3: 522, May 
1929. Lagos, Nigeria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 385 

Hylenomys Thomas 

Hylenomys Thomas, Ann. Mag. Nat. Hist. (9) 15: 667, June 1925. Genotype 
Hylenomys callewaerii Thomas. 

Hylenomys callew^rti Thomas 

Hylenomys calleuserti Thomas, Ann. Mag. Nat. Hist. (9) 15: 668, June 1925. 
Luluabourg, Lualaba, southern Congo Beige, 610 m. 

Hylomyscus Thomas. Climbing Wood Mice 

Praomys Thomas, Ann. Mag. Nat. Hist. (8) 15: 477, Dec. 1915, in part. 
Hylomyscus Thomas, Ann. Mag. Nat. Hist. (9) 17: 178, Jan. 1926. Genotype 
Epimys seta Thomas; with list of included species. 

Hylomyscus ^ta ^ta (Thomas) 

Epimys seta Thomas, Ann. Mag. Nat. Hist. (8) 7: 591, June 1911; (8) 10: 43, 
July 1912 (additional notes on size). Bitye, Ja River, Cameroons. 

Hylomyscus mta laticeps Osgood 

Hylomyscus seta laticeps Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 247, 28 

Dec. 1936. Southwest slope of Mt. Cameroon, Cameroon Mandate, 

British Nigeria. 

Hylomyscus ^ta schoutedeni (Dollman) 

Epimys schoutedeni Dollman, Revue Zool. Africaine, 4: 82, 25 July 1914, 
Mambaka, Congo Beige. 

Hylomyscus ^ta weileri (Lonnberg and Gyldenstolpe) 

Rattus {Praomys) weileri Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17B: 

no. 5, p. 3, 2 May 1925. Burunga, western foothills of Mt. Mikeno, Bir- 

unga Volcanoes, Congo Beige, 2400 m. 

Hylomyscus alleni alleni (Waterhouse) 

Mus alleni Waterhouse, Proc. Zool. Soc. London, for 1837, p. 77, 22 Jan. 1838. 
Fernando Po, Gulf of Guinea. 

Hylomyscus alleni carillus (Thomas) 

Mus carillus Thomas, Ann. Mag. N«,t. Hist. (7) 13: 418, June 1904. Pungo 
Andongo, Angola, 1200 m. 

Hylomyscus alleni simus G. M. Allen and Coolidge 

Hylomyscus alleni simus G. M. Allen and Coolidge, in R. P. Strong and others, 

The African Republic of Liberia and the Belgian Congo, 2: 599, 1930. 

Merikay, interior of Liberia. 



386 bulletin: museum of comparative zoology 

Hylomyscus DENNIS DENNi^E (Thomas) 

Mus dennise Thomas, Ann. Mag. Nat. Hist. (7) 18: 144, Aug. 1906. Thomas 

and Wroughton, Trans. Zool. Soc. London, 19: 504, Mar. 1910. Ruwen- 

zori East, Mubuku Valley, Uganda, 7000 ft. 
Epimys endorobse Heller, Smithsonian Misc. Coll., 56: no. 9, p. 3, pi. 2, 3 figs., 

22 July 1910. Western edge of Mau Forest, 25 miles north of Eldoma 

Ravine, Kenya Colony, 8600 ft. 
A sjmonym of dennise according to Hollister, Bull. U. S. Nat. Mus., no. 99, 

pt. 2, p. 79, 1919. 

Hylomyscus denni^ btjtleri (Wroughton) 

Mus butleri Wroughton, Ann. Mag. Nat. Hist. (7) 20: 503, Dec. 1907. Between 
Chak Chak and Dem Zubeir, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 

Hylomyscus dennle vulcanorum (Lonnberg and Gyldenstolpe) 

Rattus {Praomys) dennias vulcanorum Lonnberg and Gyldenstolpe, Arkiv f. 

Zool., 17B: no. 5, p. 4, 2 May 1925. Mt. Karisimbi, Birunga Volcanoes, 

Congo Beige, 3800 m. 

Hylomyscus stella stella (Thomas) 

Epimys stella Thomas, Ann. Mag. Nat. Hist. (8) 7: 590, June 1911. Ituri 
Forest, between Mawambi and Avakubi, eastern Congo Beige. 
Range: Eastern Congo to Cameroons 

Hylomyscus stella kaimos^ (Heller) 

Epimys alleni kainiosie Heller, Smithsonian Misc. Coll., 59: no. 16, p. 7, 5 July 

1912. Kaimosi, Kakumega Forest, Kenya Colony. 
Rattus Stella kaimosse Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 82, 1919. 



Komemys de Beaux. See Pelomys 
Leggada Gray. Pygmy Mice 

Leggada Gray, Charlesworth's Mag. Nat. Hist., 1 : 586, Nov. 1837. Genotype 

Leggada booduga Gray. On the generic characters see Thomas, O., Journ. 

Bombay Nat. Hist. Soc, 26: 417-420, 20 May 1919. 
Nannomys Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 480, pi. 2, f. 

4, Aug. 1876. Subgenus of Mus, type Mus setulosus Peters. 
Pseudoconomys Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 531, 8 Dec. 1896. 

Subgenus of Mus, type Mus proconodon Rhoads. 

Leggada bella bella Thomas 

Leggada bella Thomas, Ann. Mag. Nat. Hist. (8) 5: 87, Jan. 1910. Madiakos, 
Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 387 

Leqgada BELLA ENCLAVE (Heller) 

Mus bellus enclavse Heller, Smithsonian Misc. Coll., 56: no. 17, p. 8, 28 Feb. 
1911. Rhino Camp, Lado Enclave. 

LeGGADA BELLA GALLARUM ThomaS 

Leggada bella gallarum Thomas, Ann. Mag. Nat. Hist. (8) 5: 88, Jan. 1910. 
Harar, eastern Abyssinia, 6000 ft. 

Leggada bella gondokor^ (Heller) 

Mus bellus gondokorae. Heller, Smithsonian Misc. Coll., 56: no. 17, p. 8, 28 Feb. 
1911. Gondokoro, Uganda. 

Leggada bella induta Thomas 

Leggada bella induta Thomas, Ann. Mag. Nat. Hist. (8) 5: 89, Jan. 1910. 
Molopo, west of Morokwen, northern Bechuanaland. 

Leggada bella kasaica Cabrera 

Leggada bella kasaica Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 24: 
222, 28 Apl. 1924. St. Joseph de Luluabourg, Kasai, Congo Beige. 

Leggada bella marica Thomas 

Leggada bella marica Thomas, Ann. Mag. Nat. Hist. (8) 5: 88, Jan. 1910. 
Beira, Mozambique. 

Leggada bella petila (Hollister) 

Mus bellus petilus Hollister, Smithsonian Misc. Coll., 66: no. 10, p. 3, 26 Oct. 
1916. Southern Guaso Nyiro River, Kenya Colony. 

Leggada bella sybilla Thomas 

Leggada bella sybilla Thomas, Ann. Mag. Nat. Hist. (9) 2: 484, Dec. 1918. 
Benguella, Angola. 

Leggada bella vicina Thomas 

Leggada bella vicina Thomas, Ann. Mag. Nat. Hist. (8) 5: 88, Jan. 1910. 
Takaungu, near Mombasa, Kenya Colony. 

Leggada birtjngensis (Lonnberg and Gyldenstolpe) 

Mus birungensis Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17 B: no. 5, p. 5, 
2 May 1925. Mt. Mikeno, Birunga Volcanoes, Congo Beige, 3400 m. 

Leggada bufo bufo Thomas 

Leggada bufo Thomas, Ann. Mag. Nat. Hist. (7) 18: 145, Aug. 1906. Thomas 

and Wroughton, Trans. Zool. Soc. London, 19: 506, pi. 23, lower f. (col.). 

Mar. 1910. Ruwenzori East, Uganda, 6000 ft. 



388 bulletin: museum of comparative zoology 

Leggada btjfo wambutti (Lonnberg and Gyldenstolpe) 

Mus bufo wambutti Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17B: no. 5, 
p. 4, 2 May 1925. Campi ya Wambutti, Ituri Forest, eastern Congo Beige. 

Leggada deserti deserti Thomas 

Leggada deserti Thomas, Ann. Mag. Nat. Hist. (8) 5: 90, Jan. 1910. Molopo, 
west of Morokwen, northern Bechuanaland. 

Leggada deserti PRExoRiiE Roberts 

Leggada deserti pretorise Roberts, Ann. Transvaal Mus., 11: 252, 14 Sep. 1926. 
Pretoria, Transvaal. 

Leggada deserti valschensis Roberts 

Leggada deserti valschensis Roberts, Ann. Transvaal Mus., 11: 251, 14 Sep. 
1926. Bothaville, Orange Free State. 

Leggada emesi (Heller) 

Mus musculoides emesi Heller, Smithsonian Misc. Coll., 56: no. 17, p. 5, 28 

Feb. 1911. Kabula Muliro, Uganda. 
A distinct species from L. triton (Hollister, N., Bull. U. S. Nat. Mus., no. 99, 

pt. 2, p. 96, 1919). 

Leggada gerbillus G. M. Allen and Loveridge 

Leggada gerbillus G. M. Allen and Loveridge, Bull. Mus. Comp. Zool., 75: 
112, Feb. 1933. Dodoma, Ugogo, Tanganyika Territory. 

Leggada grata grata Thomas and Wroughton 

Leggada grata Thomas and Wroughton, Trans. Zool. Soc. London, 19: 507, 

Mar. 1910. Mubuku Valley, Ruwenzori East, Uganda. 
Mus gratus sungarse. Heller, Smithsonian Misc. Coll., 56: no. 17, p. 7, 28 Feb. 

1911. West Kenya Forest Station, Kenya Colony, 7500 ft. Synonym of 

L. grata (see Hollister, N., Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 102, 

1919). 

Leggada grata soricoides (Heller) 

Mus gratus soricoides Heller, Smithsonian Misc. Coll., 63: no. 7, p. 10, 24 June 
1914. Mt. Mbololo, Taita Hills, Kenya Colony. 

Leggada haussa Thomas and Hinton 

Leggada haussa Thomas and Hinton, Novitates Zool., 27: 319, 15 June 1920. 
Farniso, near Kano, northern Nigeria, 1700 ft. 

Leggada mahomet (Rhoads) 

Mus mahomet Rhoads, Proc. Acad. Nat. Sci. Philadelphia, p. 532, 8 Dec. 1896. 
Sheikh Mahomet, western British Somaliland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 389 

Leggada matschiei (Wettstein) 

Mus (Leggada) matschiei Wettstein, Anzeiger K. Akad. Wiss. Wien, math.- 

nat. Kl., 53: 160, 1916. Gebel Debri, south of Gebel Gulfan, southern 

Kordofan, Anglo-Egyptian Sudan. 

Leggada minutoides minutoides (A. Smith) 

Mus minutoides A. Smith, South African Quart. Journ., 2: 157, 1834. Thomas, 

Proc. Zool. Soc. London, p. 13, June 1888 (synonymy). Near Cape Town, 

Cape of Good Hope. 
Mu^ minimus Peters, Ber. iiber Verhandl. K. Preuss. Akad. Wiss., Berhn, p. 

274, 1852. Tette, Buio, Macanga, Mozambique. Not Mv^ minimus 

White, 1789, which is a Micromys. 

Leggada minutoides neavei Thomas 

Leggada neavei Thomas, Ann. Mag. Nat. Hist. (8) 5: 90, Jan. 1910. Petauke, 
eastern Loangwe district, northeastern Rhodesia, 2400 ft. 

Leggada minutoides orangi^ Roberts 

Leggada orangix Roberts, Ann. Transvaal Mus., 11: 251, 14 Sep. 1926. Krui- 
sementfontein, Viljoens Drift, Orange Free State. 

Leggada minutoides umbrata Thomas 

Leggada minutoides umbrata Thomas, Ann. Mag. Nat. Hist. (8) 5: 86, Jan. 
1910. Wakkerstroom, southeastern Transvaal, 5900 ft. 

Leggada musculoides (Temminck) 

Mus muscidoides Temminck, Esquisses Zool. sur la Cote de Guin6, p. 161, 

1853. "Cote de Guine," West Africa, 
ilfiis (Nannomys) setidosus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, 

p. 480, pi. 2, f. 4 (col.), 1876. Victoria, Cameroons. 

Leggada paulina Thomas 

Leggada paulina Thomas, Ann. Mag. Nat. Hist. (9) 2: 485, Dec. 1918. Bitye, 
Ja River, Cameroons, 2000 ft. 
Probably a race of musculoides (fide Thomas). 

Leggada proconodon proconodon (Rhoads) 

Mtts (Pseudoconomys) proconodon Rhoads, Proc. Acad. Nat. Sci. Philadelphia, 
p. 531, 8 Dec. 1896. Sheikh Husein, western British Somaliland. 

Leggada proconodon pasha Thomas 

Leggada pasha Thomas, Ann. Mag. Nat. Hist. (8) 5: 89, Jan. 1910. Tingasi, 

Monbuttu, Uele district, Congo Beige. 
Mus proconodon pasha Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 247, 28 

Dec. 1936. 



390 bulletin: museum of comparative zoology 

Leggada sorella Thomas 

Leggada sorella Thomas, Ann. Mag. Nat. Hist. (8) 4: 548, Dec. 1909. Kirui, 
Mt. Elgon, Kenya Colony, 6000 ft. 

Leggada spreta (Lataste) 

Mas spretus Lataste, Actes Soc. Linn, de Bordeaux, (4) 7: 27, text-f. 5, 6, 20 

Mar. 1883. Oued Magra, between M'sila and Barika, north of Hodna, 

high plateau of Algeria. 
The cranial proportions are like those of a Leggada as measured on Lataste's 
figures, but possibly this name is a synonym of a form of Mus. 

Leggada tenella tenella Thomas 

Leggada tenella Thomas, Proc. Zool. Soc. London, 1 : 298, 6 Aug. 1903. Roseires, 
Blue Nile, Anglo-Egyptian Sudan. 

Leggada tenella acholi (Heller) 

Mus tenellus acholi Heller, Smithsonian Misc. Coll., 56: no. 17, p. 6, 28 Feb. 
1911. Rhino Camp, Lado Enclave. 

Leggada tenella stjahelica Thomas 

Leggada tenella suahelica Thomas, Ann. Mag. Nat. Hist. (8) 6: 312, Sep. 1910. 
Taveta, foot of Mt. Kilimanjaro, Tanganyika Territory. 

Leggada triton triton Thomas 

Leggada triton Thomas, Ann. Mag. Nat. Hist. (8) 4: 548, Dec. 1909. Kirui, 

Mt. Elgon, Kenya Colony, 6000 ft. 
Leggada naivashse Heller, Smithsonian Misc. Coll., 54: no. 1924, p. 2, pi. 2, a, 

28 Feb. 1910. Base of Aberdare Mts., eastern edge of Naivasha Plains, 

Kenya Colony. 

Leggada triton fors Thomas 

Leggada fors Thomas, Ann. Mag. Nat. Hist. (8) 4: 548, Dec. 1909 (in descrip- 
tion of L. triton where skull length is given, as noted by HoUister, in Bull. 
U. S. Nat. Mus., no. 99, pt. 2, p. 96, 1919). Butagu Valley, Ruwenzori 

West. 

See also Thomas and Wroughton, Trans. Zool. Soc. London, 19: 506, Mar. 
1910. 

Leggada triton murilla Thomas 

Leggada triton murilla Thomas, Ann. Mag. Nat. Hist. (8) 5: 91, Jan. 1910. 
Machakos, Kenya Colony, 5400 ft. 

Leggada wam^e (Heller) 

Mus wamse Heller, Smithsonian Misc. Coll., 56: no. 17, p. 5, 28 Feb. 1911. 
Kapiti Plains, Kenya Colon5\ 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 391 

Lemniscomys Trouessart. Striped Grass Mice 

Lemniscomys Trouessart, Bull. Soc. d'Etudes Sci., Angers, 10: 124, 1881. Sub- 
genus of Mus; type Mas barbarus Linnseus. 

For diagnostic characters, see Thomas, O., Ann. Mag. Nat. Hist. (8) 18: 68, 
July 1916. 

Lemniscomys barbarus barbarus (Linnaeus) 

Mies barbarus Linnseus, Syst. Nat., ed. 12, 1: pt. 2, addenda, unpaged at end 
of vol., 1767. "Barbaria," i.e. Morocco. 

Lemniscomys barbarus albolineatus (Osgood) 

Arvicanthis barbarus albolineatus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: 11, 16 Feb. 1910. Lukenya Mt., Ulukenia Hills, Kenya Colony. 

Lemniscomys barbarus convictus (Osgood) 

Arvicanthis barbarus convictus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: 10, 16 Feb. 1910. Voi, Kenya Colony. 

Lemniscomys barbarus fasciatus (Wroughton) 

Arvicanthis fasciatus Wroughton, Ann. Mag. Nat. Hist. (7) 17: 377, Apl. 1906. 
Agoulerie, Anambra Creek, southern Nigeria. 

Lemniscomys barbarus ifniensis Morales Agacino. Morocco Striped Grass 

Mouse 
Lemniscomys barbarus ifniensis Morales Agacino, Bol. Soc. Espanola Hist. 

Nat., Madrid, 35: 390, text-f. 1, 25 Aug. 1935. Sidi Ifni, Ifni, Spanish 

Morocco. 

Lemniscomys barbarus manteufeli Matschie 

Lemniscomys barbarus maiiteufeli Matschie, Sitzb. Ges. Naturf. Freunde, Ber- 
lin, p. 338, Oct. 1911. Mwanza, south coast of Lake Victoria, Tanganyika 
Territory. 

Lemniscomys barbarus Nigeria (Thomas) 

Arvicanthis barbarus nigeriae Thomas, Ann. Mag. Nat. Hist. (8) 9: 272, Feb. 
1912. Panyan Plateau, northern Nigeria. 

Lemniscomys barbarus spekei (De Winton) 

Arvicanthis spekei De Winton, Ann. Mag. Nat. Hist. (6) 20: 318, Sep. 1897. 

Unyamuezi, Tanganyika Territory. 
Lemniscomys barbarus spekei Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 338, Oct. 1911. 



392 bulletin: museum of comparative zoology 

Lemniscomys barbarus zebra (Heuglin) 

Mas zebra Heuglin, Nova Acta Acad. Caes. Leop. -Carol., Halle, 31: Abhandl., 
no. 7, p. 10, 1864. Country of the Req negroes, Djur and Bongo, Anglo- 
Egyptian Sudan. 

Lemniscomys dunni dunni (Thomas) 

Arvicanthis dunni Thomas, Proc. Zool. Soc. London, 1 : 297, 6 Aug. 1903. Kaga 
Hills, western Kordofan, Anglo-Egyptian Sudan. 
A race of L. barbarus according to Brsestrup (Vidensk. Medd. fra Dansk 
Naturh. Foren., Kjobenhavn, 99: 120, 11 Sep. 1935). 

Lemniscomys dunni nubalis Thomas and Hinton 

Lemniscomys dunni nubalis Thomas and Hinton, Proc. Zool. Soc. London, p. 
263, 6 July 1923. Talodi, Nuba Country, southern Kordofan, Anglo- 
Egyptian Sudan. 

Lemniscomys dunni olqa Thomas and Hinton 

Lemniscomys alga Thomas and Hinton, Novitates Zool., 28: 9, 14 May 1921. 
Takoukout, Damergou, French West Africa. 

Lemniscomys griselda griselda (Thomas). Single-striped Grass Mouse 
Arvicanthis dorsalis griselda Thomas, Ann. Mag. Nat. Hist. (7) 13: 414, June 
1904. Muene Coshi, Jinga country, Angola. 
For use of griselda as a specific name in place of dorsalis, see Thomas, O., 
Proc. Zool. Soc. London, p. 385, 12 July 1927. 

Lemniscomys griselda calidior (Thomas and Wroughton) 
Arvicanthis dorsalis calidior Thomas and Wroughton, Proc. Zool. Soc. London, 
p. 545, 30 Oct. 1908. Tambarara, Gorongoza Mts., Zambesia, Mozam- 
bique. 

Lemniscomys griselda fitzsimonsi Roberts 

Lemniscomys griselda fitzsiynonsi Roberts, Ann. Transvaal Mus., 15: 11, 1 Oct. 
1932. Kaotwe Pan, central Kalahari, Bechuanaland. 

Lemniscomys griselda linulus (Thomas) 

Arvicanthis linulus Thomas, Ann. Mag. Nat. Hist. (8) 6: 429, Oct. 1910. 
Gamon, French Gambia. 

Lemniscomys griselda maculosus (Osgood) 

Arvicanthis dorsalis maculosus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 

10: no. 3, p. 17, 7 Apl. 1910. Voi, Kenya Colony. 
Arvicanthis dorsalis phasotis Thomas, Ann. Mag. Nat. Hist. (8) 6: 429, Oct. 

1910. Mazeras, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 393 

Lemniscomys griselda mearnsi Heller 

Lemniscomys dorsalis mearnsi Heller, Smithsonian Misc. Coll., 63: no. 7, p. 12, 
24 June 1914. Fort Hall, Kenya Colony. 

Lemniscomys griselda rosalia (Thomas) 

Arvicanthis dorsalis rosalia Thomas, Ann. Mag. Nat. Hist. (7) 13: 414, June 
1904. Monda, Nguru Mts., Tanganyika Territory. 

Lemniscomys griselda sabulatus Thomas 

Lemniscomys griselda sabulala Thomas, Proc. Zool. Soc. London, p. 385, 12 
July 1927. Sandfontein, Gobabis district, Southwest Africa. 

Lemniscomys griselda spinalis Thomas 

Mus dorsalis A. Smith, Illustr. Zool. South Africa, Mamm., pi. 46, f. 2, and 
text, Oct. 1845. "North of the Orange River"; assumed to be the western 
Transvaal (Roberts, A., Ann. Transvaal Mus., 5: 271, 1917). 
Name preoccupied by Mus dorsalis G. Fischer, 1814. 

Lemniscomys griselda spinalis Thomas, Ann. Mag. Nat. Hist. (8) 18: 69, foot- 
note, July 1916. New name to replace Miis dorsalis A. Smith, 1845, 
preoccupied by Mus dorsalis G. Fischer, 1814, a Sicista. 

Lemniscomys griselda zuldensis Roberts 

Lemniscomys griselda zuluensis Roberts, Ann. Transvaal Mus., 14: 235, 28 
July 1931. Manaba, northern Zululand, Cape of Good Hope. 

?Lemniscomys lineato-affinis (Hedenborg) 

Mus lineato-affinis Hedenborg, Oken's Isis, p. 5, 1832. Sennaar. Status un- 
certain; probably not intended as a technical name. 

Lemniscomys lynesi Thomas and Hinton 

Lemniscomys lynesi Thomas and Hinton, Proc. Zool. Soc. London, p. 267^ 

6 July 1923. Central Jebel Marra, Darfur, Anglo-Egyptian Sudan, 

7700-9750 ft. 

Lemniscomys macculus macctjlus (Thomas and Wroughton) 

Arvicanthis macculus Thomas and Wroughton, Trans. Zool. Soc. London, 19: 

515, pi. 24 (left-hand fig.), Mai:. 1910. Moki, southeastern Ruwenzori, 

Uganda. 
Lemniscomys macculus macculus Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, 

p. 138, 1919. 

Lemniscomys macculus akka (Thomas) 

Arvicanthis 7nacculus akka Thomas, Ann. Mag. Nat. Hist. (8) 16: 479, Dec. 
1915. Tingasi, Bomakandi River, Congo Beige. 



394 bulletin: museum of comparative zoology 

Lemniscomys oweni oweni (Thomas) 

Arvicanthis oweni Thomas, Ann. Mag. Nat. Hist. (8) 8: 120, July 1911. Gemen- 
julla, French Gambia. 

Lemniscomys oweni orientalis Hatt 

Lemniscomys oweni orientalis Hatt, Amer. Mus. Novitates, no. 790, p. 2, 
10 Apl. 1935. Faradje, Congo Beige. 

Lemniscomys striatus striatus (Linnaeus). Striped Grass Mouse 

Mus striatus Linnaeus, Syst. Nat., ed. 10, 1: 62, 1758. "India" = Sierra Leone 

(see Thomas, Proc. Zool. Soc. London, p. 148, Mar. 1911). 
Mus orientalis Desmarest, Nouv. Dictionnaire d'Hist. Nat., ed. 2, 29: 59, 

1819. "de rinde." Based on Seba; not Mus orientalis Cretzschmar, 

1826, =Mus musculus Linnaeus. 
Golunda pulchella Gray, Proc. Zool. Soc. London, p. 57, July 1864. "West 

Africa." 

Lemniscomys striatus ardens (Thomas) 

Arvicanthis pulcheUus ardens Thomas, Ann. Mag. Nat. Hist. (8) 6: 313, Sep. 

1910. Rombo, Mt. Kilimanjaro, Tanganyika Territory, 6000 ft. 

Lemniscomys striatus lulu^ Matschie 

Lemniscomys luluse Matschie, Zeitschr. f. Saugetierk., 1: 112, 24 Dec. 1926. 
Shibakala, near Luluabourg, Lulua River, Congo Beige. 

Lemniscomys striatus massaicus (Pagenstecher) 

Mus {Lemniscomys) barbarus L. var. massaicus Pagenstecher, Jahrb. Ham- 
burg. Wiss. Anstalt, Hamburg, 2: 45, 1885. Lake Naivasha, and Nguru- 
man, Kenya Colony. 

Arvicanthis pidchellus micropus Heller, Smithsonian Misc. Coll., 56: no. 17, 
p. 9, 28 Feb. 1911. Rhino Camp, Lado Enclave. 

Lemniscomys pulcheUus spermophilus Heller, Smithsonian Misc. Coll., 59: 
no. 16, p. 11, 5 July 1912. Mt. Gargues, Mathews Range, Kenya Colony. 
See Hollister, N., Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 139 ,1919. 

Lemniscomys striatus pulcher (Wroughton) 

Arvicaydhis pulcher Wroughton, Ann. Mag. Nat. Hist. (7) 17: 378, Apl. 
1906. Agoulerie, Anambra Creek, southern Nigeria. 

Lemniscomys striatus venustus (Thomas) 

Arvicanthis striatus venustus Thomas, Ann. Mag. Nat. Hist. (8) 7: 461, May 

1911. Panyam, northern Nigeria, 4000 ft. 

Lemniscomys striatus wroughtoni (Thomas) 

Arvicanthis wroughtoni Thomas, Ann. Mag. Nat. Hist. (8) 5: 85, Jan. 1910. 
Nono, west of Addis Ababa, Abyssinia (= Ethiopia). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 395 



LoPHUROMYS Peters 

Lasiomys Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 409, pi., f. 5-8, 

1866 (not of Burmeister, 1854). Genotype Lasiomys afer Peters = Mus 

sikapusi Temminck. 
Lophuromys Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 234, 1874. 

Genotype Lasiomys afer Peters = Mus sikapusi Temminck. New name 

in place of Lasiomys Peters, not of Burmeister, 1854. 

Lophuromys aquilus aquilus (True) 

Mus aquilus True, Proc. U. S. Nat. Mus., 15: 460, text-f. 1, 26 Oct. 1892. Mt. 

Kilimanjaro, Tanganyika Territory, 8000 ft. 
Lophuro7nys rubecula DoUman, Ann. Mag. Nat. Hist. (8) 4: 551, Dec. 1909. 

Elgonyi, Mt. Elgon, Kenya Colony, 7000 ft. 

Lophuromys aquilus chrysopus Osgood 

Lophuromys aquilus chrysopus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 242, 
28 Dec. 1936. Allata, Sidamo, Ethiopia. 

Lophuromys aquilus laticeps Thomas and Wroughton 

Lophuromys laticeps Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 
383, May 1907. Lake Kivu, Congo Beige, 4900 ft. 

Lophuromys aquilus margarett^ Heller 

Lophuromys aquilus margaretlse Heller, Smithsonian Misc. Coll., 59: no. 16, p. 7, 
5 July 1912. Mt. Gargues, Mathews Range, Kenya Colony, 6000 ft. 

Lophuromys aquilus zena Dollman 

Lophuromys zena Dollman, Ann. Mag. Nat. Hist. (8) 4: 550, Dec. 1909. East 
side of Aberdare Range, near Nyeri, Kenya Colony, 9581 ft. 
A recognizable race according to Osgood (Zool. Ser. Field Mus. Nat. Hist., 
20: 243, 28 Dec. 1936). 

Lophuromys brevicaudus Osgood 

Lophuromys brevicaudus Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 241, 
28 Dec. 1936. Mt. Albasso, Chilalo Mts., Arusi, Ethiopia. 

Lophuromys flavopunctatus flavopunctatus Thomas 

Lophuromys flavo-punctatus Thomas, Proc. Zool. Soc. London, p. 14, footnote, 
Jime 1888. Shoa, Abyssinia; "type . . . probably obtained at Ankober, 
about 100 miles N. E. of Addis Ababa" (Thomas, Proc. Zool. Soc. London, 
for 1902, 2: 314, Api. 1903). 



396 bulletin: museum of comparative zoology 

LOPHUROMYS FLAVOPUNCTATTJS BRTJNNEUS ThomaS 

Lophuromys aquilus brunneus Thomas, Ann. Mag. Nat. Hist. (7) 18: 305, Oct. 
1906. Manno, Jimma, Abyssinia, 4200 ft. 
According to Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 240, 18 Dec. 
1936) this is related only to L. flavopundatus and is probably a synonym of 
L. /. zaphiri. 

Lophuromys flavopunctatus simensis Osgood 

Lophuromys flavopundatus simensis Osgood, Zool. Ser. Field Mus. Nat. Hist., 
20: 238, 28 Dec. 1936. Ras Dashan (Mt. Geech), Simien Mts., north- 
eastern Ethiopia. 

Lophuromys flavopunctatus zaphiri Thomas 

Lophuromys zaphiri Thomas, Ann. Mag. Nat. Hist. (7) 18: 304, Oct. 1906. 

Bodeli, Walamo, Abyssinia, 6200 ft. 
Lophuromys flavopundatus zaphiri Osgood, Zool. Ser. Field Mus. Nat. Hist., 

20: 241, 28 Dec. 1936. 

Lophuromys luteogaster Hatt 

Lophuromys luteogaster Hatt, Amer. Mus. Novitates, no. 708, p. 4, 4 Apl. 1934. 
Medje, Ituri district, Congo Beige. 

Lophuromys major Thomas and Wroughton 

Lophuromys major Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 382, 
May 1907. Rwanda, Ubangi River, Congo Beige. 

Lophuromys naso Thomas 

Lophuromys naso Thomas, Ann. Mag. Nat. Hist. (8) 7: 381, Apl. 1911. Gaboon. 

Lophuromys nudicaudus Heller 

Lophuromys nudicaudus Heller, Smithsonian Misc. Coll., 56: no. 17, p. 11, 28 
Feb. 1911. Efulen, Bula Country, Cameroons. 

Lophuromys rita DoUman 

Lophuromys rita DoUman, Ann. Mag. Nat. Hist. (8) 5: 179, Feb. 1910. Lufupa 
River, Katanga, Congo Beige, 4000 ft. 

Lophuromys sikapusi sikapusi (Temminck) 

Mus sikapusi Temminck, Esquisses Zool. sur la Cote de Guin6, p. 160, 1853. 

Dabacrom, Gold Coast. 
Lasiomys afer Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 409, pi., 

f. 5-8, 1866. "Guinea." 
Lophuromys tullbergi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 337, 

Oct. 1911 (new name for Lophuromys afer Tullberg, not Peters). Rio del 

Rey region, near coast of northern Cameroons. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 397 

LoPHUROMYS siKAPUSi ANSORGEi De Winton 

Lophiiromys ansorgei De Winton, Proc. Zool. Soc. London, p. 607, pi. 27, Oct. 
1896. Mumia's, Nyanza Province, Kenya Colony. 

LoPHUROMYS SIKAPUSI MANTEUFELI Matschie 

Lophiiromys sikapusi manteufeli Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 335, Oct. 1911. Mwanza, south coast of Victoria Nyanza, Tanganyika 
Territory. 

LoPHUROMYS SIKAPUSI PYRRHUS Heller 

Lophiiromys pyrrhus Heller, Smithsonian Misc. Coll., 56: no. 17, p. 10, 28 Feb. 
1911. Rhino Camp, Lado Enclave. 

LoPHUROMYS woosNAMi woosNAMi Thomas 

Lophiiromys woosnami Thomas, Ann. Mag. Nat. Hist. (7) 18: 146, Aug. 1906; 
Thomas and Wroughton, Trans. Zool. Soc. London, 19: 512, pi. 24, lower 
f. (col.), Mar. 1910. Mubuku Valley, Ruwenzori East, Uganda, 6000 ft. 

LoPHUROMYS WOOSNAMI PRiTTiEi Thomas 

Lophurornys prittiei Thomas, Ann. Mag. Nat. Hist. (8) 8: 377, Sep. 1911. 
Kigezi, Mfumbiro region, southwestern Uganda. 

Malacomys a. Milne-Edwards 

Malacomys A. Milne-Edwards, Bull. Soc. Philomathique, Paris, (6) 12: pt. 2, 
p. 10, 1877. Genotype Malacomys longipes Milne-Edwards. 

Key to described forms: Hayman, R. W., Proc. Zool. Soc. London, for 1935, 
p. 932, 10 Jan. 1936. 

Malacomys longipes longipes A. Milne-Edwards 

Malacomys longipes A. Milne-Edwards, Bull. Soc. Philomathique, Paris, (6) 
12: pt. 2, p. 10, 1877. Gaboon River. 

Malacomys longipes centralis De Winton 

Malacomys centralis De Winton, Ann. Mag. Nat. Hist. (6) 19: 465, Apl. 1897. 
Monbuttu, Tingasi, Congo Beige. 

Malacomys longipes edwardsi R(Jchebrune 

Malacomys edwardsi Rochebrune, Bull. Soc. Philomathique, Paris, (7) 9: 
87, 1885. Liberia, Mellacoree River. 

Malacomys longipes wilsoni Thomas 

Malacomys wilsoni Thomas, Ann. Mag. Nat. Hist. (8) 18: 238, Aug. 1916, 
Inkongo, Congo Beige. 



398 bulletin: museum of comparative zoology 



Mastomys Thomas. Multimammate Rats 

Mastomys Thomas, Ann. Mag. Nat. Hist. (8) 16: 477, Dec. 1915. As a sub- 
genus of Epimys { = RaUus), type Mus concha A. Smith. 

Mastomys coucha coucha (A. Smith) 

Mus coucha A. Smith, App. to Rept. Exped. for Exploring Central Africa 

from Cape of Good Hope, June 23, 1834, p. 43, 1836. Between Orange 

River and the Tropic, Bechuanaland. 

Mastomys coucha blainei (Wroughton) 

Mus blainei Wroughton, Ann. Mag. Nat. Hist. (7) 20: 502, Dec. 1907. Chak 
Chak, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 
A Mastomys (see Brajstrup, F. W., Vidensk. Medd. fra Dansk Naturh. 
Foren., Kjobenhavn, 99: 117, 11 Sep. 1935). 

Mastomys coucha bradfieldi Roberts 

Mastomys coucha bradfieldi Roberts, Ann. Transvaal Mus., 11: 257, 14 Sep. 

1926. Quickborn, Okahandja, Southwest Africa. 
Mastomys coucha ovamboensis Roberts, Ann. Transvaal Mus., 11: 258, 14 Sep. 

1926. Namutoni, Etosha Pan, Grootfontein district, (not Ovamboland), 

Southwest Africa. 
Synonym of M. c. bradfieldi according to Thomas (Proc. Zool. Soc. London, 
p. 388, 12 July 1927). 

Mastomys coucha cuninghamei (Wroughton) 

Mus cuninghamei Wroughton, Ann. Mag. Nat. Hist. (8) 1: 256, Mar. 1908. 
Chivi Island, Lake Victoria, Kenya Colony. 

Mastomys coucha durum.e (Heller) 

Epimys coucha durumse Heller, Smithsonian Misc. Coll., 59: no. 16, p. 9, 5 
July 1912. Mazer as, Kenya Colony. 

Mastomys coucha effectus (Dollman) 

Epimys effectus Dollman, Ann. Mag. Nat. Hist. (8) 7: 524, May 1911. Lake 
Baringo, Kenya Colony. 

Epimys evelyni Dollman, Ann. Mag. Nat. Hist. (8) 7: 526, May 1911. Lake 
Baringo, Kenya Colony, 4000 ft. Probably a synonym of effectus (see 
HoUister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 87, 1919). 

Mastomys coucha erythroleucus (Temminck) 

Mus erythroleucus Temminck, Esquisses Zool. sur la Cote de Guine, p. 160,^ 
1853. Guinea. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 399' 

Mastomys couch a gambianus (Thomas) 

Epimys gambianus Thomas, Ann. Mag. Nat. Hist. (8) 8: 122, July 1911. 
Gamon, French Gambia. 
A synonym of Mastomys c. erythroleucus Temminck according to Wroughton, 
Ann. Mag. Nat. Hist. (8) 1 : 256, Mar. 1908. 

Mastomys coucha gardulensis (Frick) 

Epimys hildebranti (sic) gardulensis Frick, Ann. Carnegie Mus., 9: 18, 6 June 
1914. Gardula, southern Abyssinia, 4000 ft. 

Mastomys coucha hildebrandtii (Peters) 

Mus hildebrandtii Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 200, 
1878. Ndi, Taita, Kenya Colony. 

Mastomys coucha huberti (Wroughton) 

Mus huberti Wroughton, Ann. Mag. Nat. Hist. (8) 1 : 255, Mar. 1908. Zungeru, 
northern Nigeria. 

Mastomys coucha ismaille (Heller) 

Epimys concha (sic) ismailise Heller, Smithsonian Misc. Coll., 63: no. 7, p. 9, 
24 June 1914. Gondokoro, Uganda. 

Mastomys coucha itigiensis Hatt 

Mastomys coucha itigiensis Hatt, Amer. Mus. Novitates, no. 791, p. 3, 11 
Apl. 1935. Gwao's, near Itigi, Tanganyika Territory. 

Mastomys coucha komatiensis Roberts 

Mastomys coucha komatiensis Roberts, Ann. Transvaal Mus., 11: 259, 14 Sep. 
1926. Arnhemburg, Komati River, Transvaal. 

Mastomys coucha lateralis (Heuglin) 

Mus lateralis Heuglin, Reise in Nordost-Afrika, 2: 71, 1877. Dembea Province, 

Ethiopia. 
Mus tacaziena Heuglin, Reise in Nordost-Afrika, 2: 72, 1877. Takazie River, 

Abyssinia. 
Mastomys coucha lateralis Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 249, 

28 Dec. 1936. 

Mastomys coucha limpopoensis (Roberts) 

Mus limpopoensis Roberts, Ann. Transvaal Mus., 4: 183, 22 Aug. 1914. Sand 
River, northeastern Transvaal. 



400 bulletin: museum of comparative zoology 

Mastomys coucha macrolepis (Sundevall) 

Mus macrolepis Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 
1842, p. 218, 1843. Sennaar, Anglo-Egyptian Sudan. 
Osgood (Zool. Ser. Field Mus. Nat. Hist., 20: 249, 28 Dec. 1936) treats this 

as a species distinct from M. coucha. 

Mus limbatus Wagner, Arch. f. Naturgesch., 11: sect. 1, p. 149, 1845. Sennaar, 
Anglo-Egyptian Sudan. A Mastomys, see Thomas and Hinton, Proc. Zool. 
Soc. London, p. 266, 6 July 1923; Bra?strup, F. W., Vidensk. Medd. fra 
Dansk Naturh. Foren., Kjobenhavn, 99: 117, 11 Sep. 1935. 

Mus kerensis Heuglin, Reise in Nor dost- Afrika, 2: 67, 1877. Keren, Bogos 
country, Abyssinia, in dwellings. 

Epimys azrek Wroughton, Ann. Mag. Nat. Hist. (8) 8: 460, Oct. 1911. Roseires, 
Blue Nile, Anglo-Egyptian Sudan. Based on a young specimen of Mas- 
tomys c. macrolepis. 

Mastomys coucha marikquensis (A. Smith) 

Mus marikquensis A. Smith, App. Rept. Exped. for Exploring Central Africa 

from Cape of Good Hope, June 23, 1834, p. 43, 1836. "Country beyond 

Kurrichaine," Transvaal. 
Mastomys coucha marikquensis Roberts, Ann. Transvaal Mus., 11: 258, 14 Sep. 

1926 (valid race). 
Mus breyeri Roberts, Ann. Transvaal Mus., 5: 120, 28 Aug. 1915. Moorddrift, 

Waterberg district, Transvaal. 
Mastomys coucha breyeri Roberts, Ann. Transvaal Mus., 11: 258, 14 Sep. 1926. 
Mus sodalis Roberts, Ann. Transvaal Mus., 4: 88, 20 Oct. 1913. Wonder- 

fontein, Transvaal. Not Mus socialis Pallas. 
Mastomys coucha sicialis Shortridge, Mamm. South West Africa, 1: 303, 1934 

(misprint). 
According to Roberts (Ann. Transvaal Mus., 11 : 258, 14 Sep. 1926) this is a 
synonym of M. c. marikquensis (A. Smith). 

Mastomys coucha microdon (Peters) 

Mus microdon Peters, Reise nach Mossambique, Saugeth., p. 149, pi. 35, f. 5, 6; 
pi. 36, f. 1, 1852. Tette, Boror, Mozambique. 

Mastomys coucha natalensis (A. Smith) 

Mus natalensis A. Smith, Illustr. Zool. South Africa, Mamm., pi. 47, f. 2, Oct. 

1847. Port Natal, Natal. 
Mus illovoensis Jentink, Zool. Jahrb., Syst., 28: 248, text-f. L, 15 Oct. 1909. 

Lower Illovo, Natal. 
Mus coucha zuluensis Thomas and Schwann, Proc. Zool. Soc. London, 1: 268, 

10 Aug. 1905. LTmvolosi Station, Zululand, Natal. 
Probably same as M. c. microdon (see Thomas and Wroughton, Proc. Zool. 
Soc. London, p. 546, 30 Oct. 1908), in which case natalensis supersedes. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 401 

Mastomys coucha neumanni (Heller) 

Epimys coucha neumani (sic) Heller. Smithsonian Misc. Coll., 59: no. 16, p. 8, 
5 July 1912. Neumann's Boma, Northern Guaso Nyiro, Kenya Colony. 

Mastomys coucha pallida (Dollman) 

Epimys coucha pallida Dollman, Abstr. Proc. Zool. Soc. London, no. 131, p. 25, 

14 Apl. 1914; Proc. Zool. Soc. London, p. 314, 16 June 1914. Kamchuru, 

Lobor, Central Province, Uganda. 

Mastomys coucha panya (Heller) 

Epimys panya Heller, Smithsonian Misc. Coll., 56: no. 9, p. 2, pi. 2, 3 figs., 22 
July 1910. Juja Farm, Athi Plains, Kenya Colony. 

Mastomys coucha peregrinus (De Winton) 

Mus peregrinus De Winton, Proc. Zool. Soc. London, for 1897, p. 959, 1 Apl. 

1898. Ras-el-Ain, Haha, Morocco. 
For figure of skull, see Cabrera, A., Los mamiferos de Marruecos, in Trab. 
Mus. Nac. de Cienc. Nat., Madrid, zool. ser., no. 57, p. 270, text-f. 32, 30 Dec. 
1932. 
Mus calopus Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 6: 365, July 

1906. Mogador, Morocco. 

Mastomys coucha silaceus (Wagner) 

Mus silaceus Wagner, Arch. f. Naturgesch., 8: sect. 1, p. 11, 1842. Cape of 

Good Hope. 
Mastomys coucha silaceus Roberts, Ann. Transvaal Mus., 11: 257, 14 Sep. 1926. 

A valid race from the Albany district, Cape of Good Hope. 

Mastomys coucha tinctus (Hollister) 

Rattus coucha tinctus HoUister, Smithsonian Misc. Coll., 68: no. 10, p. 1, 16 Jan. 
1918. Kaimosi, Kenya Colony. 

Mastomys coucha ugand^e (De Winton) 

Mus ugandse De Winton, Ann. Mag. Nat. Hist. (6) 20: 317, Sep. 1897. Entebbe, 

Uganda. 
Rattus somereni Kershaw, Ann. Mag. Nat. Hist. (9) 11: 594, May 1923. Ka- 

baroni, Bukedi district, western foothills of Mt. Elgon, eastern Uganda, 

7000 ft. 
Range: Lado to northern Victoria Nyanza. 

Mastomys coucha victoria (Matschie) 

Mus (Epimys) microdon victorix Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 342, Oct. 1911. Mwanza, Speke Gulf, Lake Victoria Tanganyika Terri- 
tory. 



402 bulletin: museum of comparative zoology 

Mastomys rufidorsalis (Heuglin) 

Mus (?) rufidorsalis Heuglin, Reise in Nordost-Afrika, 2: 70, 1877. Simen and 

Wogara, Abyssinia. An Arvicanthis according to Trouessart (Cat. Mamm. 

Viv. Foss., Suppl., p. 375, 1904). 

Muriculus Thomas 

Muriculus Thomas, Proc. Zool. Soc. London, for 1902, 2: 314, Apl. 1903. 
Genotype Mus imberbis Riippell. 

Muriculus imberbis imberbis (Riippell) 

Mus imberbis Riippell, Mus. Senckenbergianum, 3: 110, pi. 6, f. 4, 1842. Simen, 
Ethiopia, 3300 m. 

Muriculus imberbis chilaloensis Osgood 

Muriculus imberbis chilaloensis Osgood, Zool. Ser. Field Mus. Nat. Hist., 20: 
245, 28 Dec. 1936. Mt. Albasso, northeastern Chilalo Mts., Arusi, Ethio- 
pia. 

Mus Linnaeus. House Mice 

Mus Linnseus, Syst. Nat., ed. 10, 1: 59, 1758. Genotype Mus musculus Lin- 
naeus. 

Musculus Rafinesque, Precis d. Decouvertes Somiologiques, p. 13, 1814. Sub- 
stitute for Mus. 

Drymomys Tschudi, Fauna Peruana, Therologie, p. 178, pi. 13, f. 1, 1844. 
Genotype Drymomys parvulus Tschudi = Mus musculus Linnseus. 

Mus musculus musculus Linnseus. House Mouse 

Mus rnusculus Linnseus, Syst. Nat., ed. 10, 1: 62, 1758. Sweden. 

Mus orientalis Cretzschmar, Rlippell's Atlas z. d. Reise im Nordliche Afrika, 

p. 76, pi. 30a, 1826 (not of Desmarest, Nouv. Diet. d'Hist. Nat., ed. 2, 29: 

59, 1819, which is Mus striatus Linnseus, 1758). Massawa, Ethiopia. 
M[us] modestus Wagner, Arch. f. Naturg., 8: sect. 1, p. 14, 1842; Schreber's 

Saugthiere, Suppl., 3: 432, pi. 181 A, 1843. Cape of Good Hope. 
Mus vignaudii Demurs and Prevost, in Lefebvre's Voy. en Abyssinie, Atlas, 

Zool., pi. 5, f. 2, 1850. Abyssinia. 
Mus pallesceus (sic) Heuglin, Reise in Nord-Ost Afrika, 2: 72, 1877. Northern 

Sennaar. Synonym of Mus musculus orientalis according to Thomas. 

Mus musculus brevirostris Waterhouse 

Mus brevirostris Waterhouse, Proc. Zool. Soc. London, p. 19, 21 Nov. 1837. 

Maldonado, Uruguay (introduced from Spain to Morocco). 
Mus azoricus Schinz, Syst. Verzeichn. Sjiugeth., 2: 161, 1845. In insulis 

Azoricis. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 403 

[Mus musculus, var.] flavescens Lataste, Actes Soc. Linn, de Bordeaux, (4) 7 = 

37: 21, 1883 (not of Fitzinger). Southern France. 
Mus musculus brevirostris Trouessart, Cat. Mamm. Viv. Foss., ed. 2, p. 380, 

1904. Cabrera, Trab. Mus. Nac. de Cienc. Nat., Madrid, zool. ser., no. 57, 

p. 255, 30 Dec. 1932. 
Range: Northern Morocco. 

Mus MUSCULUS FAR Cabrera 

Mus musculus Jar Cabrera, Real Soc. Espanola Hist. Nat., Madrid, 50th-anniv. 
vol., p. 46, 1921. Mogador, Morocco. 

Mus MUSCULUS QENTiLis Brants. White-bellied House Mouse 
Mus gentilis Brants, Het Geslacht der Muizen, p. 126, 1827. Egypt and Nubia. 
Mus musculus candidus Laurent, Bull. Soc. Sci. Nat. Moroc, 17: 1, 31 Mar. 
1937. Berguent, eastern Morocco. 

Mus BACTRiANUS LYNESi Cabrera 

Mus spicilegus lynesi Cabrera, Bol. Real Soc. Espaiiola Hist. Nat., Madrid, 23: 
430, 31 Dec. 1923. Tazarot, Yebala, northeastern Morocco. 
Mus spicilegus (type locality Hungary) is but a race of Mus bactrianus 
(type locality Kandahar, Afghanistan). 

Mus BACTRIANUS MOGREBiNUS Cabrera 

Mus spicilegus mogrebinus Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 
11: 555, Dec. 1911. Taguidert, Haha Province, Morocco. 

Mus BACTRIANUS RiFENSis Cabrera 

Mus spicilegus rifensis Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 
23: 431, 31 Dec. 1923. Melilla, eastern Rif, Morocco. 



Mylomys Thomas 

Mylomys Thomas, Ann. Mag. Nat. Hist. (7) 18: 224, Sep. 1906. Genotype 
Mylomys cuninghamei Thomas. 

Mylomys alberti Thomas 

Mylomys alberti Thomas, Ann. Mag. Nat. Hist. (8) 16: 148, Aug. 1915. Poko,. 
upper Uele River, Congo Beige. 

Mylomys christyi Thomas 

Mylomys christyi Thomas, Ann. Mag. Nat. Hist. (8) 20: 362, Nov. 1917. Mt 
Baginzi, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 



404 bulletin: museum of comparative zoology 

Mylomys cxjninghamei cuninghamei Thomas 

Mylomys cuninghamei Thomas, Ann. Mag. Nat. Hist. (7) 18: 225, Sep. 1906. 
East of Aberdare Mts., Kenya Colony, 4480 ft. 
Probably the various described "species" are all geographic races of this. 

Mylomys cuninghamei lowei Haj^man 

Mylomys cuninghamei lowei Hayman, Proc. Zool. Soc. London, for 1935, p. 934, 
10 Jan. 1936. Wenchi, Ashanti, Gold Coast. 

Mylomys cuninghamei massaicus Lonnberg 

Mylomys cunninghamei (sic) massaicus Lonnberg, Arkiv f. Zool., 10: no. 12, 
p. 8, 19 May 1916. Donya Sabuk, Kenya Colony. 

Mylomys lutescens Thomas 

Mylomys lutescens Thomas, Ann. Mag. Nat. Hist. (8) 16: 149, Aug. 1915. 
Nalasanji, southwestern Uganda, 5000 ft. 

Mylomys roosevelti (Heller) 

Pelomys roosevelti Heller, Smithsonian Misc. Coll., 54: no. 1924, p. 1, pi. 1, a, 

28 Feb. 1910. Nzoia River, Guas Ngishu Plateau, Kenya Colony. 
Mylomys roosevelti Thomas, Ann. Mag. Nat. Hist. (8) 20: 363, Nov. 1917. 



Myomys Thomas 

Myomys Thomas, Ann. Mag. Nat. Hist. (8) 16: 477, Dec. 1915. As a subgenus 
of Epimys {=Rattus), type Epimys colonus (A. Smith) =Af us colonus 
Brants. 

Myomys albipes albipes (Riippell) 

Mus albipes Riippell, Mus. Senckenbergianum, 3: 107, pi. 6, f. 2, 1842. Mas- 
saw a, Abyssinia ( = Ethiopia) . 

Epimys rufidorsalis ankoberensis Frick, Ann. Carnegie Mus., 9: 18, 6 June 1914. 
Ankober, Shoa, Abyssinia, 7500 ft. 
Probably a synonym of Myomys albipes (see Osgood, Zool. Ser. Field Mus. 

Nat. Hist., 20: 250, 28 Dec. 1936). 

Epimys rufidorsalis alettensis Frick, Ann. Carnegie Mus., 9: 17, 6 June 1914. 
Aletta, Sidamo, southern Abyssinia, 6000 ft. 
Probably a synonym of Myomys albipes (see Osgood, Zool. Ser. Field Mus. 

Nat. Hist., 20: 250, 28 Dec. 1936). 

Myomys albipes fuscirostris (Wagner) 

Mus fuscirostris Wagner, Arch. f. Naturgesch., 11: sect. 1, p. 149, 1845. Sen- 

naar, Anglo-Egyptian Sudan. 
Mus albipes fuscirostris Trouessart, Cat. Mamm. Viv. Foss., pt. 3, p. 481, 1897. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 405 

Myomys angolensis (Bocage) 

Mus angolensis Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 2: 12, Sep. 
1890. Capangombe, interior of Mossamedes, Angola. 

Myomys avunculus avunculus (Thomas) 

Mus avunculus Thomas, Ann. Mag. Nat. Hist. (7) 13: 417, June 1904. Pungo 

Andongo, Angola. 
Mthomys avunculus Hill and Carter, Amer. Mus. Novitates, no. 913, p. 3, 8 

Mar. 1937. 

Myomys avunculus phippsi (Hill and Carter) 

.■Ethomys avunculus phippsi Hill and Carter, Amer. Mus. Novitates, no. 913, p.. 
3, f. 2, 8 Mar. 1937. Humpata, Angola, 6300 ft. 

Myomys brockmani (Thomas) 

Mus brockmani Thomas, Ann. Mag. Nat. Hist. (7) 18: 298, Oct. 1908. Upper 
Sheikh, British Somaliland. 

Myomys colonus colonus (Brants) 

Mus colonus Brants, Het Geslacht der Muizen, p. 124, 1827. Algoa Bay, South 
Africa, Cape of Good Hope. 

Myomys colonus shortridgbi St. Leger 

Myomys shortridgei St. Leger, Proc. Zool. Soc. London, p. 411, 30 June 1933. 
Okavango-Omatako junction, Grootfontein district, Southwest Africa. 

Myomys daltoni daltoni (Thomas) 

Mus daltoni Thomas, Ann. Mag. Nat. Hist. (6) 10: 181, Aug. 1892. West 
Africa, probably the Island of Fernando Po. 

Myomys daltoni saturatus Ingoldby 

Myomys daltoni saturatus Ingoldby, Ann. Mag. Nat. Hist. (10) 3: 520, May 
1929. Near Kintampo, northern Ashanti, Gold Coast. 

Myomys fumatus pumatus (Peters). Rock Rat 

Mus fumatus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 200, 1878. 

Ukamba, Kenya Colony. 
Rattus fumatus fumatus Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 82, 

1919 (with notes on the type and synonymy). 
Mus niveiventris Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 12, 16 

Feb. 1910. Voi, Kenya Colony. 
Epimys niveiventris ulse Heller, Smithsonian Misc. Coll., 56: no. 9, p. 3, pi. 2, 

3 figs., 22 July 1910. Ulukenia Hills, Kenya Colony. 

Myomys fumatus subfuscus (Osgood) 

Mus niveiventris subfuscus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 
12, 16 Feb. 1910. Lake Elmenteita, Kenya Colony. 



406 bulletin: museum of comparative zoology 

Myomys fumatus tana (True) 

Mus tana True, Proc. U. S. Nat. Mus., 16: 602, 25 Oct. 1893. Tana River, 
between the coast and Hameye, Kenya Colony. 
On the status of this rat see Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, 
p. 86, 1919. 

Myomys granti (Wroughton) 

Mus granti Wroughton, Ann. Mag. Nat. Hist. (8) 1 : 257, Mar. 1908. Deelfon- 
tein, Cape of Good Hope. 

Myomys pernanus (Kershaw) 

Rattus pernanus Kershaw, Ann. Mag. Nat. Hist. (9) 8: 568, Nov. 1921. Amala 

(or Mara) River, rising in the Sotik, Kenya Colony, and entering Lake 

Victoria at 1° 30' S. 

Myomys verroxii (A. Smith) 

Mus verroxii A. Smith, South African Quart. Journ., 2: 156, 1834. Near Cape 

Town, Cape of Good Hope. 
Mus verreauxi W. L. Sdater, Mamm. South Africa, 2: 45, 1901 (emendation). 



Nesokia Gray. Bandicoots 

Nesokia Gray, Ann. Mag. Nat. Hist. (1) 10: 264, Dec. 1842. Genotype Mus 
hardwickii Gray ,= Arvicola indica Gray and Hardwicke. (See Thomas, 
Ann. Mag. Nat. Hist. (7) 20: 203, Sep. 1907.) 

Nesocia Blanford, Mamm. British India, p. 421, 1891, emendation. 

Nesokia suilla Thomas 

Nesokia suilla Thomas, Ann. Mag. Nat. Hist. (7) 20: 203, Sep. 1907. Shaluf, 

Suez, Egypt. 
Nesokia bacheri Anderson and De Winton, Zool. Egypt, Mamm., p. 286, pi. 50, 

1902 (in part). 
Hange: Lower Egypt, west to Fayum and extreme west of the delta. 



Nilopegamys Osgood 

Nilopegamys Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 12: 185, 21 Nov. 
1928. Genotjqje Nilopegamys plumbeus Osgood. 

Nilopegamys plumbeus Osgood. Water Rat 

Nilopegamys plumbeus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 12: 185, 

pi. 15, text-f. 1 (skull), 21 Nov. 1928. Tributary of Little Abbai, between 

Sakalla and Njabarra, Gojjam, Abyssinia, 8500 ft. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 407 



OcHROMYS Thomas 

Ochromys Thomas, Ann. Mag. Nat. Hist. (9) 5: 142, Jan. 1920. Genotype Mus 
woosnami Schwann. 

Ochromys woosnami (Schwann) 

Mus woosnami Schwann, Proc. Zool. See. London, 1 : 108, pi. 6, 7 June 1906. 
Molopo, Bechuanaland. 



Oenomys Thomas. Rufous-nosed Rats 

Oenomys Thomas, Ann. Mag. Nat. Hist. (7) 13: 416, June 1904. Genotype 

Mus hypoxanthus Pucheran. 
Aenomys Kershaw, Revue Zool. Africaine, 11: 365, 27 Dec. 1923; misprint 

for Oenomys. 

See Thomas and Wroughton, Trans. Zool. Soc. London, 19: 510, Mar. 1910, 
for key. 

Oenomys hypoxanthus hypoxanthus (Pucheran) 

Mus hypoxanthus Pucheran, Revue et Mag. de Zool. (2) 7: 206, Apl. 1855. 

Gaboon. 
Mus rufinus Matschie, Deutsch-Ost-Afrika, 3: pt. 1, Saugeth., p. 52, 1895, not 

of Temminck, 1855. Ukondjo, Tanganyika. 

Oenomys hypoxanthus anchiet^ (Bocage) 

Mus anchietse Bocage, Jorn. Sci. Math., Phys. e Nat., Lisboa, (2) 2: 11, pi., 
f. 3, 3a, Sep. 1890 (the description of the male only). Ambacca, Angola. 

Oenomys hypoxanthus bacchante (Thomas) 

Mus hypoxanthus bacchante Thomas, Ann. Mag. Nat. Hist. (7) 12: 342, Sep. 

1903. Nandi Station, edge of Nandi Escarpment, Kenya Colony (see 

Hollister, N., Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 64, 1919). 
Oenomys bacchante maerens Thomas, Ann. Mag. Nat. Hist. (8) 7: 379, Apl. 

1911. Solai, west slope of Mt. Kenya, Kenya Colony, 8000 ft. Probably 
an aberrant individual of Oe. h. bacchante. 
Oenomys oris Thomas, Ann. Mag. Nat. Hist. (8) 7: 380, Apl. 1911. Mt. Kin- 

angop, Aberdare Range, Kenya Colony, 11,000 ft. (Probably based on an 

unusually large individual.) 

Oenomys hypoxanthus editus Thomas and Wroughton 

Oenomys bacchante editus Thomas and Wroughton, Trans. Zool. Soc. London, 
19: 509, Mar. 1910. Mubuku Valley, Ruwenzori East, Uganda. Accord- 
ing to Hollister, may be indistinguishable from Oe. h. unyori. 



408 bulletin: museum of comparative zoology 

Oenomys hypoxanthus unyori (Thomas) 

Mus hypoxanthus unyori Thomas, Ann. Mag. Nat. Hist. (7) 12: 342, Sep. 1903. 

Fadjas on Victoria Nile, Unyoro. 
Mus unyoroi Trouessart, Cat. Mamm. Viv. Foss., ed. 2, p. 375, 1904. 

Oenomys hypoxanthus vallicola Heller 

Oenomys hypoxanthus vallicola Heller, Smithsonian Misc. Coll., 63: no. 7, 

p. 11, 24 June 1914. Lake Naivasha, Kenya Colony. 
Oenomys bacchante vallicola HoUister, Bull. U. S. Nat. Mus., no. 99, pt. 2, 

p. 66, 1919. 

Oenomys oenatus Thomas 

Oenomys ornatus Thomas, Ann. Mag. Nat. Hist. (8) 7: 378, Apl. 1911. Bibi- 
anaha, near Dunkwa, Gold Coast. 



Pelomys Peters. Creek Rats 

Pelomys Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 275, May 1852 
(as a subgenus of Mus); Reise nach Mossambique, Saugeth., p. 157, pi. 
33, f.3; pi. 35, f.9, 1852 (as a genus). Type species Mus (Pelomys) fallax 
Peters. 

Subgenus Pelomys Peters 

Pelomys campan^ (Huet) 

Golunda campanse Huet, Le NaturaUste (2) 10: no. 31, p. 143, text-f. 1-3, 
15 June 1888. Landana, west coast of Angola. 

Pelomys dybowskii (Pousargues) 

Golunda dybowskii Pousargues, Bull. Soc. Zool. de France, 18: 163, 1893. 
Kemo River, French Congo, 6° 17' N., 17° 15' E. 

Pelomys fallax fallax (Peters) 

Mus (Pelomys) fallax Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 

275, May 1852. Caya district, Zambesi River, and Boror, Licuare River, 

Mozambique. 

Pelomys fallax australis Roberts 

Pelomys australis Roberts, Ann. Transvaal Mus., 4: 90, 20 Oct. 1913. Mazam- 
beti, Beira, Mozambique. 

Pelomys fallax concolor Heller 

Pelomys fallax concolor Heller, Smithsonian Misc. Coll., 59: no. 16, p. 13, 5 
July 1912. Kiduha, Lake Mutanda, Uganda. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 409 

Pelomys fallax insignatus Osgood 

Pelomys fallax insignatus Osgood, Ann. Mag. Nat. Hist. (8) 5: 276, Mar. 
1910. Fort Hill, northern Nyasaland. 

Pelomys fallax iridescens Heller 

Pelomys fallax iridescens Heller, Smithsonian Misc. Coll., 59: no. 16, p. 12, 
5 July 1912. Mt. Mbololo, Taita Mts., Kenya Colony. 

Pelomys fallax rhodesi^ Roberts 

Pelomys fallax rhodesiae Roberts, Ann. Transvaal Mus., 13: 118, 6 July 1929. 
Machile River, northwestern Rhodesia. 

Pelomys frater Thomas 

Pelomys frater Thomas, Ann. Mag. Nat. Hist. (7) 13: 415, June 1904. Brag- 
anza, Angola. 

Pelomys lultj^ Matschie 

Pelomys luluse Matschie, Zeitschr. f. Saugetierk., 1: 113, 24 Dec. 1926. Ditapa, 
near Luluabourg, Lulua River, Congo Beige. 



Subgenus Komemys de Beaux 

Koynemys de Beaux, Ann. Mus. Civ. Stor. Nat. Genova, 51: 207, 12 Apl. 1924. 
Genotype Komemys isseli de Beaux. 
According to Hatt (Amer. Mus. Novitates, no. 790, p. 4, 10 Apl. 1935) this 
"cannot at best be considered more than a subgenus." 

Pelomys isseli (de Beaux) 

Komemys isseli de Beaux, Ann. Mus. Civ. Stor. Nat. Genova, 51 : 207, 12 Apl. 
1924. Kome Island, Lake Victoria, Uganda. 

Pelomys minor Cabrera and Ruxton 

Pelomys minor Cabrera and Ruxton, Ann. Mag. Nat. Hist. (9) 17: 601, May 

1926. St. Joseph de Luluabourg, Congo Beige. 
Pelomys (Komemys) minor Hatt, Amer. Mus. Novitates, no. 790, p. 4, 10 

Apl. 1935. 

Praomys Thomas. Short-haired Rats 

Praomys Thomas, Ann. Mag. Nat. Hist. (8) 15: 477, Dec. 1915. As a subgenus 
of Epimys ( =RaUus), type Epimys tullbergi Thomas. 
Redefined as a more circumscribed genus in Ann. Mag. Nat. Hist. (9) 17: 
178, Jan. 1926, with fist of included species. 



410 bulletin: museum of comparative zoology 

Praomys jacksoni jacksoni (De Winton) 

Musjacksoni De Winton, Ann. Mag. Nat. Hist. (6) 20: 318, Sep. 1897. Entebbe, 
Uganda. 

Praomys jacksoni delectorum (Thomas) 

Epimys delectorum Thomas, Ann. Mag. Nat. Hist. (8) 6: 430, Oct. 1910. 
Mlanji Plateau, southern Nyasaland, 5500 ft. 

Praomys jacksoni melanotus G. M. Allen and Loveridge 

Praomys tullbergi melanotus G. M.Allen and Loveridge, Bull. Mus. Comp. Zool., 

75: 106, Feb. 1933. Nyamwanga, Poroto Mts., northwest end of Lake 

Nyasa, Tanganyika Territory. 

Praomys jacksoni minor Hatt 

Praomys jacksoni minor Hatt, Amer. Mus. Novitates, no. 708, p. 11,4 Apl. 1934; 
Lukolela, middle Congo, Congo Beige. 

Praomys jacksoni montis (Thomas and Wroughton) 

Mus jacksoni montis Thomas and Wrougliton, Trans. Zool. Soc. London, 19: 
503, Mar. 1910. Mubuku Valley, Ruwenzori East, Uganda, 6000 ft. 

Praomys jacksoni peromyscus (Heller) 

Mus -peromyscus Heller, Smithsonian Misc. Coll., 52: 472, pi. 56, fig. b, 13 Nov. 

1909. Njoro O Nyiro, Sotik, Kenya Colony. 
Rattus tullbergi peromyscus Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 77, 

1919. 

Praomys jacksoni viator (Thomas) 

Epimys jacksoni viator Thomas, Ann. Mag. Nat. Hist. (8) 7: 461, May 1911. 
Panyam, northern Nigeria, 4000 ft. 

Praomys morio (Trouessart) 

Mus maura Gray, Proc. Zool. Soc. London, p. 181, Sep. 1862 (not Mus ynaurus 
Waterhouse; not Tullberg). Cameroon Mountains, Cameroons, 7000 ft. 

Mus morio Trouessart, Bull. Soc. d'Etudes Sci., Angers, 10: 121, 1881. Cam- 
eroon Mts., Cameroons. 
New name for Mus maurus Gray, not Waterhouse. 

Praomys tait^ (Heller) 

Epi?mjs taitse Heller, Smithsonian Misc. Coll., 59: no. 16, p. 9, 5 July 1912. 
Mt. Mbololo, Taita Hills, Kenya Colony, 5000 ft. 

Praomys tullbergi tullbergi (Thomas) 

Mus burtoni Thomas, Ann. Mag. Nat. Hist. (6) 10: 182, Aug. 1892. Ankober 

River, Wasa, Ashanti. Not Mus burtoni Ramsay, 1887, an Australian 

species. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 411 

Mus tullbergi Thomas, Ann. Mag. Nat. Hist. (6) 13: 205, Feb. 1894. New name 
for Mus hurloni Thomas, not Ramsay, 1887. 
A species distinct from P. jacksoni (see Hatt, R. T., Amer. Mus. Novitates, 
no. 708, p. 13, 4 Apl. 1934). 

Praomys tullbergi ltjkolel^ Hatt 

Praomys tullbergi lukolelae Hatt, Amer. Mus. Novitates, no. 708, p. 13, 4 Apl. 
1934. Lukolela, middle Congo, Congo Beige. 

Praomys tullbergi rostratus (Miller) 

Mus tullbergi rostratus Miller, Proc. Washington Acad. Sci., 2: 637, 28 Dec. 
1900. Mount Coffee, Liberia. 



Rattus Fischer. Typical Rats 

Ruttus (misprint for Rattus) Fischer, National-mus.Naturg. Paris, 2: 128, 1803. 
Genotj^e Mus norvegicus Erxleben. 

See HoUister, N., Proc. Biol. Soc. Washington, 29: 126, 6 June 1916, as to 
genotype. 



Subgenus Epimys Trouessart. Roof Rats 

Epimys Trouessart, Cat. Mamm. Viv. et Foss., in Bull. Soc. d'Etudes Sci. 
d'Angers, 10: 117, 1881. Subgenus of Mus; type Mus rattus Linnaeus. 

Rattus auratus (G. Grandidier) 

Mus auratus G. Grandidier, Bull. Mus. National d'Hist. Nat. Paris, (1) 5: 277, 
1899. Near Morondava, west coast of Madagascar. 
Probably an introduced rat of the Rattus rattus group. 

Rattus rattus rattus (Linnaeus). Black Rat 

Mris rattus Linnajus, Syst. Nat., ed. 10, 1: 61, 1758. Sweden. 

Mus tectorum var.fuliginosus Bonaparte, Iconogr. Fauna Ital., 1 : fasc. 3, pi. 22, 

f. 1, 1833. Italy. 
Rattus domesticus Fitzinger, Sitzb. K. Akad. Wiss., Wien, math.-nat. CI., 56: 

pt. 1, p. 64, 1867. Germany. Lists also the varieties /asciis, varius, ful- 

vaster, albus and ater, now regarded as synonyms. 
Mus rattus ater Millais, Zoologist, (4) 9: 205, pi. 4, 15 June 1905. London, 

England. 
Range: Introduced (?) in Morocco (see Laurent, P., Bull. Soc. Hist. Nat. 

de I'Afrique du Nord, 27: 165-172, May 1936); or may be a melanistic 

variant of the native R. r. alexandrinus. 



412 bulletin: museum of comparative zoology 

Rattus rattus alexandrinus (E. Geoff roy and Audouin). Alexandrine Rat 
Mas alexandrinus E. Geoffroy and Audouin, Description de I'EgjTjte, 2: 733, 

Atlas, pi. 5, f. 1, 1829. Alexandria, Egypt. 
Mus alexandrinus var. Mus tettensis Peters, Reise nach Mossambique, Saugeth., 

p. 156, 1852. Tette, Mozambique. 
Doubtless a synonym of one of the introduced races of Rattus rattus. 
Mus samharensis Heuglin, Reise in Nordost-Afrika, 2: 67, 1877. Mekulu, 

inland from Massawa, Eritrea, a house rat. = Rattus alexandrinus? 

Rattus rattus frugivorus (Rafinesque). Cream-bellied House Rat 
Musculus frugivorus Rafinesque, Precis des Decouvertes et Travaux Somio- 

logiques, p. 13, 1814. Sicily. Probably originally from northwestern India, 

Indus Valley (see Schwarz, E., Nature, London, 133: 573, 14 Apl. 1934; 

Proc. Zool. Soc. London, p. 723-726, 1934). 
Mus flavigaster Heuglin, Nova Acta Akad. Ca;s. Leop.-Carol., Halle, 20: no. 10, 

p. 4, pi. 2, fig. 2, 1861. Error for Mus flaviventris as shown by reference. 

Rattus rattus rufescens (Gray) 

Mus rufescens Gray, Charlesworth's Mag. Nat. Hist., 1 : 585, Nov. 1837. India. 
Introduced on Zanzibar (see Schwarz, E., Nature, London, 133: 573, 14 Apl. 
1934). 

Rattus rattus kijabius (J. A. Allen) 

Mus kijabius J. A. Allen, Bull. Amer. Mus. Nat. Hist., 26: 169, 19 Mar. 1909. 

Kijabe, Kenya Colony. This rat may have been introduced from India; 

specimens are "to a certain extent intermediate in color between" R. r. 

rattus and R. r. alexandrinus (Hollister, N., Bull. U.S. Nat. Mus., no. 99, 

pt. 2, p. 68, 1919). 
Mvs (Epimys?) muansas Matschie, Sitzb. Ges. Naturf. Freunde, Berhn, p. 340, 

Oct. 1911. Mwanza, on Lake Victoria, Tanganyika Territory. 
Mus (Epimys) rattiforrnis Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 98, 

Apl. 1915. Amani, Usambara, Tanganyika Territory. 
Epimys jujensis Lonnberg, Arkiv f. Zool., 10: no. 12, p. 10, 19 May 1916. Juja 

Farm, Kenya Colony. See Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, 

p. 68, 1919. 

Rattus rattus nericola Cabrera 

Rattus rattus nericola Cabrera, Real Soc. Espanola Hist. Nat., Madrid, 50th- 
anniv. vol., p. 54, 1921. Wad Saf-saf, lower Muluya, Morocco. 
Range: Northern Morocco, Yebala and the Rif. 

Rattus rattus sueirensis Cabrera 

Rattus rattus chionogaster Cabrera, Real Soc. Espanola Hist. Nat., Madrid, 50th- 
anniv. vol., p. 51, 1921. Mogador, Morocco. Not Epimys {=Rattus) 
chionogaster Lonnberg and Mjoberg, 1916, an Australian species. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 413 

Rattus rallus sueirensis Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 
21: 159, 25 July 1921. Environs of Mogador, Morocco. New name for 
Rattus rattus chionogaster Cabrera, not Lonnberg and Mjoberg. 

Rattus rattus wroughtoni Hinton 

Rattus rattus wroughtoni Hinton, Journ. Bombay Nat. Hist. Soc, 26: 384, 20 
May 1919. Coonoor, Nilgiri Hills, India. Introduced into Zanzibar (see 
Schwarz, E., Nature, London, 133: 573, 14 Apl. 1934). 



Subgenus Rattus Fischer. Brown Rats 

Rattus norvegicus norvegicus (Berkenhout). Norway Rat 

Mus norvegicus Berkenhout, Outl. Nat. Hist. Great Britain, 1: 5, 1769. Norway. 

Mus norvegicus Erxleben, Syst. Regn. Anim., 1: 381, 1777. Norway. 

Mus decumanus Pallas, Novse Spec. Quadrup. e Glir. Ord., p. 91, 1778. "Cum 
alls Indiarum noxis in Europeam . . . invecta." 

Mus sxcrmulottus Severinus, Tentamen Zool. Hungar., p. 73, 1779. 

Mus hibernicus Thompson, Proc. Zool. Soc. London, p. 52, 20 Dec. 1837, 
Rathfriland, County Down, Ireland. 

Mus leucosternwm Riippell, Mus. Senckenbergianum, 3: 108, pi. 7, f. 2, 1842. 
Massawa, Eritrea, in houses. 
See Mertens, R., Senckenbergiana, 7: 25, 12 Feb. 1925. 

Mus decaryi G. Grandidier, Bull. Mus. National d'Hist. Nat. Paris, (2) 6: 478, 
text-f. 1-3, Nov. 1934. Two or three kilometers from Tamatave, Mad- 
agascar, This is apparently an individual oi Rattus norvegicus (Berkenhout) . 
Introduced in the Nile Valley to Khartoum (Flower, S.S., Proc. Zool. Soc. 

London, p. 410, 8 July 1932); also in Fernando Po (Cabrera, A., Mem.. Real 

Soc. Espaiiola Hist. Nat., Madrid, 16: 87, 10 July 1929). 



Rhabdomys Thomas. Four-striped Grass Mice 

Rhabdomys Thomas, Ann. Mag. Nat. Hist. (8) 18: 69, July 1916. Genotype 
Mus pumilio Sparrman. 

Review: Wroughton, R. C, Notes on the various forms of Arvicanthis pumilio 
Sparrm. Ann. Mag. Nat. Hist. (7) 16: 629-639, Dec. 1905. 

Rhabdomys pumilio pumilio (Sparrman) 

Mus pumilio Sparrman, Kungl. Svehska Vet.-Akad. Handl., Stockholm, p. 236, 

pi. 6, 1784. Sitzicamma forest, on Snake River, east of Cape of Good Hope. 
Mus donavani (sic) Lesson, Manuel de Mammalogie, p. 268, 1827 (based on 

Donovan, Mag. du Naturaliste, pt. 26). Cape of Good Hope. 
M{us\ pumilio var. major Brants, Het Geslacht der Muizen, p. 105, 1827. Based 

on account in Desmarest's Mammalogie; inland from Cape of Good Hope. 
Mus lineatus F. Cuvier, Hist. Nat. des Mammiferes, pi. 161 and text, 1829. 



414 bulletin: museum of comparative zoology 

M[us\ vittatus Wagner, Arch. f. Naturg., 8: sect. 1, p. 11, 1842. Cape of Good 

Hope. 
Mus septemvittatus Schinz, Syst. Verzeichn. d. Saugeth., 2: 155, 1845. Cape of 

Good Hope. 

Rhabdomys pumilio angoljE (Wroughton) 

Arvicanthis pumilio angolse Wroughton, Ann. Mag. Nat. Hist. (7) 16: 636, 
Dec. 1905. Caconda, Angola, 4700 ft. 

Rhabdomys pumilio BECHUANiE (Thomas) 

Isomys p[u7nilio] bechuanx Thomas, Proc. Zool. Soc. London, for 1892, p. 
551, Apl. 1893. Bechuanaland; but Shortridge (Mamm. South West 
Africa, 1: 280, 1934), considers it advisable to fix the type locality as 
Rooibank, near Walvis Bay, Southwest Africa. 

Rhabdomys pumilio namibensis Roberts, Ann. TransvaalMus., 11: 255, 14 Sep. 
1926. Swakopmund, Southwest Africa. Shortridge (Mamm. South West 
Africa, 1: 280, 1934) regards this as a synonym of R. p. bechuanae. 

Rhabdomys pumilio chak^ (Wroughton) 

Arvicanthis pumilio chakse Wroughton, Ann. Mag. Nat. Hist. (7) 16: 636, Dec. 
1905. Sibudeni, Zululand, Natal. 

Rhabdomys pumilio cinereus (Thomas and Schwann) 

Arvicanthis pumilio cinereus Thomas and Schwann, Abstr. Proc. Zool. Soc. 
London, no. 2, p. 5, 9 Feb. 1904; Proc. Zool. Soc. London, p. 179, 9 June 

1904. Klipfontein, Namaqualand, Cape of Good Hope. 

Rhabdomys pumilio dilectus (De Winton) 

Arvicanthis pumilio dilectus De Winton, Proc. Zool. Soc. London, for 1896, 
p. 803, Apl. 1897. Mazoe, Mashonaland, Southern Rhodesia. 

Rhabdomys pumilio diminutus (Thomas) 

Isomys pumilio diminutus Thomas, Proc. Zool. Soc. London, for 1892, p. 551, 
Apl. 1893. Mianzini, east of Lake Naivasha, Kenya Colony. 

Rhabdomy's pumilio griqu^ (Wroughton) 

Arvicanthis pumilio griquas Wroughton, Ann. Mag. Nat. Hist. (7) 16: 632, Dec. 

1905. Kuruman, Bechuanaland. 

Arvicanthis pumilio deserti Dollman, Ann. Mag. Nat. Hist. (8) 6: 399, Oct. 
1910. Lehutitung, Kalahari, Bechuanaland, 3000 ft. 

Rhabdomys pumilio intermedius (Wroughton) 

Arvicanthis pumilio intermedius Wroughton, Ann. Mag. Nat. Hist. (7) 16: 635, 
Dec. 1905. Deelfontein, Cape of Good Hope. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 415 

Rhabdomys pumilio meridionalis (Wroughton) 

Arvicanthis 'pumilio meridionalis Wroughton, Ann. Mag. Nat. Hist. (7) 16: 632, 
Dec. 1905. Tokai Retreat, Cape Town, Cape of Good Hope. 

Rhabdomys pumilio moshesh (Wroughton) 

Arvicanthis -pumilio moshesh Wroughton, Ann. Mag. Nat. Hist. (7) 16: 638, Dec. 
1905. Maseru, Basutoland, Cape of Good Hope, 5000 ft. 

Rhabdomys pumilio nyas^ (Wroughton) 

Arvicanthis -pumilio nyasse Wroughton, Ann. Mag. Nat. Hist. (7) 16: 639, Dec» 
1905. Mlanje Plateau, Nyasaland, 6000 ft. 



Stenocephalemys Frick. Narrow-headed Rat 

Stenocephalemys Frick, Ann. Carnegie Mus., 9: 7, 6 June 1914. Genotype 
Stenocephalemys albocaudata Frick. 

Stenocephalemys albocaudata Frick 

Stenocephalemys albocaudata Frick, Ann. Carnegie Mus., 9: 8, pi. 1, f. 1-5; pi. 2, 
f. 1-5, 6 June 1914. Inyala Camp, Chilalo Mts., southern Abyssinia. 



Stochomys Thomas 

Stochomys Thomas, Ann. Mag. Nat. Hist. (9) 17: 176, Jan. 1926. Genotype 
Dasymys longicaudatus Tullberg. 

Stochomys longicaudatus longicaudatus (Tullberg). Bristly Rat 

Mus hypoleucus Pucheran, Revue et Mag. de Zool. (2) 7: 206, Apl. 1855. 

Gaboon. 
Not Mus hypoleucus Sundevall; according to Trouessart (Cat. Mamm. Viv. 
Foss., ed. 1, pt. 2, p. 485, 1897) this is the same as Dasymys longicaudatus Tull- 
berg = a Stochomys. 
D[asymys] longicaudatus Tullberg, Nova Acta Reg. Soc. Sci. Upsala, (3) 16: art. 

12, p. 36, pi. 2, f. 39-46; pi. 3, f. 5, 48, 49; pi. 4, f. 10, 1893. Cameroons. 
Mus sebastianus De Win ton, Ann. Mag. Nat. Hist. (6) 19: 463, Apl. 1897. 

Efulen, Cameroons. 

Stochomys longicaudatus ituricus (Thomas) 

Epimys longicaudatus ituricus Thomas, Ann. Mag. Nat. Hist. (8) 16: 149,. 
Aug. 1915. Upper Ituri River, Congo Beige. 



416 bulletin: museum of comparative zoology 



Thallomys Thomas. White-bellied Tree Rats 

Thallomys Thomas, Ann. Mag. Nat. Hist. (9) 5: 141, Jan. 1920. Genotype Mus 
nigricauda Thomas. 

See Thomas, O., and Hinton, M. A. C, Proc. Zool. Soc. London, p. 493, 
3 Sep. 1923, for key to then-known forms. 

Thallomys damarensis damarensis (De Winton). Damara Tree-rat 
Mus damarensis De Winton, Ann. Mag. Nat. Hist. (6) 19: 349, Mar. 1897. 
Damaraland (perhaps the Grootfontein-Okavango region, specimens from 
which most nearly match the type, according to Shortridge, Mamm. 
South West Afr., 1: 270, footnote, 1934). 

Thallomys damarensis herero Thomas 

Thallomys herero Thomas, Proc. Zool. Soc. London, p. 303, 29 Apl. 1926. 
Ondongwa, Ovamboland, Southwest Africa. 

Thallomys damarensis rhodesi^ (Osgood) 

Mus damarensis rhodesiie Osgood, Ann. Mag. Nat. Hist. (8) 5: 277, Mar. 
1910. Petauke, East Loangwa district. Northern Rhodesia. 

Thallomys damarensis scotti Thomas and Hinton 

T[hallomys] scotti Thomas and Hinton, Proc. Zool. Soc. London, p. 493, 494, 

3 Sep. 1923. Yata Plains, east of Thika River where it joins the Tana, 

Kenya Colony, 4000 ft. 
Range: Eastern Kenya Colony to Dodoma, Tanganyika Territory. 

Thallomys damarensis stevensoni Roberts 

Thallomys stevensoni Roberts, Ann. Transvaal Mus., 15: 269, 1 Aug. 1933. 
Bembesi, 30 miles north of Bulawayo, Southern Rhodesia. 

Thallomys moggi moggi (Roberts) 

Mus moggi Roberts, Ann. Transvaal Mus., 4: 85, 20 Oct. 1913. Zoutpan, 
Pretoria district, Transvaal. 
For Ernest Mogg. 

Thallomys moggi acacia (Roberts) 

Mus moggi acacise Roberts, Ann. Transvaal Mus., 5: 120, 28 Aug. 1915. 
Woodbush, and Moorddrift, Transvaal. 

Thallomys moggi lebomboensis Roberts 

Thallomys moggi lebomboensis Roberts, Ann. Transvaal Mus., 14: 234, 28 July 
1931. Mkuzi River, Ubombo, northern Zululand, Natal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 417 

Thallomys nigricauda nigricauda (Thomas). Black-tailed Tree Rat 
Mus nigricauda Thomas, Proc. Zool. Soc. London, p. 266, pi. 14, f.l, June 
1882. Hountop ( = Hudup or Hutop) River, Great Namaqualand, South- 
west Africa. 
T[hallomys] longicauda Thomas, Proc. Zool. Soc. London, p. 493, 3 Sep. 1923. 
Lapsus calami for T. nigricauda. 

Thallomys nigricauda bradfieldi Roberts 

Thallomys leuconoe bradfieldi Roberts, Ann. Transvaal Mus., 15: 268, 1 Aug. 
1933. Quickborn Farm, Okahandja, Southwest Africa. 

Thallomys nigricauda kalaharicus (DoUman) 

Epimys nigricauda kalaharicus Dollman, Ann. Mag. Nat. Hist. (8) 8: 544, 
Oct. 1911. Molopo River, Kalahari, Bechuanaland. Regarded by Short- 
ridge (Mamm. South West Africa, 1: 266, footnote, 1934) as a synonym 
of T. n. nigricauda. 

Thallomys nigricauda leuconoe Thomas 

Thallomys leuconoe Thomas, Proc. Zool. Soc. London, p. 303, 29 Apl. 1926. 

Osohama, Etosha Pan, Ovamboland, Southwest Africa. 
Thallomys nigricauda leuconoe Shortridge, Mamm. South West Africa, 1: 

266, 1934. 

Thallomys nigricauda loringi (Heller) 

Thamnomys loringi Heller, Smithsonian Misc. Coll., 52: pt. 4, p. 471, pi. 56, 
fig. a, 13 Nov. 1909. Lake Naivasha, Kenya Colony. 

Thallomys nigricauda molopensis Roberts 

Thallomys leuconoe molopensis Roberts, Ann. Transvaal Mus., 15: 269, 1 Aug. 

1933. Midway between Setlagoli and Molopo River at Pitsani Location, 

Bechuanaland. 

Thallomys nigricauda nitela Thomas and Hinton 

T[hallomys] nitela Thomas and Hinton, Proc. Zool. Soc. London, p. 493, 
3 Sep. 1923. Bombone, Mossamedes, Angola, 3200 ft. 

Thallomys nigricauda shortridgei Thomas and Hinton 

Thallomys shortridgei Thomas and Hinton, Proc. Zool. Soc. London, p. 492, 

3 Sep. 1923. Louisvale, near "Upington, south bank of Orange River, 

Southwest Africa. 



Thamnomys Thomas. Thicket Rats 

Thamnomys Thomas, Ann. Mag. Nat. Hist. (7) 19: 121, Jan. 1907. Geno- 
type Thamnomys venustus Thomas. 



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Subgenus Thamnomys Thomas 

See Thomas, O., Ann. Mag. Nat. Hist. (8) 16: 150, Aug. 1915, for char- 
acters and list of species. 

Thamnomys kempi kempi Dollman 

Thamnomys kempi Dollman, Ann. Mag. Nat. Hist. (8) 8: 658, Nov. 1911. 
Buhamba, near Lake Kivu, Congo Beige, 6000 ft. 

Thamnomys kempi major Hatt 

Thamnomys kempi major Hatt, Amer. Mus. Novitates, no. 708, p. 10, 4 Apl. 

1934. Lukumi, north slope of Mt. Karisimbi, Kivu volcanoes, Congo 

Beige, 12,000 ft. 

Thamnomys kuru Thomas and Wroughton 

Thamnomys kuru Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 
381, May 1907. Angu, Uele River. 

Thamnomys rutilans rutilans (Peters) 

Mus rutilans Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 478, pi. 2, 
f. 2, 1876. Limbareni, West Africa. 

Thamnomys rutilans centralis Dollman 

Thamnomys rutilans centralis Dollman, Revue Zool. Africaine, 4: 83, July 1914. 
Fundi, Congo Beige. 

Thamnomys venustus venustus Thomas 

Thamnomys venustus Thomas, Ann. Mag. Nat. Hist. (7) 19: 122, Jan. 1907. 
Ruwenzori East, Uganda, 7000 ft. 

Thamnomys venustus schoutedeni Hatt 

Thamnomys venustus schoutedeni Hatt, Amer. Mus. Novitates, no. 708, p. 9, 
4 Apl. 1934. Medje, Ituri district, Congo Beige. 



Subgenus Grammomys Thomas (x-cusp reduced) 

Grammomys Thomas, Ann. Mag. Nat. Hist. (8) 16: 150, Aug. 1915. Genotype 
Miis dolichurus Smuts. A synonym of Thamnomys according to Hollister 
(Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 57, 1919). 

Thamnomys baliolus Osgood 

Thamnomys baliolus Osgood, Ann. Mag. Nat. Hist. (8) 5: 278, Mar. 1910. 
Woodbush Hills, northeast of Pietersburg, Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 419 

Thamnomys BUNTING! Thomas 

Thamnomys buntingi Thomas, Ann. Mag. Nat. Hist. (8) 7: 381, Apl. 1911. 
Gonyon, Bassa, Liberia. 

Thamnomys cometes Thomas and Wroughton 

Thamnomys cometes Thomas and Wroughton, Proc. Zool. Soc. London, p. 549, 
30 Oct. 1908. Inhambane, Mozambique. 

Thamnomys dolichurus dolichurus (Smuts) 

Mlus] dolichurus Smuts, Enumerat. Mamm. Capens., p. 38, pi. 2, 1832. Near 
Cape Town, South Africa. 

Thamnomys dolichurus tongensis (Roberts) 

Grammomys to7igensis Roberts, Ann. Transvaal Mus., 14: 234, 28 July 1931. 
Manaba, northern Zululand, Natal. 

Thamnomys dryas Thomas 

Thamnomys dryas Thomas, Ann. Mag. Nat. Hist. (7) 19: 123, Jan. 1907. Ru- 
wenzori East, Kenya Colony, 6-7000 ft. 

Thamnomys gigas DoUman 

Thamnomys gigas DoUman, Ann. Mag. Nat. Hist. (8) 7: 527, May 1911. Solai, 
Mt. Kenya, Kenya Colony, 9000 ft. 

Thamnomys ibeanus ibeanus Osgood 

Thamnomys ibeanus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: no. 2, 
p. 8, 16 Feb. 1910. Molo, Kenya Colony. 

Thamnomys ibeanus lutosus Dollman 

Thamnomys surdaster lutosus Dollman, Ann. Mag. Nat. Hist. (8) 8: 657, Nov. 
1911. Mt. Nyiro, Kenya Colony. 

Thamnomys macmillani macmillani Wroughton 

Thamnomys macmillani Wroughton, Ann. Mag. Nat. Hist. (7) 20: 504, Dec. 
1907. Wouida, north of Lake Rudolf, Ethiopia (Abyssinia). 

Thamnomys macmillani aridulus (Thomas and Hinton) 

GramTnomys aridulus Thomas and Hinton, Proc. Zool. Soc. London, p. 268, 

6 July 1923. Kulme, Wadi Aribo, Darfur, Anglo-Egyptian Sudan. 
Grammomys macmillani aridulus Braestrup, Vidensk. Medd. fra Dansk Naturh. 

Foren., Kjobenhavn, 99: 115, 11 Sep. 1935. 

Thamnomys macmillani gazell,e Thomas 

Thamnomys macmillani gazellx Thomas, Ann. Mag. Nat. Hist. (8) 5: 282, 
Mar. 1910. Chak Chak, Bahr-el-Ghazal, Anglo-Egyptian Sudan. 



420 bulletin: museum of comparative zoology 

Thamnomys macmillani oblitus Osgood 

Thamnomys oblitus Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: no. 3, 
p. 16, 7 Apl. 1910. Vol, Kenya Colony. 

Thamnomys macmillani ochraceus G. M. Allen 

Thamnomys ochraceus G. M. Allen, Bull. Mus. Comp. Zool., 54: 442, Apl. 1912. 
Meru River, Northern Guaso Nyiro, Kenya Colony. 

Thamnomys macmillani tuareg (BrsDstrup) 

Gramtnomys macmillani tuareg Bra3strup, Vidensk. Medd. fra Dansk Naturh. 
Foren., Kjobenhavn, 99: 113, 11 Sep. 1935. Timbuctu, French Sudan. 

Thamnomys ruddi Thomas and Wroughton 

Thamnomys ruddi Thomas and Wroughton, Proc. Zool. Soc. London, p. 549, 
30 Oct. 1908. Tette, Mozambique. 

Thamnomys surdaster surdaster Thomas and Wroughton 

Thamnomys surdaster Thomas and Wroughton, Proc. Zool. Soc. London, p. 

550, 30 Oct. 1908. Zomba, Nyasaland. 
Thamnomys usambarse Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 99, 

Apl. 1915. Amani, Usambara, Tanganyika Territory. 

Thamnomys surdaster angolensis (Hill and Carter) 

Gratnmoinys surdaster angolensis Hill and Carter, Amer. Mus. Novitates, no. 
913, p. 4, 8 Mar. 1937. Chitau, Angola, 4930 ft. 

Thamnomys surdaster callithrix Hatt 

Thafunomys {Grammomys) surdaster callithrix Hatt, Amer. Mus. Novitates, 
no. 708, p. 11, 4 Apl. 1934. Garamba, upper Uele district, Congo Beige. 

Thamnomys surdaster discolor Thomas 

Thamnomys discolor Thomas, Ann. Mag. Nat. Hist. (8) 5: 283, Mar. 1910. 
Kakamega Forest, Kisumu, Kenya Colony. 

Thamnomys surdaster elgonis Thomas 

Thamnomys surdaster elgonis Thomas, Ann. Mag. Nat. Hist. (8) 5: 282, Mar. 

1910. Malikisi, south slope of Mt. Elgon, Kenya Colony, 5000 ft. 

Thamnomys surdaster insignis Dollman 

Thamnomys surdaster insignis Dollman, Ann. Mag. Nat. Hist. (8) 7: 528, May 

1911. South face of Mt. Elgon, Kenya Colony, 9000 ft. 

Thamnomys surdaster littoralis Heller 

Thamnomys dolichurus littoralis Heller, Smithsonian Misc. Publ., 59: no. 16, 
p. 10, 5 July 1912. Mazeras, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 421 

Thamnomys surdaster polionops Osgood 

Thamnomys surdaster ■polionops Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 
10: no. 2, p. 8, 16 Feb. 1910. Lukenya Mt., Ulukenia Hills, Kenya Colony. 

Thamnomys vxtmbensis (Roberts) 

Grammomys vumbensis Roberts, Ann. Transvaal Mus., 19: 245, 31 Oct. 1938. 
Vumba, Southern Rhodesia. 



Uranomys Dollman 

Uranomys Dollman, Ann. Mag. Nat. Hist. (8) 4: 551, Dec. 1909. Genotype 
Uranomys ruddi Dollman. 

Uranomys acomyoides Ingoldby 

Uranomys acomyoides Ingoldby, Ann. Mag. Nat. Hist. (10) 3: 523, May 1929. 
Kintampo, northern Ashanti, Gold Coast. 

Uranomys foxi Thomas 

Uranomys foxi Thomas, Ann. Mag. Nat. Hist. (8) 9: 273, Feb. 1912. Panyam 
Plateau, northern Nigeria. 

Uranomys oweni Thomas 

Uranomys oweni Thomas, Ann. Mag. Nat. Hist. (8) 6: 431, Oct. 1910. Mara- 
kissa, French Guinea. 

Uranomys ruddi Dollman 

Uranomys ruddi Dollman, Ann. Mag. Nat. Hist. (8) 4: 552, Dec. 1909. Kirui, 
Mt. Elgon, Kenya Colony, 6000 ft. 
Probably all the named forms will prove to be subspecies of this. 

Uranomys tenebrosus Hinton 

Uranomys tenebrosus Hinton, Ann. Mag. Nat. Hist. (9) 7: 370, Apl. 1921. 
N'sana, northwestern Nyasaland. 

Uranomys ugand^ Heller. 

Uranomys ugandse Heller, Smithsonian Misc. Coll., 56: no. 17, p. 12, 28 Feb. 
1911. Kikonda, Uganda. 

Uranomys woodi Hinton 

Uranomys woodi Hinton, Ann. Mag. Nat. Hist. (9) 7: 369, Apl. 1921. Cholo, 
Nyasaland. 



422 bulletin: museum of comparative zoology 



Zelotomys Osgood. Broad-headed Mice 

Zelotomys Osgood, Publ. Field Mus. Nat. Hist., zopl. ser., 10: no. 2, p. 7, 16 
Feb. 1910. Genotype Mus hildegardex Thomas. 

Zelotomys HiLDEOARDEiE hildegarde^ (Thomas) 

Mus hildegardex Thomas, Ann. Mag. Nat. Hist. (7) 9: 219, Mar. 1902. Mach- 
akos, Kenya Colony. 

Zelotomys hildegarde^ vinacetjs Heller 

Zelotomys hildegardse (sic) vinaceus Heller, Smithsonian Misc. Coll., 59: no. 16, 
p. 10, 5 July 1912. Ndi, Mt. Mbololo, Taita Hills, Kenya Colony, 3000 ft. 

Zelotomys instans Thomas 

Zelotomys instans Thomas, Ann. Mag. Nat. Hist. (8) 16: 480, Dec. 1916 (mam- 
mary formula given). Poko, Uele, Congo Beige. 

Zelotomys shortridgei shortridgei Hinton 

Zelotomys shortridgei Hinton, Ann. Mag. Nat. Hist. (9) 6: 242, Aug. 1920. 
N'dola, Northern Rhodesia. 

Zelotomys shortridgei kuvelaiensis St. Leger 

Zelotomys shortridgei kuvelaiensis St. Leger, Ann. Mag. Nat. Hist. (10) 17: 470, 

Apl. 1936. Kuvelai River, about 50 km. north of Mupa, Mossamedes 

(17° S., 16°30'E.). 



Family JACULIDiE. Jerboas 

Subfamily JACULIN.E 

Note: Alactaga arimdinis F. Cuvier (Trans. Zool. Soc. London, 2: 134, 4 May 
1838) North Africa, is probably an error, for the genus is not known from 
Africa. Based on Shaw's Voyage en Barbarie, French transl., 1: 321, in which 
there is an evident error in describing the hind toes as four long and two short. 

Jaculus Erxleben. Jerboa 

Jaculus Erxleben, Syst. Regni Animalia, p. 404, 1777. Genotype Jaculus orien- 
talis Erxleben. 

Jaculus hirtipes (Lichtenstein) 

Dipus hirtipes Lichtenstein, Verzeichn. d. Doubletten Zool. Mus. Berlin, p. 5, 
1823. "E deserto prope Sakharum." 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 423 

Jaculus JACULUS JACULUS (Linnseus). Jerboa. 

Mus jaculus Linnseus, Syst. Nat., ed. 10, 1: 62, 1758. Giza Pyramids, Egypt; 

"in Arabia, Calmukia." 
Dipus segyplius Lichtenstein, Darstellungen Neue Saugeth., pi. 22 and text, 

1827. Egypt. 

Jaculus jaculus airensis Thomas and Hinton 

Jaculus jaculus airensis Thomas and Hinton, Novitates Zool., 28: 10, 14 May 
1921. Aderbissinat, northern Nigeria. 

Jaculus jaculus butleri Thomas 

Jaculus jaculus butleri Thomas, Ann. Mag. Nat. Hist. (9) 9: 296, Apl. 1922. 
Khartoum, Anglo-Egyptian Sudan. 
Range : Khartoum to Sennaar. 

Jaculus jaculus centralis Thomas and Hinton 

J[aculus] j{aculus] centralis Thomas and Hinton, Novitates Zool., 28: 11, 14 
May 1921. Oued-el-Abiad, north of In-salah, central Sahara of Algeria. 

Jaculus jaculus desertii (Loche) 

Dipus desertii Loche, Cat. Mamm. de I'Algerie, p. 21, 1858; Exploration Sci. de 

I'Algerie, Zool., Mammiferes, p. 100, 1867. Oasis of Ouargla, southern 

Algeria. 
Dipus darricarrerei Lataste, Anal. Mus. Civ. Stor. Nat. Genova, 18: 661, 11 

June 1883. Bou Saada, Algerian Sahara. A synonym of J.j. desertii (see 

Thomas and Hinton, Novitates Zool., 28: 10, 14 May 1921). 
Range: Algeria between the high plateaux and the Sahara, and in Tunisia. 

Jaculus jaculus favonicus Thomas 

Jaculus jaculus favonicus Thomas, Ann. Mag. Nat. Hist. (8) 11 : 483, May 1913. 
Trarza country, southwestern Mauritania. 

Jaculus jaculus gordoni Thomas 

Jaculus gordoni Thomas, Proc. Zool. Soc. London, 1: 299, 6 Aug. 1903. Kaga 
Hills, western Kordofan, Anglo-Egyptian Sudan. 
Range: Kordofan, Egyptian Sudan. 

Jaculus jaculus sefrius Thomas and Hinton 

Jaculus jaculus sefrius Thomas and Hinton, Novitates Zool., 28: 10, 14 May 

1921. Ain-Sefra, southwestern Algeria. 
Range: Desert region of Morocco, to the southwest of the Great Atlas, and 

the adjacent parts of the sandy Sahara. 



424 bulletin: museum of comparative zoology 

Jaculus jaculus vulturnus Thomas 

Jaculus jaculus vulturnus Thomas, Ann. Mag. Nat. Hist. (8) 11 : 485, May 1913. 
Berbera, British Somahland. 

Jaculus microtis (Reichenow) 

Dipus microtis Reichenow, Zool. Anzeiger, 10: 369, 11 July 1887. Samar, 

northeast Africa. Based on a young specimen, see Thomas, O., Proc. Zool. 

Soc. London, 1: 299, 6 Aug. 1903. 

Jaculus orientalis Erxleben 

Jaculus orientalis Erxleben, Syst. Regni Animalis, p. 404, 1777. Egypt, in the 
mountains separating Egypt from Arabia. 

Dipus abyssinicus F. A. A. Meyer, Syst.-summarisch. Uebersicht d. Entdec- 
kungen in Neu-Holland u. Africa, p. 82, 1793. Sandy deserts of Africa 
(Abyssinia implied) ; based on Bruce's Travels to discover Source of Nile, 
5: pi. 27, 1791, German ed. 
Range: Coastal districts of western Lower Egypt, Tripoli, Tunisia and Al- 
geria (S.S. Flower). 



Scarturus Gloger. Four-toed Jerboa 

Scarturus Gloger, Hand- u. Hilfsbuch d. Naturgesch., 1: pp. xx.xi, 106, 1841. 

Genotype Dipus tetradactylus Lichtenstein (see Thomas, Ann. Mag. Nat. 

Hist. (6) 15: 191, Feb. 1895). 
Scirtomys Brandt, Bull. Acad. Imp. des Sci., St. Petersbourg, CI. phys.-math., 

2: 220, 20 Jan. 1844. As a subgenus of Alactaga, type Alactaga (Dipus) 

tetradactylus (Lichtenstein). 

Scarturus tetradactylus (Lichtenstein) 

Dipus tetradactylus Lichtenstein, Verzeichn. d. Doubletten in Zool. Mus. Ber- 

Un, p. 2, 1823. Libyan Desert between Siwa and Alexandria, Egypt. 
Dipus brucii Lesson, Manuel de Mammalogie, p. 253, 1827. Based on the 

gerboa described by Bruce from Barca. 
Range: Known from the type locality, and districts of Mariut and Mersa 

Matruh, Egypt. 

SciRTOPODA Brandt 

Scirtopoda Brandt, Bull. Acad. Imp. des Sci., St. Petersbourg, CI. phys.- 
math., 2: 212, 20 Jan. 1844. As a subgenus of Dipus, type Dipus mauri- 
tanicus Duvernoy. 

Haltomys Brandt, Bull. Acad. Imp. des Sci., St. Petersbourg, 2: 215, 20 Jan. 
1844. As a section of Scirtopoda, type not specified, but by elimination is 
Dipus jnauritanicus Duvernoy. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 425 

Haliicus Brandt, Bull. Acad. Imp. des Sci., St. Petersbourg, 2: 213, 20 Jan. 
1844. As a section of Scirtopoda, type Dipus halticus lUiger; preoccupied 
by Halticus Hahn, 1831, in Hemiptera. 

Scirtopoda gerboa gerboa (Olivier) 

Dipus gerboa Olivier, Bull, des Sci., Soc. Philomathique, Paris, (1) 2: no. 39, 

p. 121, 1801. Egypt. 
Dipus locusta lUiger, Abhandl. K. Preuss. Akad. Wiss., Berlin, 1804-11, 

p. 77, 1815, (nomen nudum). 
Dipus bipes Lichtenstein, Verzeichn. d. Doubletten Zool. Mus., Berlin, p. 5, 

1823. "InAegypto." 
Range: Egypt, west toward Morocco. 

Scirtopoda gerboa maxjritanica (Duvernoy) 

Dipus mauritanicus Duvernoy, Mem. Soc. du Mus. d'Hist. Nat. Strasbourg, 

3: pt. 2, art. QQ, p. 30, pi. 3, 4, 1840. Mauritania. 
Scirtopoda mauritanicus Pocock, Proc. Zool. Soc. London, p. 680, 28 Sep. 

1922. 
Scirtopoda gerboa mauritanica Cabrera, Trab. Mus. Nac. de Ciencias Nat., 

Madrid, zool. ser., no. 57, p. 277, text-f. 33 (skull), 30 Dec. 1932. 
Range: Algeria and in Morocco the Rif, to the high plateaux. 



Family BATHYERGID^. Mole-rats 

On the tooth formulae and homologies of the genera see Thomas, O., Ann. 
Mag. Nat. Hist. (8) 4: 110-112, Aug. 1909. 



Bathyergus Illiger 

Bathyergus Illiger, Prodromus Syst. Mamm. et Avium, p. 86, 1811. Genotype 

Mus maritimus Gmelin = Mus suillus Schreber. 
Orycterus F. Cuvier, Dictionnaire des Sci. Nat., 59: 481, 1829. Genotype Mus 

maritimus Gmelin. 

Bathyergus suillus suillus (Schreber). Sand-mole 

Mus suillus Schreber, Saugthiere, 4: 715, pi. 204B, 1782. Cape of Good Hope. 

(On dates of publication, see Sherborn, C. D., Proc. Zool. Soc. London, 

for 1891, 587-592, Apl. 1892.) 
Mus maritimus Gmelin, Linnseus's Syst. Nat., ed. 13, 1: pt. 1, p. 140, 1788. 

Cape of Good Hope. 
Arctomis (sic) africana Lamarck, in Voyages de C. P. Thunberg au Japon, 

etc., 4: 348, 1796. Africa, Cape of Good Hope implied. 



426 bulletin: museum of comparative zoology 

Bathyergus suillus intermedius Roberts 

Bathyergus suillus intermedius Roberts, Ann. Transvaal Mus., 11: 261, 14 Sep. 
1926. Klaver, Cape of Good Hope. 

Bathyergus suillus janetta Thomas and Schwann 

Bathyergus janetta Thomas and Schwann, Abstr. Proc. Zool. Soc. London, no. 2, 

p. 6, 9 Feb. 1904; Proc. Zool. Soc. London, 1 : 180, pi. 6, f. 2, 9 June 1904. 

Port Nolloth, Namaqualand, Southwest Africa. 
Bathyergus suillus janetta Shortridge, Mamm. South West Africa, 1 : 318, 1934. 



Cryptomys Gray. Mole-rats 

■Cryptomys Gray, Proc. Zool. Soc. London, p. 124, text-f. 3, 6, July 1864. As a 
. subgenus of Georychus, type Georychus holosericeus Wagner. 

Typhloryctes Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. Kl., 55: sect. 1, 
p. 502, 1867. Genotype not specified; included Georychus ochraceo-cinereus 
Heuglin and Bathyergus caecutiens Brants. 

Ca-tomys Gray, Proc. Zool. Soc. London, p. 125, text-f. 4, 5, July 1864. A sub- 
genus of Georychus, type not specified, but included Bathyergus csecutiens 
Brants and B. damarensis Ogilby. 

Cryptomys aberrans (Roberts) 

Georychus aberrans Roberts, Ann. Transvaal Mus., 4: 97, 20 Oct. 1913. Port 
St. Johns, Cape of Good Hope. 

Cryptomys albus (Roberts) 

Georychus albus Roberts, Ann. Transvaal Mus., 4: 100, 20 Oct. 1913. ?Wyn- 
berg, Cape of Good Hope. 

Cryptomys amatus (Wroughton) 

Georychus amatus Wroughton, Mem. and Proc. Manchester Lit. and Phil. Soc, 

51 : no. 5, p. 28, 13 Mar. 1907. Road to Chiwali's, Alala plateau, Rhodesia, 

4000 ft. 

Cryptomys anomalus (Roberts) 

Georychus anomalus Roberts, Ann. Transvaal Mus., 4: 96, text-f. 10 (skull), 
20 Oct. 1913. Skinners Court Valley, Pretoria, Transvaal. 

Cryptomys ansorgei (Thomas and Wroughton) 

Georychus ansorgei Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 16: 175, 
Aug. 1905. Kukema River, Bihe, Angola, 5900 ft. 

Cryptomys arenarius (Roberts) 

Georychus arenarius Roberts, Ann. Transvaal Mus., 4: 96, text-f. 9 (skull), 
20 Oct. 1913. Rietondale (East), Pretoria, Transvaal. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 427 

Cryptomys beir^ (Thomas) 

Georychus heirse Thomas, Proc. Zool. Soc. London, for 1907, p. 780, 4 June 1908. 
Beira, Mozambique. 

Cryptomys bigalkei Roberts 

Cryptomys bigalkei Roberts, Ann. Transvaal Mus., 10: 73, 31 Jan. 1924. Glen, 
Orange Free State. 

Cryptomys blainei Hinton 

Cryploviys blainei Hinton, Ann. Mag. Nat. Hist. (9) 7: 372, Apl. 1921. Chi- 
songwe, Luando River, central Angola. 

Cryptomys bocagei (De Winton) 

Georychus bocagei De Winton, Ann. Mag. Nat. Hist. (6) 20: 323, Sep. 1897. 
Hanha, Angola. 

Cryptomys cradockensis Roberts 

Cryptomys cradockensis Roberts, Ann. Transvaal Mus., 10: 73, 31 Jan. 1924. 
Cradock, Cape of Good Hope. 

Cryptomys damarensis damarensis (Ogilby) 

Bathyergtis da?narensis Ogilby, Proc. Zool. Soc. London, p. 5, July 1838. 
"Country of the Damaras," i.e. Damaraland, Cape of Good Hope. 

Cryptomys damarensis lugardi (De Winton) 

Georychus lugardi De Winton, Ann. Mag. Nat. Hist. (7) 1: 253, Mar. 1898. 

Kalahari Desert, between Palapye and Lake Ngami, northern Bechu- 

analand. 
Cryptomys damarensis lugardi Shortridge, Mamm. South West Africa, 1 : 320, 

1934. 

Cryptomys damarensis micklemi (Chubb) 

Georychus micklemi Chubb, Ann. Mag. Nat. Hist. (8) 3: 35, Jan. 1909. Kataba 

River, upper Zambesi, northwestern Rhodesia. 
Cryptomys damarensis micklemi Shortridge, Mamm. South West Africa, 1 : 320, 

1934. Possibly a synonym of C. d. lugardi (De Winton). 

Cryptomys darlingi (Thomas) 

Georychus darlingi Thomas, Ann. Mag. Nat. Hist. (6) 16: 239, Sep. 1895. 
Salisbury, Cape of Good Hope. 

Cryptomys foxi (Thomas) 

Georychus foxi Thomas, Ann. Mag. Nat. Hist. (8) 7: 462, May 1911. Panyam, 
northern Nigeria, 4000 ft. 



428 bulletin: museum of comparative zoology 

Cryptomys holosericeus (Wagner) 

G[eorychus] holosericeus Wagner, Schreber's Saugth., SuppL, 3: 373, 1843. 
Graaf Reinet, Cape of Good Hope (selected as type locality and species 
regarded as valid by Thomas, O., and Schwann, H., Proc. Zool. Soc. 
London, 1: 166, 7 June 1906). 

Cryptomys hottentotus hottentotus (Lesson) 

Bathyergus hottentotus Lesson, Voyage autour du Monde sur la Coquille, Zool., 

1 : 166, pi. 2, f . 2, 1826. Pearl Village twenty miles from Cape Town, Cape 

of Good Hope. 
Bathyergus csecutiens Brants, Het Geslacht der Muizen, p. 37, 1827. Cape of 

Good Hope. (Brants credits the name to Lichtenstein.) 
Bathyergus ludwigii A. Smith, Zool. Journ., 4: 439, May 1829. "Various parts 

of South Africa." 
Georychus exenticus Trouessart, Cat. Mamm. Viv. Foss,. app., ed. 1, p. 1338, 

1899. Mistake for caecutiens Brants. 

Cryptomys hottentotus occlustjs G. M. Allen and Loveridge 
Cryptomys hottentotus occlusus G. M. Allen and Loveridge, Bull. Mus. Comp. 
Zool., 75: 125, Feb. 1933. Kigogo, Uzungwe Mts., southwestern Tan- 
ganyika Territory. 

Cryptomys hottentotus talpoides (Thomas and Schwann) 
Georychus hottentottus (sic) talpoides Thomas and Schwann, Proc. Zool. Soc. 
London, p. 166, 7 June 1906. Knysna, Cape of Good Hope. 

Cryptomys jamesoni (Roberts) 

Georychus jamesoni Roberts, Ann. Transvaal Mus., 4: 95, 20 Oct. 1913. Hough- 
ton Estate, Johannesburg, Transvaal. 

Cryptomys jorisseni (Jameson) 

Georychus jorisseni Jameson, Ann. Mag. Nat. Hist. (8) 4: 466, Nov. 1909. 
Waynek, Waterberg district, Transvaal. 

Cryptomys junodi Roberts 

Cryptomys junodi Roberts, Ann. Transvaal Mus., 11: 260, 14 Sep. 1926. 
Masiene, Mozambique. 

Cryptomys komatiensis (Roberts) 

Georychus komatiensis Roberts, Ann. Transvaal Mus., 5: 272, 16 May 1917. 
Arnhemburg, Carolina, Transvaal. 

Cryptomys kubanqensis (Monard) 

Georhychus (sic) kubangensis Monard, Bull. Soc. Neuchateloise des Sci. Nat., 

Neuchatel, 57: 58, text-f. 1-4, 1932. Angola, specimens from Rio Mbal6, 

Vila da Ponte, Chimporo, and Tumbole. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 429 

Cryptomys kummi (Thomas) 

Georychus kummi Thomas, Ann. Mag. Nat. Hist. (8) 7: 592, June 1911. French 
Shari Protectorate, Ironstone plateau, about 8° N., 22° E. 

Cryptomys lechei (Thomas) 

Georychus lechei Thomas, Ann. Mag. Nat. Hist. (6) 16: 241, Sep. 1895. BeUima, 
Monbuttu, Congo Beige. 

Cryptomys mahali (Roberts) 

Georychus mahali Roberts, Ann. Transvaal Mus., 4: 108, 20 Oct. 1913. Rosslyn, 
Rustenburg district, Transvaal. 

Cryptomys mechowii (Peters) 

Georychus mechovoii Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 133, 1881. 
Malange, Benguela, Angola. 

Cryptomys melanoticus Roberts 

Cryptomys melanoticus Roberts, Ann. Transvaal Mus., 11: 260, 14 Sep. 1926. 
Balloon, Makoetsi River, northeastern Transvaal. 

Cryptomys mellandi (Thomas) 

Georychus mellandi Thomas, Ann. Mag. Nat. Hist. (7) 17: 178, Feb. 1906. 
Mpika, northeastern Rhodesia, 5000 ft. 

Cryptomys molyneuxi (Chubb) 

Georychus molyneuxi Phubb, Ann. Mag. Nat. Hist. (8) 2: 451, Nov. 1908. 
Loano valley, northwestern Rhodesia. 

Cryptomys montanus Roberts 

Cryptomys montanus Roberts, Ann. Transvaal Mus., 11: 260, 14 Sep. 1926. 
Klapperkop, Pretoria, Transvaal. 

Cryptomys natalensis natalensis (Roberts) 

Georychus natalensis Roberts, Ann. Transvaal Mus., 4: 94, text-f. 8 (skull), 20 
Oct. 1913. Wakkerstroom, Transvaal. 

Cryptomys natalensis nemo new name 

Georychus natalensis pallidus Roberts, Ann. Transvaal Mus., 5: 278, 16 May 

1917. Manetsi River, near Malala, Zoutpansberg district, Transvaal. 

Not Georychus pallidus Gray, 1864, a Heliophobius. 

Cryptomys nimrodi (De Winton) 

Georychus nimrodi De Winton, Proc. Zool. Soc. London, for 1896, p. 808, pi. 40, 
f. 1, Apl. 1897. Essex Farm, near Bulawayo, Southern Rhodesia. 



430 bulletin: museum of comparative zoology 

Cryptomys ochraceo-cinereus (Heuglin) 

Georychus ochraceo-cinereus Heuglin, Nova Acta Acad. Caes. Leop.-Carol.^ 
Dresden, 31: art. 7, p. 3, 1864. Upper Bahr-el-Ghazal, 7-8° N., 24-26° E. 

Cryptomys orangije Roberts 

Cryptomys orangise Roberts, Ann. Transvaal Mus., 11 : 259, 14 Sep. 1926. Glen, 
Orange Free State. 

Cryptomys palki (Roberts) 

Georychus palki Roberts, Ann. Transvaal Mus., 6: 5, 28 June 1917. Venter- 
skroon, Vaal River, Potchefstroom district, Transvaal. 

Cryptomys Pretoria (Roberts) 

Georychus pretorix Roberts, Ann. Transvaal Mus., 4: 99, 20 Oct. 1913. Skinners 
Court Valley, Pretoria, Transvaal. 

Cryptomys rufulus (Roberts) 

Georychus rufulus Roberts, Ann. Transvaal Mus., 5: 272, 16 May 1917. Tzan- 
een, Transvaal. 

Cryptomys stellatus (Roberts) 

Georychus stellatus Roberts, Ann. Transvaal Mus., 5: 272, 16 May 1917. Koma^ 
tipoort, Gazaland, Transvaal. 

Cryptomys transvaalensis Roberts 

Cryptomys transvaalensis Roberts, Ann. Transvaal Mus., 10: 73, 31 Jan. 1924. 
Boekenhoutfontein farm, Pretoria district, Transvaal. 

Cryptomys van dami (Roberts) 

Georychus van dami Roberts, Ann. Transvaal Mus., 5: 273, 16 May 1917. 
Griffin Mine, Leydsdorp, Transvaal. 

Cryptomys vetensis Roberts 

Cryptomys vetensis Roberts, Ann. Transvaal Mus., 11: 259, 14 Sep. 1926. Taai- 
boschspruit. Vet River, Orange Free State. 

Cryptomys vryburgensis (Roberts) 

Georychus vryburgensis Roberts, Ann. Transvaal Mus., 5: 274, 16 May 1917. 
Vryburg, southern Bechuanaland. 

Cryptomys whytei (Thomas) 

Georychus whytei Thomas, Proc. Zool. Soc. London, p. 432, 1 Oct. 1897. Kar- 
onga, Lake Nyasa, Nyasaland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 431 

Cryptomys zechi (Matschie) 

Georhychus (sic) zechi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 146, 
1900. Middle Volta, near Kete-ELradji station, Togo, West Africa. 



Georychus lUiger. Blesmols 

Georychus Illiger, Prodromus Systematis Mamm. et Avium, p. 87, 1811. Geno- 
type Mus capeusis Pallas (see J. A. Allen, Bull. Amer. Mus. Nat. Hist., 7: 
183, June 1895). 

Georhychus Wagner, Schreber's Saugthiere, Suppl., 3: 369, 1843, emendation. 

Fossor Lichtenstein, in Forster's Descr. Animalium in Iter, ad Maris Austr. 
Terras Suscepto, (edited by Lichtenstein), p. 364, 1844. Genotype Fossor 
leucops Lichtenstein = Georychus capensis (Pallas). 

Georychus capensis capensis (Pallas) 

Mus capensis Pallas, Nov. Spec. Quadrup. e Glir. Ord., pp. 76, 172, pi. 7, 1778. 

Cape of Good Hope. 
[Mu^] hufonii F. Cuvier, Ann. des Sci. Nat., zool., (2) 1: 196, 1834. Cape of 

Good Hope. 
Fossor leucops Lichtenstein, in J. R. Forster's Descrip. Animalium in Iter, ad 

Maris Austr. Terras Suscepto, (ed. by Lichtenstein), p. 364, 1844. Cape 

of Good Hope. 

Georychus capensis canescens Thomas and Schwann 

Georychus capensis canescens Thomas and Schwann, Proc. Zool. Soc. London, 1 1 
165, 7 June 1906. Knysna, Cape of Good Hope. 

Georychus capensis yatesi Roberts 

Georychus yatesi Roberts, Ann. Transvaal Mus., 4: 92, 20 Oct. 1913. Belfast,. 
Transvaal. 

Heliophobius Peters 

Heliophobius Peters, Ber. iiber Verhandl. K. Preuss. Akad. Wiss., Berlin, p. 

259, Aug. 1846. Genotype Heliophobius argenteocinereus Peters. 
Myoscalops Thomas, Proc. Zool. Sop. London, p. 448, Oct. 1890. Proposed to 

replace Heliophobius on the ground that it is preoccupied by Heliophobus 

Boisduval, in Lepidoptera, 1829. 

Heliophobius albifrons (Gray) 

Georychus albifrons Gray, Proc. Zool. Soc. London, p. 123, July 1864. "E. 
Africa (Capt. Speke)" hence probably coast of Tanganyika Territory. 



432 bulletin: museum of comparative zoology 

Heliophobitts angonicus Thomas 

Heliophobius angonicus Thomas, Ann. Mag. Nat. Hist. (8) 20: 314, Oct. 1917. 
Bua River, central Angoniland, Rhodesia. 

Heliophobius argenteocinereus Peters 

Heliophobius argenteocinereus Peters, Ber. liber Verhandl. K. Preuss. Akad. 
Wiss., Berhn, p. 259, Aug. 1846; Reise nach Mossambique, Saugethiere, 
p. 140, pi. 31, f. 2; pi. 35, f. 2, 1852. Tette, Mozambique. 
For skull characters distinguishing this from H. albifrons Gray, see Thomas, 

O., Ann. Mag. Nat. Hist. (6) 16: 241, Sep. 1895. 

Georychus pallidus Gray, Proc. Zool. Soc. London, p. 124, text-f. 2, 7, July 1864. 
"E, Africa (Capt. Speke)." 

Heliophobius marungensis Noack 

Heliophobius marungensis var. of argenteocinereus Noack, Zool. Jahrb., Syst., 
2: 223, pi. 9, f. 23-27, 7 May 1887. Marungu, Tanganyika Territory. 
A distinct species according to Thomas (Ann. Mag. Nat. Hist. (8) 20: 315, 
Oct. 1917). 

Heliophobius emini Noack 

Heliophobius emini Noack, Zool. Jahrb., Syst., 7: 559, pi. 18, f. 5-9, 23 Dec. 1893. 
Simba Muenne, Tanganyika Territory. 

Heliophobius kapiti (Heller) 

Georychus kapiti Heller, Smithsonian Misc. Coll., 52: 469, pi. 55 (skull), 24 Sep. 

1909. Potha, Kapiti Plains, Kenya Colony. 
Heliophobius kapiti HoUister, Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 161, pi. 38, 

lower three figs, (type skull), 1919. 

Heliophobius mottoulei (Schouteden) 

Myoscalops mottoulei Schouteden, Revue Zool. Africaine, 2: 303, text-f. 1-4 
(skull) , 30 May 1913. Kilongwe, near Lake Kisale, Congo Beige. 

Heliophobius robustus Thomas 

Heliophobius robustus Thomas, Ann. Mag. Nat. Hist. (7) 17: 179, Feb. 1906. 
Mpika, northeastern Rhodesia, 5000 ft. 

Heliophobius spalax Thomas 

Heliophobius spalax Thomas, Ann. Mag. Nat. Hist. (8) 6: 315, Sep. 1910. 
Taveta, foot of Mt. Kilimanjaro, Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 433 



Heterocephalus Riippell. Naked Mole-rats 

Heterocephalus Riippell, Mus. Senckenbergianum, 3: 99, pi. 8, f. 1; pi. 10, f. 

3 a-c, 1842. Genotype Heterocephalus glaber Riippell. 
Fornarina Thomas, Proc. Zool. Soc. London, for 1903, 2: 336, 1 Apl. 1904. 

Genotype Heterocephalus phiUipsi Thomas. On the status of this name see 

de Beaux, Atti Soc. Ital. di Sci. Nat. e Mus. Civ. Stor. Nat. Milano, 73: 

290, Dec. 1934. 

Review and critical discussion: de Beaux, O., Atti Soc. Ital. di Sci. Nat. e Mus. 
Civ. Stor. Nat., Milano, 73: 283-292, pi. 7-9, Dec. 1934. 

Heterocephalus glaber glaber Riippell 

Heterocephalus glaber Riippell, Mus. Senckenbergianum, 3: 99, 175, pi. 8, f. 1; 

pi. 10, f. 3 a-c, 1842. Shoa, southern Abyssinia. 
Heterocephalus phillipsi Thomas, Proc. Zool. Soc. London, p. 612, pi. 55, f. 1-4, 

Oct. 1885; ibid., p. 845, Apl. 1886. Ogardain, Gerlogobie, central Somali- 
land. 
Heterocephalus ansorgeiThomas, Proc. Zool. Soc. London, for 1903, 2: 336, 1 Apl- 

1904. Between Ngomeni and Kinani, Makindu Country, Kenya Colony. 
Regarded as a synonym of H. glaber by HolKster (Bull. U. S. Nat. Mus., 

no. 99, pt. 2, p. 160, pi. 6, 7, 1919). 
Heterocephalus dunni Thomas, Ann. Mag. Nat. Hist. (8) 4: 109, Aug. 1909. 

Wardairi, central Somaliland. 
Heterocephalus stygius G. M. Allen, Bull. Mus. Comp. Zool., 54: 444, Apl. 1912- 

Neumann's Boma, Northern Guaso Nyiro, Kenya Colony. 
A synonym of H. glaber (see HoIIister, N., Bull. U. S. Nat. Mus., no. 99, pt. 2, 

p. 160, pi. 6, 7, 1919). 
Heterocephalus glaber progrediens Lonnberg, Kimgl. SvenskaVet.-Akad.Handl., 

Stockholm, (2) 48: no. 5, p. 102, pi. 3, f. 2-5, 26 June 1912. Thornbush 

country north of Northern Guaso Nyiro, Kenya Colony. 
Regarded by HoIIister (Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 160, pi. 6, 7, 
1919) as a synonym of H. glaber. 

Heterocephalus glaber scorteccii de Beaux 

Heterocephalus glaber scorteccii de Beaux, Atti Soc. Ital. di Sci. Nat. e Mus. Civ. 

Stor. Nat., Milano, 73: 283, pi. 7, pi. 8, upper figs.; pi. 9, upper figs., Dec. 

1934. Gardo, northern Italian Somaliland. 



Family THRYONOMYID.E. Cane-rats 

Review: Thomas, O. On the animals known as "Ground-hogs" or "Cane- 
rats" in Africa. Ann. Mag. Nat. Hist. (9) 9: 389-392, Apl. 1922. 



434 bulletin: museum of comparative zoology 



Chceromys Thomas. Lesser Cane-rats 

Choeromys Thomas, Ann. Mag. Nat. Hist. (9) 9: 390, Apl. 1922. Genotype 
Thryonomys gregorianus Thomas. 

Chceromys gregorianus gregorianus (Thomas) 

Aulacodus gregorianus Thomas, Ann. Mag. Nat. Hist. (6) 13: 202, Feb. 1894. 
Lniji Reru River, Konu, Kikuyu, Kenya Colony, 5700 ft. 

Chceromys gregorianus pusillus (Heller) 

Thryonomys gregorianus pusillus Heller, Smithsonian Misc. Coll., 59: no. 16, 
p. 17, 5 July 1912. Ndi, Taita Hills, Kenya Colony. Regarded as a dis- 
tinct species by Hollister (Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 163, 1919). 

Chceromys harrisoni harrisoni (Thomas and Wroughton) 
Thryonomys harrisoni Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 
384, May 1907. Loka, 60 miles southwest of Fort Berkeley, Lado Enclave. 

Chceromys harrisoni congicus Thomas 

Chceromys harrisoni congicus Thomas, Ann. Mag. Nat. Hist. (9) 9: 390, Apl. 
1922. Lower Uele River, Congo Beige. 

Chceromys harrisoni rutshuricus (Lonnberg) 

Thryonomys rutshuricus Lonnberg, Kungl. Svenska Vet.-Akad. Handl., Stock- 
holm, (2) 58: art. 2, p. 78, 1 Sep. 1917. Rutshuru, Congo Beige. 
Doubtfully distinct from C. harrisoni according to Thomas. 

Chceromys harrisoni sclateri (Thomas) 

Thryonomys sclateri Thomas, Proc. Zool. Soc. London, p. 432, 1 Oct. 1897. 
Nyika plateau, Nyasaland, 6000-7000 ft. 

Chceromys logonensis Jeannin 

Choeromys logonensis Jeannin, Mamm. Sauvages du Cam^roun, (Encycl. Biol., 

16:) 178, 1936. Borders of Logone, Chad district, French Equatorial 

Africa (by implication). 



Thryonomys Fitzinger. Larger Cane-rats or "Ground-hogs" 

Thryonomys Fitzinger, Sitzb. K. Akad. Wiss. Wien, math.-nat. Kl., 56: sect. 1, 

p. 141, 1867. Genotype Aulacodus semipalmatus Heuglin. 
Aulacodus Temminck, Monographies de Mammalogie, 1: 245, pi. 25, 1827. 

Genotype Aulacodus swinderianus Temminck. Preoccupied by Aulacodus 

Eschscholtz, 1822, in Coleoptera. 
Triaulacodus Lydekker, Geogr. Hist. Mammals, pp. 91, 240, and footnote, 

1896. A substitute for Aulacodus Temminck (not Eschscholtz). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 435 

Thryonomys swindbrianus swinderiantjs (Temminck) 

Aulacodus swinderianus Temminck, Monographies de Mammalogie, 1: 248, 

1827. Sierra Leone (see Thomas, Ann. Mag. Nat. Hist. (9) 9: 391, Apl. 

1922). 
Range: Gambia to northern Nigeria. 

Thryonomys swinderianus angola Thomas 

Thryonomys swinderianus angolse Thomas, Ann. Mag. Nat. Hist. (9) 9: 392, 
Apl. 1922. Junction of Luando and Cuje Rivers, Angola, 4500 ft. 

Thryonomys logani Romer and Nesbitt 

Thryonomys logani Romer and Nesbitt, Ann. Mag. Nat. Hist. (10) 6: 687, Dec. 

1930. Based on skull and skeleton from valley of Wad Odiemet, 140 km. 

southwest of Fort Leperrine, 4° 30' E., 22° N., Tanezrouft region, central 

Sahara, probably of "early Recent" age. 

Thryonomys swinderianus raptorum Thomas 

Thryonomys svnnderianus raptorum Thomas, Ann. Mag. Nat. Hist. (9) 9: 392, 
Apl. 1922. Lagos, Nigeria. 
Range: Forest region of lower Nigeria and Gold Coast, east through the 
Congo region. 

Thryonomys swinderianus variegatus (Peters) 

Aulacodus variegatus Peters, Reise nach Mossambique, Saugeth., p. 138, 1852. 

Mozambique (Tette, Macauga, Sena, and Boror). 
Aulacodus seinipalmalus Heuglin, Nova Acta Acad. Cses. Leop. -Carol., Halle, 

31: art. no. 7, p. 5, pi. 1, f. 1-4, 1864. West of the Djur River, to Fertit 

and Niam-niam country, upper Bahr-el-Ghazal, Anglo-Egyptian Sudan. 
Aulacodus calamophagus de Beerst in Pousargues, Bull. Mus. d'Hist. Nat. 

Paris, 3: 160, 1897. Saint Jacques de Lusaka, Nyasaland. 
Range: Bahr-el-Ghazal and Uganda to eastern Cape of Good Hope. 



Family CTENODACTYLID.E 

Ctenodactylus Gray 

Ctenodactylus Gray, Spicilegia Zoologica, 2: 10, 1 Aug. 1830. Genotype Cteno- 
dactylus massonii Gray. 

Palmer (North Amer. Fauna, no. 23, p. 303, 23 Jan. 1904) agrees with Lataste 
that Gundi as used by Shaw, and by Fischer is a vernacular use of the 
name and not a generic term. 



436 bulletin: museum of comparative zoology 

Ctenodactylus gundi gundi (Rothmann). Gundi 

Mus gundi Rothmann, in Schlozer's Briefwechsel, 1: 339, 1776. Hills of Ghar- 
ian ("Garean"), 80 km. south of Tripoli, Tunisia (see Thomas, O., A note 
on the type-locality and geographical races of the gundi {Ctenodactylus 
gundi Rothm.), Novitates Zool., 27: 506-507, Nov. 1920). 
Range: Northwestern Tripoli and southern Tunisia. 

Ctenodactylus gundi massonii Gray. Biskra Gundi 

Ctenodactylus massonii Gray, Spicilegia Zoologica, 2: 11, pi. 10, 1 Aug. 1830. 

"Cape of Good Hope," but doubtless North Africa. 
Ctenodactylus typicus A. Smith, South African Quart. Journ., 2: 151, 1834. 

"Barbary." 
Range: Southern slopes of the Algerian Atlas, Biskra. 

Ctenodactylus joleaudi Heim de Balsac 

Ctenodactylus joleaudi Heim de Balsac, Suppl. au Bull. Biol, de France et de 
Belgique, Paris, 21: 315 (brief description); 378, f. 10, no. 7 (skull); 381, 
f. 12, A (skull), 405, 1936; Bull. Soc. Zool. de France, 62: no. 5, p. 329, 14 
Aug. 1937 (formal description). Beni Ounif, Jebel Melias, Algeria. 

Ctenodactylus vali Thomas 

Ctenodactylus vali Thomas, Proc. Zool. Soc. London, 2:11, Oct. 1902. Wadi Bey, 
northwest of Bonjem, Tripoli. 
Range: Middle Tripoli. 

Felovia Lataste 

Felovia Lataste, Le Naturaliste, 8: no. 36, p. 287, 15 June 1886. Proposed as a 
subgenus of Massoutiera with type Felovia vse Lataste; recognized as a 
valid genus by Thomas, O., Novitates Zool., 20: 31, 24 Feb. 1913; St. 
Leger, Proc. Zool. Soc. London, p. 978, 30 Sep. 1931. 

Felovia v^ Lataste 

Felovia vse Lataste, Le Naturaliste, 8: no. 36, p. 287, 15 June 1886. Medina 
district, upper Senegal River, Senegal. 



Massoutiera Lataste 

Massoutiera Lataste, Le Naturaliste, 7: no. 3, p. 21, 1 Feb. 1885. Genotype 
Ctenodactylus mzabi (Lataste.) 

Massoutiera mzabi mzabi (Lataste) 

Ctenodactylus ynzabi Lataste, Bull. Soc. Zool. de France, 6: 314, text-f. 1, 3, 5-7, 
1881. Gardaia, Mzab, Algeria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 437 

Massoutiera mzabi harterti Thomas 

Massoutiera harterti Thomas, Novitates ZooL, 20: 31, 24 Feb. 1913. Oued Mya, 
south of Fort Miribel, western Sahara, about 28° 30' N., 3° E. 

Massoutiera mzabi rothschildi Thomas and Hinton 

Massoutiera rothschildi Thomas and Hinton, Novitates ZooL, 28: 11, 14 May 
1921. Mt. Baguezan, Asben, northern Nigeria. 



Pectinator Blyth 

Pedinator Blyth, Journ. Asiatic Soc. Bengal, for 1855, (2) 24: 294, 1856. Geno- 
type Pectinator spekei Blyth. 

Petrobates Heuglin, Zeitschr. f. Gesammt. Naturw., Berlin, 16: 413, 1860. 
Genotype Pectinator spekei Blyth. 

Pectinator spekei spekei Blyth 

Pectinator spekei Blyth, Journ. Asiatic Soc. Bengal, for 1855, (2) 24: 294, pi. 
2, f. 1, 1856. [Between Goree Bunder and Nogal], Somaliland. 

Pectinator spekei leger^ de Beaux 

Pectinator spekei legerse de Beaux, Atti Soc. Ital. Sci. Nat.eMus. Civ. Stor. Nat., 
Milano, 73: 293, Oct. 1934. Assab, Eritrea. 

Pectinator spekei meridionalis de Beaux 

Pectinator spekei meridionalis de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. 
Stor. Nat., Milano, 61: 27, Feb. 1922. Dolo, Somaliland. 



Petromus a. Smith. Dassie Rats 

Petromus A. Smith, South African Quart. Journ., 1: no. 5, p. 10-11 (misprint 
for p. 2), Oct. 1831. Genotype Petromus typicus A. Smith. 

Petromys A. Smith, South African Quart. Journ., 2: no. 2, p. 146, Jan.-Mar. 
1834; Illustr. ZooL South Africa, Mamm., pt. 9, pi. 20; pi. 21, f. 1 a-e, Jan. 
1840 (emendation). 

Petromus typicus typicus A. Smith 

Petromus typicus A. Smith, South African Quart. Journ., 1: no. 5, p. 11 (mis- 
print for p. 2), Oct. 1831. Mountains toward the mouth of Orange River, 
Little Namaqualand, Cape of Good Hope. 

Petromus typicus cunealis Thomas 

Petromys cunealis Thomas, Proc. ZooL Soc. London, p. 307, 29 Apl. 1926. 
Cunene Falls, northwestern Ovamboland, Southwest Africa, 3350 ft. 



438 bulletin: museum of comparative zoology 

Petromus typicus marjori^ Bradfield 

Petromys typicus marjoriae Bradfield, Description of New Races of Kalahari 
Birds and Mammals, (privately printed), Benoni, South Africa, p. 1, 26 
Sep. 1935; The Auk, 53: 131, Jan. 1936 (reprinted). Khan River, South- 
west Africa. 

Petromus typicus tropicalis Thomas and Hinton 

Petromys typicus tropicalis Thomas and Hinton, Proc. Zool. Soc. London, p. 
241, 3 Apl. 1925. Karibib, Southwest Africa. 



Family PEDETIDiE. 

Pedetes lUiger. Spring-haas 

Yerhua Forster, Kongl. Svenska Vet.-Akad. Handl. Stockholm, (1) 39: 108, 

pi. 3, 1778 (in part, as to Yerhua capensis Forster). 
Pedetes Illiger, Prodromus Syst. Mamm. et Avium, p. 81, 1811. Genotype 

Dipus cafer (Gmelin) = Mus cafer Pallas. 
Helamys F. Cuvier, Regne Animal, ed. 1,*1 : 202, footnote, 1817. Genotype Mus 

cafer Pallas. 
Helamis F. Cuvier, Dictionnaire des Sci. Nat., 20: 341, 1821, variant. 
Pedestes Gray, List Spec. Mamm. Brit. Mus., p. 130, 13 May 1843 (lapsus cal.). 

Pedetes cafer cafer (Pallas) 

Mus cafer Pallas, Nov. Spec. Quadrup. e Glir. Ord., p. 87, 1778. Cape of Good 

Hope. 
Yerbua capensis Forster, Kongl. Svenska Vet.-Akad. Handl., Stockholm, (1) 

39: 108, pi. 3, 1778. Cape of Good Hope. 
Pedetes typicus A. Smith, South African Quart. Journ., 2: 169, 1834. Eastern 

districts of Cape Colony. 

Pedetes cafer angola Hinton 

Pedetes angolse Hinton, Ann. Mag. Nat. Hist. (9) 6: 102, July 1920. Cholinde, 
20 miles northeast of Bihe, Angola. 

Pedetes cafer damarensis Roberts 

Pedetes caffer damarensis Roberts, Ann. Transvaal Mus., 11: 261, 14 Sep. 1926. 
Quickborn, Okahandja, Southwest Africa. 

Pedetes cafer dentatus Miller 

Pedetes cafer dentatus Miller, Proc. Biol. Soc. Washington, 40: 113, 26 Sep. 1927. 
Near Dodoma, Tanganyika Territory. 



ALLEj/: CHECKLIST OF AFRICAN MAMMALS 439 

Pedetes cafer orangi^ Wroughton 

Pedetes cafer orangise Wroughton, Ann. Mag. Nat. Hist. (7) 20: 32, July 1907. 
Aberfeldy district. Orange Free State. 

Pedetes cafer salin^e Wroughton 

Pedetes cafer salinse Wroughton, Ann. Mag. Nat. Hist. (7) 20: 32, 33, July 1907. 
Woodbush, Zoutpansberg district, northwestern Transvaal. 

Pedetes cafer taboRjE G. M. Allen and Loveridge 

Pedetes cafer taborse G. M. Allen and Loveridge, Proc. Boston Soc. Nat. Hist., 
38: 438, Dec. 1927. Tabora, Tanganyika Territory. 

Pedetes sxjrdaster surdaster Thomas 

Pedetes surdaster Thomas, Ann. Mag. Nat. Hist. (7) 9: 440, June 1902. More- 
dat, mile 365 of Uganda Railway, Naivasha Province, Kenya Colony. 
A distinct species, according to Hollister, Bull. U. S. Nat. Mus., no. 99, pt. 2, 
p. 156, 1919. 

Pedetes surdaster currax Hollister 

Pedetes surdaster currax Hollister, Smithsonian Misc. Coll., 68: no. 10, p. 3, 16 

Jan. 1918; Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 157, pis. 39, 40 (skull), 

1919. Kabalolot Hill, Sotik, Kenya Colony. 

Pedetes surdaster larvalis Hollister 

Pedetes surdaster larvalis HoUister, Smithsonian Misc. Coll., 68: no. 10, p. 2, 16 
Jan. 1918; Bull. U. S. Nat. Mus., no. 99, pt. 2, p. 157, pis. 3, 4, 5 (lower 
figs.), 1919. Ulukenia Hills, Athi Plains, Kenya Colony. 



Family HYSTRICID^. Old World Porcupines 
Subfamily ATHERURIN^ 

Atherurus G. Cuvier. Brush-tailed Porcupines 

Atherurus G. Cuvier, Dictionnaire des Sci. Nat., 59: 483, 1829. Genotype not 
given but fixed as Hystrix fasci&ulata Shaw of Malacca = Hystrix viacroura 
Linnaeus. 

Atherura G. Cuvier, Regne Animal, ed. 2, 1: 215, 1829 (emendation). 

Atherurus africanus (Gray) 

Atherura africana Gray, Ann. Mag. Nat. Hist. (1) 10: 261, Dec. 1842. Sierra 
Leone. 



440 bulletin: museum of comparative zoology 

Atherurus armatus Gervais 

Atherura arrnata Gervais, Les Trois Regnes de la Nature, Hist. Nat. des Mam- 
mif., p. 334, 1854. Gambia. 

Atherurus burrowsi Thomas 

Atherura burrowsi Thomas, Ann. Mag. Nat. Hist. (7) 9: 270, Apl. 1902. Lower 
Aruwimi River, Congo Beige. 

Atherurus centralis Thomas 

Atherura centralis Thomas, Ann. Mag. Nat. Hist. (6) 15: 89, Jan. 1895. Mon- 
buttu, = Niangara, Congo Beige. 

Atherurus turneri St. Leger 

Atherura turneri St. Leger, Ann. Mag. Nat. Hist. (10) 10: 231^ Sep. 1932. 
Kakumega Forest, near Kaimosi, Kenya Colony. 



Subfamily HYSTRICIN^ 

Hystrix Linnfeus. Crested Porcupines 

Hystrix Linnaeus, Syst. Nat., ed. 10, 1 : 56, 1758. Included five species; genotype 

Hystrix cristata Linnseus. 
Oedocephalus Gray, Proc. Zool. Soc. London, p. 308, Sep. 1866. Genotype Acan- 

thion cuvieri Gray = Hystrix cristata Linnseus, in part. 

See Muller, Ferdinand. Besprechung einiger Hystrix-Schiidel aus Deutsch 
Ost-Afrika. Sitzb. Ges. Naturf. Freunde, Berlin, p. 309-315, 1910. 

The status of all the described forms is in need of revision; probably those 
that are recognizable should be regarded as subspecies of H. cristata Linnaeus. 

Hystrix aerula Thomas 

Hystrix aerula Thomas, Ann. Mag. Nat. Hist. (9) 16: 196, July 1925. Aouderas, 
Asben, French West Africa. 

Hystrix afric^australis africvEaustralis Peters 

Hystrix africse australis Peters, Reise nach Mossambique, Saugeth., p. 170, pi. 

32, f. 6, 7 (skull), 1852. Querimba coast and Tette, Mozambique. 
Hystrix capensis Grill, Kongl. Svenska Vet.-Akad. Handl., Stockholm, (2) 2: 

no. 10, p. 19, 1858. Salt River, near Knysna, Cape of Good Hope. 

Hystrix afric^australis prittwitzi F. Miiller 

Hystrix africse-australis prittwitzi Muller, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 311, text-f. 2, 313, Oct. 1910. Tabora, Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 441 

Hystrix afric^australis zuluensis Roberts 

Hystrix africse-australis zuluensis Roberts, Ann. Transvaal Mus., 18: 240, 21 
Feb. 1936. Umfolosi River, Transvaal. 

Hystrix cristata Linnaeus 

Hystrix cristata Linnaeus, Syst. Nat., ed. 10, 1: 56, 1758. "Asia" but restricted 

to "Rome" by Thomas, where probably introduced from North Africa 

(Cabrera). 
H[ystrix] cristata europsea Kerr, Linnaeus's Animal Kingdom, p. 213, 1792. No 

locality. 
Acanthion cuvieri Gray, Proc. Zool. Soc. London, p. 102, 5 Aug. 1847 (in part). 

Locality unknown, but given in 1866 (Proc. Zool. Soc. London, p. 308, 

under Oedocephalus cuvieri) as North Africa. 
Range: Northern Africa, from at least eastern Morocco to Egypt, and south 

to the Sahara. 

Hystrix galeata galeata Thomas 

Hystrix galeata Thomas, Ann. Mag. Nat. Hist. (6) 11 : 230, Mar. 1893. Lamu,. 
Kenya Colonj'. 

Hystrix galeata ambigua Lonnberg 

Hystrix galeata ambigua Lonnberg, Wiss. Ergebn. Schwedisch. Zool. Exped. 

Kilimandjaro unter Leitung v. Sjostedt, Mamm., p. 29, pi. 5, f. 3 (skull), 

1908. Kibonoto, Tanganyika Territory. 

Hystrix galeata conradsi F. Miiller 

Hlystrix] galeata conradsi Miiller, Sitzb. Ges. Naturf. Freunde, Berlin, p. 314, 

Oct. 1910. Mwanza and Ukerewe Island, Victoria Nyanza, Tanganyika 

Territory. 

Hystrix galeata lademanni F. Miiller 

H[ystrix] galeata ludemanni (sic) Miiller, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 314, Oct. 1910. Kondoa Irangi, and lower Bubu southwest of Mt. Kili- 
manjaro, Tanganyika Territory. 

Hystrix galeata lonnbergi F. Miiller 

Hystrix galeata lonnbergi Miiller, Sitzb. Ges. Naturf. Freunde, Berlin, p. 315, 
Oct. 1910. Mt. Kilimanjaro, Tanganyika Territory. 

• 

Hystrix galeata somalensis Lonnberg 

Hystrix galeata somalensis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 48: art. 5, p. 109, 26 June 1912. Near Nyoro, Northern 

Guaso Nyiro, Kenya Colony. 



442 bulletin: museum of comparative zoology 

Hystrix occidanea Cabrera 

Hystrix occidanea Cabrera, Bol. Real Soc. Espanola Hist. Nat., Madrid, 24: 
220, text-f. 1, 28 Apl. 1924; Trab. Mus. Nac. de Cien. Nat., Madrid, zool. 
ser., no. 57, p. 289, text-f. 34 B (skull), 30 Dec. 1932. Mogador, Morocco. 
Range: Southeastern Morocco. 

Hystrix senegalica F. Cuvier 

Hystrix senegalica F. Cuvier, Mem. Mus. d'Hist. Nat., Paris, 9: 430, 1822. 
Senegal (based on a young specimen). 

Hystrix stegmanni F. Miiller 

Hystrix stegmanni F. Miiller, Arch. f. Naturgesch., 76: sect. A, pt. 1, no. 2, 
p. 186, Oct. 1910; Sitzb. Ges. Naturf. Freunde, Berlin, p. 312, text-f. 3, 
Oct. 1910. Kissenji, northeast of Lake Kivu, Congo Beige. 



Order HYRACOIDEA. Dassies 
Family PROCAVIID^. Dassies or Hyraxes 

Review: Hahn, Herbert. Die Familie der Procaviidse. Zeitschr. f. Sauge- 
tierk., 9: 207-358, 69 text-f., pi. 13-16 (maps), 30 Dec. 1934. 

Hatt, Robert T. Hyraxes collected by the American Museum Congo Expe- 
dition. Bull. Amer. Mus. Nat. Hist., 72: 117-141, text-f. 1-3, pi. 18-22, 28 
Aug. 1936. 

Dendrohyrax Gray. Tree Hyraxes 

Dendrohyrax Gray, Ann. Mag. Nat. Hist. (4) 1 : 48, Jan. 1868. Included three 
species; genotype fixed as Hyrax arboreus A. Smith (see Sclater, W. L., 
Mamm. South Africa, 1: 310, 1900). 

Dendrohyrax arboreus arboreus (A. Smith) 

Hyrax arboreus A. Smith, Trans. Linn. Soc. London, 15: 468, 1827. Forests, 

Cape of Good Hope. 
Dendrohyrax arboreus Gray, Ann. Mag. Nat. Hist. (4) 1 : 49, Jan. 1868. 

Dendrohyrax arboreus adolfi-friederici (Brauer) 

Procavia {Dendrohyrax) adolfi-friederici Brauer, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 132, Feb. 1913. Bugoie Forest, north of Lake Kivu, Congo 

Beige. 
Procavia (Heterohyrax) schubotzi Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 134, Feb. 1913. Bugoie Forest, north of Lake Kivu, Congo Beige. 
Procavia (Heterohyrax) helgei Lonnberg and Gyldenstolpe, Arkiv f. Zool., 17 B: 

no. 9, p. 4, 14 May 1925. Mt. Mikeno, Birunga Volcanoes, Congo Beige. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 443 

Dendrohyrax arboreus bettoni (Thomas and Schwann) 

Procavia bettoni Thomas and Schwann, Abstract Proc. Zool. Soc. London, no. 6, 

p. 23, 26 Apl. 1904; Proc. Zool. Soc. London, 1: 463, 2 Aug. 1904. Rogoro, 

mile 346 of Uganda Railway, Kenya Colony. 
Procavia (Dendrohyrax) scheffleri Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 130, Feb. 1913. Kibwezi, Kenya Colony. 
Procavia (Dendrohyrax) vilhelmi Lonnberg, Arkiv f. Zool., 10: no. 12, p. 26, 

19 May 1916. Donya Sabuk, Kenya Colony. 

Dendrohyrax arboreus braueri Hahn 

Dendrohyrax arboreus braueri Hahn, Zeitschr. f. Saugethierk., 8: 278, 20 Dec. 
1933. Elizabethville, Congo Beige. 

Dendrohyrax arboreus crawshayi (Thomas) 

Procavia (Dendrohyrax) crawshayi Thomas, Proc. Zool. Soc. London, p. 178, 

1 June 1900. Western slope of Mt. Kenya, Kenya Colony, 10,000 ft. 
Dendrohyrax crawshayi laikipia Dollman, Ann. Mag. Nat. Hist. (8) 8: 131, 

July 1911. Rumruti, Laikipia Plateau, Kenya Colony. 

Dendrohyrax arboreus mimus (Thomas) 

Procavia mima Thomas, Ann. Mag. Nat. Hist. (7) 6: 387, Oct. 1900. Northern 
Nyasaland, Nyasa-Tanganyika plateau. 

Dendrohyrax arboreus ruwenzorii (Neumann) 

Procavia (Dendrohyrax) ruwenzorii Neumann, Proc. Zool. Soc. London, 2: 143, 
Oct. 1902. Mt. Ruwenzori, Congo Beige, 11-15,000 ft. 

Dendrohyrax arboreus stuhlmanni (Matschie) 

Procavia stuhlmanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 110, 
1892. Bukoba, Victoria Nyanza, Tanganyika Territory. 

Dendrohyrax scheelei Matschie, Deutsch-Ost-Afrika, 3: Saugeth., p. 90, foot- 
note, 1895. Khutu, near Ruaha River, Tanganyika Territory. 

Dendrohyrax dorsalis dorsalis (Fraser) 

Hyrax dorsalis Fraser, Proc. Zool. Soc. London, for 1852, p. 99, pi. 33 (Mamm.), 
23 May 1854. Island of Fernando Po. 

Dendrohyrax dorsalis emini Thomas 

Dendrohyrax emini Thomas, Ann."" Mag. Nat. Hist. (5) 20: 440, Dec. 1887. 

Tingasi, Monbuttu, Congo Beige. 
Dendrohyrax rubriventer Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 295, 
10 Sep. 1917. Kalumenda-Beni, Congo Beige. 
Omitted by Hahn (1934) but probably a synonym of D. dorsalis emini 
Thomas according to Hatt (Bull. Amer. Mus. Nat. Hist., 72: 118, 28 Aug. 
1936). 



444 bulletin: museum of comparative zoology 

Dendrohyrax beniensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 295, 10 
Sep. 1917. Beni Zambo, Kalumenda-Beni, Congo Beige. 

Dendrohyrax congoensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 295, 
10 Sep. 1917. Beni (Cartoushi), Congo Beige. 

Dendrohyrax brevimaculatus Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 296, 
10 Sep. 1917. Lesse, Ituri district, Congo Beige. 

Dendrohyrax dorsalis latrator (Thomas) 

Procavia emini latrator Thomas, Ann. Mag. Nat. Hist. (8) 5: 285, Mar. 1910. 
, Batempa, upper Sankuru River, Congo Beige. 

Although by Hahn regarded as a synonym of D. d. emini. Hatt (Bull. Amer. 
Mus. Nat. Hist., 72: 118, 28 Aug. 1936) believes it probably recognizable. 

Dendrohyrax dorsalis marmota (Thomas) 

Procavia {Dendrohyrax) marmota. Thomas, Proc. Zool. Soc. London, 2: 88, 
1 Oct. 1901. Mengo, north of Entebbe, Uganda. 

Dendrohyrax dorsalis nigricans (Peters) 

Hyrax nigricans Peters, Sitzb. Ges. Naturf. Freunde, Berlin, p. 10, 1879. 

Chinchoxo, north of the mouth of the Congo, Kabinda, Angola. 
Procavia {Dendrohyrax) tessmanni Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 411, July 1912. Akonangi, Spanish Guinea. 
Pr{ocavia\ {Dendrohyrax) adametzi Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 412, July 1912. Lake Barombi, Johann-Albrechts Heights, Cameroons. 
Pr[ocavia] {Dendrohyrax) adametzi zenkeri Brauer, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 38, Jan. 1914. Southern Cameroons, specimens from Bipindi, 

Bibundi and Yaunde. 

Dendrohyrax dorsalis sylvestris (Temminck) 

Hyrax sylvestris Temminck, Esquisses Zool. sur la Cote de Guine, p. 182, 1853. 

Guinea coast to Ashanti, West Africa. 
Hyrax stampflii Jentink, Notes Leyden Mus., 8: 210, Oct. 1886. Chap Hill, 

opposite Schlieffelinsville, Junk River, Liberia. 
Pr[ocavia] {Dendrohyrax) aschantiensis Brauer, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 36, Jan. 1914. Abetifi, Ashanti, northwest of Begoro, Gold 

Coast. 

Dendrohyrax validus validus True 

Dendrohyrax validus True, Proc. U. S. Nat. Mus., 13: 228, 16 Sep. 1890. Mt. 
Kilimanjaro, Tanganyika Territory. 

Dendrohyrax validus neumanni (Matschie) 

Procavia neumanni Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 112, 1893. 

Zanzibar, Pangani forest. 
Dendrohyrax adersi Kershaw, Ann. Mag.. Nat. Hist. (9) 13: 557, May 1924. 

Wete, Ngesi, Pemba Island, off Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 445 

Dendrohyrax validus terricola Mollison 

Dendrohyrax terricola Mollison, Zool. Anzeiger, 29: 423, 26 Sep. 1905. Near 

Monga, eastern Usambara, Tanganyika Territory. 
Dendrohyrax terricola schusteri Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 296, 10 Sep. 1917. Uluguru Mts., eastern Tanganyika Territory. 

Dendrohyrax validus vosseleri Brauer 

Dendrohyrax terricola vosseleri Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 296, 10 Sep. 1917. Amani on the Sigi, Tanganyika Territory. 



Heterohyrax Gray Gray Hyraxes 

Heterohyrax Gray, Ann. Mag. Nat. Hist. (4) 1 : 50, Jan. 1868. As a subgenus of 
Dendrohyrax; type Dendrohyrax blainvillii Gray = Hyrax syriacus Schreber. 

Heterohyrax antine^ (Heim de Balsac and Begouen) 
Procavia {Heterohyrax) antinese Heim de Balsac and Begouen, Bull. Mus. 
d'Hist. Nat., Paris, (2) 4: 479, June 1932. Ahaggar, central Sahara. 

Heterohyrax chapini (Hatt) 

Procavia chapini Hatt, Amer. Mus. Novitates, no. 594, p. 1, 26 Jan. 1933. 

Summit of Loadi Hill, 5 km. southwest of Matadi, Lower Congo district, 

Congo Beige. 
As to status, see Hatt, Bull. Amer. Mus. Nat. Hist., 72: 132, text-f. 2, pi. 22, 
lower fig., 28 Aug. 1936. 

Heterohyrax syriacus syriacus (Schreber) 

Hyrax syriacus Schreber, Saugthiere, 4: 923, pi. 240B, 1792. Mt. Lebanon, Mt. 

Sinai; Ras Mohammed, Ethiopia. 
Hyrax brucei Gray, Ann. Mag. Nat. Hist. (4) 1 : 44, Jan. 1868. Abyssinia. 
Dendrohyrax blainvillii Gray, Ann. Mag. Nat. Hist. (4) 1 : 50, Jan. 1868. No 

locality. 
H[yrax] irroratus Gray, Ann. Mag. Nat. Hist. (4) 3: 242, Mar. 1869. Abyssinia. 

Heterohyrax syriacus albipes Hollister 

Heterohyrax brucei albipes Hollister, Proc. Biol. Soc. Washington, 35: 135, 17 
Oct. 1922. Telek River, Loita Plains, Kenya Colony. 
Regarded by Hahn (1934, p. 276) as a synonym of H. syriacus hindei 
(Wroughton). 

Heterohyrax syriacus bakeri (Gray) 

Dendrohyrax bakeri Gray, Ann. Mag. Nat. Hist. (4) 14: 133, Aug. 1874. Latiko, 
3°N. 
The only Heterohyrax on west bank of the Nile. 



446 bulletin: museum of comparative zoology 

Heterohyrax syriacus bocagei (Gray) 

Hyrax bocagei Gray, Ann. Mag. Nat. Hist. (4) 3: 242, Mar. 1869. Angola. 
Dendrohyrax grayi Bocage, Jorn. Sci. Math., Phys., e Nat., Lisboa, (2) 1: 190, 
1889. Quissange, Angola. 

Heterohyrax syriacus dieseneri Brauer 

Heterohyrax hrucei dieseneri Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 

298, 10 Sep. 1917. Guta, Speke Gulf, Uschaschi, Tanganyika Territory, 

Ikoma, and Ukerewe Island, Lake Victoria. 

Heterohyrax syriacus frommi (Brauer) 

Procavia (Heterohyrax) frommi Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 136, Feb. 1913. Mahenge district, southeast of Iringa, Tanganyika 

Territory. 

Heterohyrax syriacus granti (Wroughton) 

Procavia hrucei granti Wroughton, Ann. Mag. Nat. Hist. (8) 5: 109, Jan. 1910. 
Woodbush, Transvaal. 

Heterohyrax syriacus hindei (Wroughton) 

Procavia hrucei hindei Wroughton, Ann. Mag. Nat. Hist. (8) 5: 107, Jan. 1910. 

Fort Hall, Kenya Colony. 
Procavia hrucei maculata Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 

6, 16 Feb. 1910. Lukenya Mt., Kenya Colony. 

Heterohyrax syriacus kempi (Thomas) 

Procavia hrucei kempi Thomas, Ann. Mag. Nat. Hist. (8) 5: 200, Feb. 1910. 
Elgonyi, Mt. Elgon, Kenya Colony, 7000 ft. 

Heterohyrax syriacus lademanni Brauer 

Heterohyrax lademanni Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 298, 

10 Sep. 1917. Livingstone Mts., at Mwakete, Tanganyika Territory; 

other localities mentioned are Upangwa, Msangwa and Mlando. 

Heterohyrax syriacus manningi (Wroughton) 

Procavia hrucei manningi Wroughton, Ann. Mag. Nat. Hist. (8) 5: 109, Jan. 
1910. Mlanji, Nyasaland. 

Heterohyrax syriacus mossambicus (Peters) 

Hyrax mossambicus Peters, Sitzb. Ges. Naturf. Freunde, p. 25, 1869. Caba- 
geira peninsula, Mozambique. 

Heterohyrax syriacus munzneri (Brauer) 

Procavia (Heterohyrax) munzneri Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 137, Feb. 1913. Bismarckburg, near Lake Tanganyika, Tanganyika 

Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 447 

Heterohyrax munzneri ruckwaensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 299, 10 Sep. 1917. Rukwa Lake, and Rudshugi Post, Tanganyika 
Territory. 

Heterohyrax syriacus princeps (Thomas) 

Procavia brucei princeps Thomas, Ann. Mag. Nat. Hist. (8) 5: 199, Feb. 1910. 

Bourka, Arusi country, near Lake Zuai, Abyssinia, 5000 ft. 
Heterohyrax arboricola Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 297, 10 

Sep. 1917. Dembel Lake, north of Abbaja Lake, Galla, central Abyssinia. 

Heterohyrax syriacus prittwitzi Brauer 

Heterohyrax brucei prittwitzi Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 
299, 10 Sep. 1917. No type locality designated, but Kilimatinde, Tan- 
ganyika Territory, may be selected from those named. 

Heterohyrax syriacus pumilus (Thomas) 

Procavia pumila Thomas, Ann. Mag. Nat. Hist. (8) 5: 201, Feb. 1910. Fifty 
miles southeast of Berbera, British Somaliland. May be a distinct species. 

Heterohy'rax syriacus ruddi (Wroughton) 

Procavia brucei ruddi Wroughton, Ann. Mag. Nat. Hist. (8) 5: 108, Jan. 1910. 
Tambarara, Gorongoza Mts., Mozambique. 

Heterohyrax syriacus rudolfi (Thomas) 

Procavia pumila rudolfi Thomas, Ann. Mag. Nat. Hist. (8) 5: 202, Feb. 1910. 

North end of Lake Rudolf, Abyssinia, 2000 ft. 
Procavia brucei borana Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 66, Jan. 1912. 

North of Northern Guaso Nyiro, east of Marsabit road, Kenya Colony. 

Heterohyrax syriacus somalicus (Thomas) 

P[rocavia] brucei somalica Thomas, Proc. Zool. Soc. London, p. 71, June 1892. 

Berbera, Somaliland. 
Heterohyrax brucei hararensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 

297, 10 Sep. 1917. Near Harar, Galla country, Abyssinia. 
Heterohyrax pumila webensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 

297, 10 Sep. 1917. Web River, southern Abyssinia. 

Heterohyrax syriacus ssonge^e Brauer 

Heterohyrax brucei ssongea? Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 300, 
10 Sep. 1917. No type locality specified; specimens noted from Ssongea, 
North Matengo, Papaj-Nyassa, Rhuira, Ussangira, southern Tanganyika 
Territory, of which the first may be designated as type locality. 



448 bulletin: museum of comparative zoology 

Heterohyrax syriacus thomasi (Neumann) 

Procavia thomasi Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 240, 1901. 

Kaffa, Gimirra, and Binescho, Omo Basin and basin of the Gelo (one of the 

sources of the Sobat), Anglo- Egyptian Sudan. 

Heterohyrax syriacus viCTORiA-NJANSiE Brauer 

Heterohyrax brucei vicloria-njansse Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 299, 10 Sep. 1917. No type locality designated; specimens from Nyan- 
gesi, Mwanza, and Mondo, near Lake Victoria, Tanganyika Territory, of 
which Mwanza may be designated type locality. 



Procavia Storr. Large-toothed Hyraxes 

Procavia Storr, Prodromus Methodi Mamm., p. 40, pi. B, 1780. Thomas, Proc. 

Zool. Soc. London, p. 51, June 1892. Genotype Cavia capensis Pallas. 
Hyrax Hermann, Tabula Affinitatum Anim., p. 115, 1783. Genotype Cavia 

capensis Pallas. 
Euhyrax Gray, Ann. Mag. Nat. Hist. (4) 1 : 46, Jan. 1868. Genotype Hyrax 

habessinicus Hemprich and Ehrenberg. 

Procavia capensis capensis (Pallas) 

Cavia capensis Pallas, Miscellanea Zool., p. 30, pis. 3, 4, 1766. Cape of Good 

Hope. 
Dendrohyrax semicircularis Gray, Cat. Carnivorous, Pachydermatous and 

Edentate Mamm. Brit. Mus., p. 285, 1869. No locality. 
P[rocavia] capensis coombsi Roberts, Ann. Transvaal Mus., 10: 76, 31 Jan. 1924. 

Hennops River, Pretoria, Transvaal. 
Procavia capensis natalensis Roberts, Ann. Transvaal Mus., 10: 76, 31 Jan. 

1924. Piggs Peak, Swaziland, Transvaal. 

Procavia capensis chiversi Roberts 

Procavia capensis chiversi Roberts, Ann. Transvaal Mus., 19: 101, 8 Nov. 1937. 
Mt. Fletcher, East Griqualand, south of Basutoland, South Africa. 

Procavia capensis letab^ Roberts 

Procavia capensis letabse Roberts, Ann. Transvaal Mus., 19: 102, 8 Nov. 1937. 
Mokeetsi, northeastern Transvaal. 

Procavia capensis marlothi Brauer 

Pr{ocavia\ capensis marlothi Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 33, 

Jan. 1914; Kranshoek and Nieuwveld Mts. near Beaufort West, northern 

Cape of Good Hope (Cape Colony). 

Procavia capensis oRANGiiE Roberts 

Procavia capensis orangise Roberts, Ann. Transvaal Mus., 19: 101, 8 Nov. 1937. 
Meadows, Orange Free ^tate. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 449 

Procavia capensis schultzei Brauer 

Pr[ocavia] capensis schultzei Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 32, 
Jan. 1914. Chamis, Great Namaqualand, Southwest Africa. 

Procavia capensis welwitschii (Gray) 

Hyrax welwitschii Gray, Ann. Mag. Nat. Hist. (4) 1: 43, Jan. 1868. Rocky 

banks of River Maiomba, Mossamedes district, Angola. 
Procavia flavimaculata Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 303, 10 

Sep. 1917. Kaokoveld, northwestern Southwest Africa. 

Procavia capensis windhuki Brauer 

Pr[ocavia\ capensis windhuki Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 30, 

Jan. 1914. Dobra and neighboring localities near Windhuk, Southwest 

Africa. 
Pr[ocavia] capensis waterbergensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 33, Jan. 1914. Waterberg, Southwest Africa. 
Prlocavia] capensis reuningi Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 31, 

Jan. 1914. Fiirstenwalde, Lichtenstein and neighborhood, near Windhuk, 

Southwest Africa. 

Procavia habessinica habessinica (Hemprich and Ehrenberg) 

Hyrax habessinicus Hemprich and Ehrenberg, Symbolse Physics;, ZooL, 

Mamm., dec. 1, folio g, Aug. 1832. Coastal mountains of Abyssinia, near 

Arkiko and Eilet. 
Euhyrax abyssinicus Gray, Ann. Mag. Nat. Hist. (4) 1: 47, Jan. 1868. 
Hyrax habyssinicus Anderson and De Winton, Zool. of Egypt, Mamm., p. 326, 

1902. 

Procavia habessinica alpini (Gray) 

Hyrax alpini Gray, Ann. Mag. Hist. Nat. (4) 1: 45, Jan. 1868. "Abyssinia." 

H[yrax] ferrugineus Gray, Ann. Mag. Nat. Hist. (4) 3: 242, Mar. 1869. Abys- 
sinia. 

Hlyrax] irroratus, var. luteogaster Gray, Ann. Mag. Nat. Hist. (4) 3: 242, Mar. 
1869. "Very probably Abyssinia." 

Hyrax irroratus, var. leucogaster Gray, Handlist Edentates, etc., Brit. Mus., 
p. 42, 1873 (lapsus calami for luteogaster). 

Procavia habessinica burtonii (Gray) 

Hyrax burtonii Gray, Ann. Mag. Nat. Hist. (4) 1: 43, Jan. 1868. "Egypt." 

Hyrax dongolanus Blanford, ex Hemprich and Ehrenberg, Proc. Zool. Soc. 

London, for 1869, p. 642, Apl. 1870. Dongola. 
Procavia ruficeps Thomas, Proc. Zool. Soc. London, p. 63, June 1892, not of 

Hemprich and Ehrenberg. 



450 bulletin: museum of comparative zoology 

Procavia habessinica butleri Wroughton 

Procavia butleri Wroughton, Ann. Mag. Nat. Hist. (8) 8: 461, Oct. 1911. Jebel 
Fazogli, Blue Nile at Abyssinian border, 3500 ft. 

Procavia habessinica d^mon Thomas 

Procavia dsemon Thomas, Ann. Mag. Nat. Hist. (8) 5: 199, Feb. 1910. Elgonyi, 

Mt. Elgon, Kenya Colony, 7000 ft. 
Procavia dsemon varians Granvik, Acta Univ. Lundensis, (2) 21: no. 3, p. 26, 

1925. East slopes of Mt. Elgon, Kenya Colony. 

Procavia habessinica erlangeri Neumann 

Procavia erlangeri Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 238, 1901. 

Headwaters of Webi Shebeli, Somaliland. 
Procavia erlangeri capillosa Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 302, 

10 Sep. 1917. Agada on the Abunass and Semano, Arusi Land, central 

Abyssinia. 
Procavia erlangeri comata Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 302, 

10 Sep. 1917. Gara Mulata, near Harar, Galla, Abyssinia. 

Procavia habessinica jacksoni Thomas 

Procavia jacksoni Thomas, Proc. Zool. Soc. London, p. 176, footnote, 1 June 

1900. Ravine Station, Kenya Colony. 
Procavia niackinderi zelotes Osgood, Publ. Field Mus. Nat. Hist., zool. ser., 10: 

no. 2, p. 5, 16 Feb. 1910. Between Naivasha and Kijabe, Kenya Colony. 

Procavia habessinica meneliki Neumann 

Procavia meneliki Neumann, Proc. Zool. Soc. London, 2: 143, Oct. 1902. Abuje 
and Badattino, Province of Gindeberat, south of Blue Nile, Abyssinia. 

Procavia habessinica minor Thomas 

P[rocavia\ abyssinica minor Thomas, Proc. Zool. Soc. London, p. 66, June 1892. 
Alali, at 13° N., Red Sea coast. 

Procavia habessinica pallida Thomas 

Procavia pallida Thomas, Ann. Mus. Civ. Stor. Nat., Genova, 30= (2) 10: 908, 
25 June 1891. Hekebo plateau, northern Somaliland. 

Procavia habessinica scioana (Giglioli) 

Hyrax scioanus Giglioli, Ann. Mus. Civ. Stor. Nat., Genova, 26= (2) 6: 21, 
5 Mar. 1888. Ankober, Shoa. 

Procavia shoana Thomas, Proc. Zool. Soc. London, p. 61, June 1892 (emenda- 
tion). 

Procavia johnstoni johnstoni Thomas 

Procavia johnstoni Thomas, Proc. Zool. Soc. London, p. 142, June 1894. Fort 

Lister, Nyasaland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 451 

Procavia johnstoni lopesi Thomas 

Procavia lopesi Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 251, 

June 1907. Kodja Hill, Gaima Range, Kibali River, Monbuttu, Congo 

Beige. 
Procavia ituriensis Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 303, 10 Sep. 

1917. Ituri, Congo Beige. 

Procavia johnstoni mackinderi Thomas 

Procavia mackinderi Thomas, Proc. Zool. Soc. London, p. 176, 1 June 1900. 
Teleki Valley, Mt. Kenya, Kenya Colony, 13,000 ft. 

Procavia johnstoni matschiei Neumann 

Procavia matschiei Neumann, Zool. Jahrb., Syst., 13: 555, 10 Oct. 1900. 
Mwanza, Tanganyika Territory. 

Procavia johnstoni volkmanni Brauer 

Pr[ocavia] volkmanni Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 35, Jan. 
1914. Franzfontein, northern Southwest Africa. 

Procavia ruficeps ruficeps (Hemprich and Ehrenberg) 

Hyrax ruficeps Hemprich and Ehrenberg, Symbolse Physicse, Zool., Mamm., 

dec. 1, folio h, pi. 2, upper fig., Aug. 1832. Dongola, Anglo- Egyptian 

Sudan. 
Hyrax dongolanus Blanford, Proc. Zool. Soc. London, for 1869, p. 642, Apl. 1870 

(in part). 

Procavia ruficeps bamend^e Brauer 

Procavia bamendse Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 125, Feb. 
1913. Bamenda, southwestern Cameroons. 

Procavia ruficeps bounhioli Kollmann 

Procavia bounhioli Kollmann, Bull. Mus. d'Hist. Nat., Paris, 18: 281, 1912. 

Ahaggar, Sahara Desert. Algeria. 
Procavia (Heterohyrax) antinese Heim de Balsac and Begouen, Bull. Mus. d'Hist. 

Nat., Paris, (2) 4:479, June 1932. Ahaggar, central Sahara, French West 

Africa. 

Procavia ruficeps buchanani Thomas and Hinton 

Procavia buchanani Thomas and Hinton, Novitates Zool., 28: 13, 14 May 1921. 
Aouderas, Asben, northern Nigeria. 

Procavia ruficeps ebneri Wettstein 

Procavia {Procavia) ebneri Wettstein, Anzeiger K. Akad. Wiss. Wien, math.- 
nat. Kl., 53: 162, 1916. Talodi, southern Kordofan, Anglo-Egyptian Sudan. 



452 bulletin: museum of comparative zoology 

Procavia ruficeps elberti Brauer 

Procavia elberti Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 300, 10 Sep. 

1917. Cave at Jade, about 7° N., 15° E., northern Cameroons. 
Procavia naumanni Brauer, Sitzb. Ges. Naturf. Freunde, Berlin, p. 300, 10 

Sep. 1917. On the Bole, tributary of the Uham, 6° N., 16° E., northern 

Cameroons. 

Procavia rtjficeps goslingi Thomas 

Procavia goslingi Thomas, Ann. Mag. Nat. Hist. (7) 15: 82, Jan. 1905. Ner- 
tang, Yerghum country, northern Nigeria. 

Procavia rtjficeps kamerunensis Brauer 

Procavia kamerunensis Brauer, Sitzb. Ges. Naturf. Freunde, p. 127, Feb. 1913. 
Lagdo Hills, southeast of Garua, northern Cameroons. 

Procavia ruficeps kerstinqi Matschie 

Procavia kerstingi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 59, 1899. 

Tshyati, 50 km. southeast of Pessi, about 7° 50' N., near Togo-Dahomy 

boundary, Nigeria. 

Procavia ruficeps latastei Thomas 

Procavia latastei Thomas, Proc. Zool. Soc. London, p. 69, June 1892. Senegal, 
cotypes from Fclou and Medine. 

Procavia ruficeps marrensis Thomas and Hinton 

Procavia ruficeps marrensis Thomas and Hinton, Proc. Zool. Soc. London, p. 

271, 6 July 1923. Central part of Jebel Marra, Darfur, Anglo-Egyptian 

Sudan, 7100 ft. 

Procavia ruficeps melfica Mertens 

Procavia sharica melfica Mertens, Senckenbergiana, Frankfurt a. M., 11: 121, 

29 Apl. 1929. Melfi Mts., East Bagirmi, Chad district, French Equatorial 

Africa. (Omitted from Hahn's monograph, 1934.) 

Procavia ruficeps oweni Thomas 

Procavia oweni Thomas, Ann. Mag. Nat. Hist. (8) 8: 123, July 1911. Near 
Daley, Fouta Djalon, French Gambia. 

Procavia ruficeps sharica Thomas and Wroughton 

Procavia sharica Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 522, 
June 1907. Kajibu, Shari River, French Equatorial Africa. 

Procavia ruficeps slatini Sassi 

Procavia slatini Sassi, Sitzb. K. K. Akad. Wiss., Wien, math.-nat. Kl., 115: 

sect. 1, pt. 6, p. 1002, 2 pis., June 1906. Hills on the White Nile, south of 

Gondokoro, 5° N., Anglo-Egyptian Sudan. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 453 

Order PROBOSCIDEA. Proboscideans 
Family ELEPHANTID.E. Elephants 

LoxoDONTA F. Cuvier. African Elephants 

Loxodonta F. Cuvier, Zool. Journ., 3: 140, Jan. 1827, based on the Loxodonte 
of E. Geoffroy and F. Cuvier, Hist. Nat. des Mammiferes, pts. 51, 52, Nov. 
1825. Genotype Elephas capensis F. Cuvier =^. africanus Blumenbach. 

Loxodon Falconer, Quart. Journ. Geol. Soc. London, 13: 315 and synopt. tab., 
1 Nov. 1857 (preoccupied). "Includes species allied to the African Ele- 
phant" hence genotype is Elephas africanus Blumenbach. 

Loxodonta africana africana (Blumenbach). South African Bush Ele- 
phant. 
Elephas africanus Blumenbach, Handbuch d. Naturgesch., ed. 5, p. 125, Atlas, 
pi. 19, f. C, 1797. No locality given, but selected as Orange River, South 
Africa (Pohle, H., Zeitschr. f. Saugethierk., 1 : 63, 31 Aug. 1926). 
Hollister (Bull. U. S. Nat. Mus., no. 99, pt. 3, p. 138, 1924) regards the state- 
ment of Roosevelt and Heller (Life Histories of African Game Animals, 2: 716, 
1914) that "we have applied the older name, africana, to the form inhabiting 
the Congo basin," as fixing the type locality; this, however, is too general a 
statement, and may be interpreted as tentative only. 
Elephas capensis F. Cuvier, Tableau Elcmentaire de I'Hist. Nat. des Anim., p. 

149, 1798. Orange River region, South Africa. 
Elephas africanus toxotis Lydekker, Proc. Zool. Soc. London, p. 385, 388, text-f. 
106, 107, 1 Aug. 1907. Western Cape of Good Hope; type mounted in 
South African Museum, Cape Town; type locality later corrected to Addo 
Bush, southeastern Cape of Good Hope. 

Loxodonta africana knochenhaueri (Matschie) 

Elephas africanus knochenhaueri Matschie, Sitzb. Ges. Naturf . Freunde, Berlin, 
p. 197, 1900. Barikiwa, southern Tanganyika Territory. 

Loxodonta africana oxyotis (Matschie) 

Elephas {Loxodonta) oxyotis Matsqhie, Sitzb. Ges. Naturf. Freunde, Berlin, 
p. 196, 1900. Upper Atbara River, Anglo-Egyptian Sudan. 

The status of the following supposed races of Bush Elephant is still uncertain, 
though probably few of them will prove to be recognizable: — 
Loxodonta africana angolensis Frade, Titulos e Trabalhos Cientificos, (Curricu- 
lum vitse), Lisboa, p. 15, June 1928; Bull. Soc. Portuguaise des Sci. Nat.,, 
Lisbonne, 11: 322, Dec. 1933. Cunene region, Angola. 



454 bulletin: museum of comparative zoology 

Elephas africanus cavendishi Lydekker, Proc. Zool. Soc. London, p. 394 
(legend to text-f. 115, of type), 395 (characters), 1 Aug. 1907. Lake Ru- 
dolf district, northern Kenya Colony. 

Elephas africanus mogambicus Frade, Bull. Soc. Portuguaise des Sci. Nat., Lis- 
bonne, 9: 133, text-f. 4, 1924. Maputo region, Mozambique. 

Elephas africanus orleansi Lydekker, Proc. Zool. Soc. London, p. 398 (legend to 
text-f. 118, of type, and in text), 1 Aug. 1907. Northern Somaliland. 

Elephas africanus peeli Lydekker, Proc. Zool. Soc. London, p. 393 (legend to 
text-f. 114 of cotype), 394, 1 Aug. 1907. Aberdare Mts., Kenya Colony 
(first coty]3e); southeast Africa (second cotype). 

Elephas africanus rothschildi Lydekker, Proc. Zool. Soc. London, p. 399, text-f. 
119 (of "Jumbo"), 1 Aug. 1907. "Some part of the French Sudan, prob- 
ably to the southward of Lake Chad." 

Elephas africanus selousi Lydekker, Proc. Zool. Soc. London, p. 387 (in legend 
to text-f. 108, of type), 388, text-f. 109, 389 (type designated), 1 Aug. 1907. 
Mashonaland, Rhodesia. 

Loxodonta africana zukowskyi Strand, Arch. f. Naturgesch., 90: sect. A, pt. 1, 
p. 68, footnote, July 1924. Kaokoveld, Southwest Africa. 

Loxodonta cyclotis (Matschie). Forest Elephant 

E[lephas] cyclotis or Elephas capensis cyclotis Matschie, Sitzb. Ges. Naturf. 

Freunde, Berlin, p. 194, 1900. Yaunde, southern Cameroons. 
Elephas africanus pumilio Noack, Zool. Anzeiger, 29: 631, 8 Jan. 1906. French 

Congo. 
Elephas africanus alhertensis Lydekker, The Field (London), 107: 1089, 30 

June 1906; Proc. Zool. Soc. London, p. 402, text-f. 121 (skull), 1 Aug. 

1907. Southwest end of Albert Nyanza, Congo Beige. 
Elephas africanus cottoni Lydekker, Proc. Zool. Soc. London, for 1907, p. 783, 

4 June 1908; based on ear figured and described in Proc. Zool. Soc. London, 

p. 390, text-f. Ill, 1 Aug. 1907. Congo. 
Elephas africanus fransseni Schouteden, Revue Zool. Africaine, 3: 396, 20 Jan. 

1914. M'Paa, near Lake Leopold II, Congo Beige. 

For summary of history and characters, see Allen, G. M., Proc. Acad. Nat- 
Sci., Philadelphia, 88: 15-44, pi. 1-4, text-f. 1-2, 10 Mar. 1936. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 455 

Order SIRENIA. Sea-cows 
Family DUGONGID.E. Dugongs 

DuGONG Lacepede 

Dugong Lacepede, Tabl. des Mamm., p. 17, 1799; Buffon, Hist. Nat., Quad- 

rupedes, Didot ed., 14: 193, 1799. Genotype Dugong indicus Desmarest. 
Platystomus G. Fischer, Das National Museum d. Naturgesch. zu Paris, 2: 353, 

1803; Zoognosia, ed. 3, 1: 15, 19, 1813. Genotype Platystomus dugong 

(Gmelin) = Trichecus dugon Miiller. 
Dugimgus Tiedemann, Zoologie, 1 : 554, 1808, emendation. 
Halicore Illiger, Prodromus Syst. Mamm. et Avium, p. 140, 1811. Genotype 

Trichechus dugong Gmelin = Trichecus dugon Miiller. 
Dugongidus Gray, London Medical Repos., 15: 309, 1 Apl. 1821. Genotype 

Trichecus dugon Miiller. 

Dugong dugon (P. L. S. Miiller). Dugong. 

Trichecus (sic) dugon P. L. S. Miiller, Linne's VoUstandigen Natursystems, 

Suppl., p. 21, 1776. "Vorgeburge der Guten Hofnung an, bis an die philippi- 

nischen Inseln." 
H[alicore\ hemprichii Ehrenberg, in Hemprich and Ehrenberg, Symbolse Physi- 

cse, Mamm., dec. 2, folio k, footnote, Sep. 1832. Barkan Island, Red Sea. 
H[alicore] lottum Hemprich and Ehrenberg, Symbolae Physicse, Mamm., dec. 

2, folio k, footnote, Sep. 1832. Hauakel Island, southern part of Red Sea. 
Halicore tahernaculi Riippell, Mus. Senckenburgianum, 1: 113, pi. 6, 1834. Red 

Sea, based on a skeleton found on Maxud Island. 
Halicore cetacea Heuglin, Reise in Nordost-Afrika, 2: 135, 1877. Red Sea. The 

name is attributed by Heuglin to Illiger. 

Family TRICHECHID.^. Manatees 

Review: Hatt, Robert T. The American Museum Congo Expedition mana- 
tee and other recent manatees. Bull. Amer. Mus. Nat. Hist., 66: 533-566, pi. 27 
(exterior), 10 Sep. 1934. 

Trichechus Linnaeus 

Trichechus Linnaeus, Syst. Nat., ed. 10, 1 : 34, 1758. Genotype Trichechus rnana- 
tus Linnseus (see Opinion 112, Internat. Comm. on Zool. Nomenclature, 
Smithsonian Misc. Coll., 73: no. 6, p. 19, 8 June 1929). 

Manatus Briinnich, Zoologise Fundamenta, pp. 34, 38, 1772. Genotype Triche- 
chus manatus Linnaeus. 



456 bulletin: museum of comparative zoology 

Oxystomus G. Fischer, Das National Museum d. Naturgesch. zu Paris, 2: 353, 

1803. Genotype Trichechus manatus Linnaeus. 
Halipsedisca Gistel, Naturgesch. d. Thierreichs f. Hohere Schulen, p. 83, 1848. 

Genotype Manatus americanus Desmarest. 

Trichechus seneqalensis Link. West African Manatee. 

Trichechus manatus a australis Gmelin, Linnseus's Syst. Nat., ed. 13, 1: pt. 1, 

p. 60, 1788 (in part). "Mari africano et americano." When, in 1800, Shaw 

restricted the name to the African manatee, the latter had already been 

named by Link. 
Trichechus senegalensis Link, Beytrage z. Naturgesch., 1: sec. 2, p. 109, 1795. 

Senegal; based mainly on Buffon's account (Hist. Nat., 13: 431, 1765; 

suppl., 6: 403, 1782) after Adanson. 
Trichechus sequatorialis Lac^pede, Tableaux d. Mammiferes, p. 17, 1799 (part), 

a nomen nudum. 
Manatus stroggylonurus Bechstein, in Pennant's AUgemeine Uebersicht d. 

Vierfiissigcn Thiere, 2: 732, 1800. 
Manatus sphserurus Illiger, Abhandl. K. Akad. Wiss., Berlin, for 1804-11, p. 79, 

87, 1815. 
T[richechus] M\anatus] africanus Oken, Lehrb. d. Naturgesch., 3: sect. 2, p. 688, 

1816. Senegal. 
Manatus atlanticus Oken, Allgemein. Naturgesch. f. alle Stande, 7: sect. 2, 

p. 1098, 1838, in part. Atlantic coast of South America, especially Antilles, 

Cayenne, Surinam. 
Manatus nasutus Wyman, Proc. Boston Soc. Nat. Hist., 2: 199, footnote. Mar. 

1847. Caracalla River, 20 miles east of Cape Palmas, Ivory Coast. 
Manatus vogelii Owen, Edinburgh New Philos. Journ., (2) 4: 346, Oct. 1856. 

Benue River, northern Nigeria. 
Manatus oweni Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 367, Nov. 1860. 

Camma country, various localities named, about the mouth of the Gaboon 

River. 
Range: West coast of Africa from Senegal to the Quanza River, Angola, 

following up the larger rivers where possible. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 457 

Order ARTIODACTYLA. Even-toed Ungulates 

Section Suina. Pigs and Hippos 

Family HIPPOPOTAMIDyE. Hippopotami 

Chceropsis Leidy. Pygmy Hippopotamus 

Choerodes Leidy, Proc. Acad. Nat. Sci. Philadelphia, 6: 52, 1852, not of White, 

1846. Genotype Hippopotamus liberiensis Morton. 
Di'protodon Duvernoy, Compt. Rend. Acad. Sci. Paris, 29: 277, 1849, not of 

Owen, 1838. A subgenus of Hippopotamus, type Hippopotamus liberiensis 

Morton. 
Chceropsis Leidy, Journ. Acad. Nat. Sci. Philadelphia, (2) 2: 213, pi. 21, Jan. 

1853. Genotype Hippopotamus liberiensis Morton. 

Chceropsis liberiensis (Morton) 

Hippopotamus minor Morton, Proc. Acad. Nat. Sci. Philadelphia, 2: 14, 1844, 
not of Desmarest, 1822. St. Paul's River, Liberia. 

Hippopotamus (Tetraprotodon) liberiensis Morton, Journ. Acad. Nat. Sci. Phila- 
delphia, (2) 1: 232, pi. 32-34, Aug. 1849. Substitute for Hippopotamus 
minor Morton, not Desmarest, 1822. 
Range: Liberia and adjacent parts of Sierra Leone. 

Hippopotamus Linnaeus 

Hippopotamus Linnaeus, Syst. Nat.,ed. 10, 1: 74, 1758. Genotype Hippopota- 
mus amphibius Linnaeus. 

Hippopotamus amphibius amphibius Linnaeus. Northern Hippo 
Hippopotamus amphibius Linnaeus, Syst. Nat., ed. 10, 1: 74, 1758. Nile River, 

Egypt. 
Hippopotamus senegalensis Desmoulins, Dictionnaire Class. d'Hist. Nat., 8: 

220, 1826. Senegal. See Schwarz, E., Ann. Mag. Nat. Hist. (10) 14: 261, 

Aug. 1934. 
Hippopotamus abyssinicus Lesson, Nouveau Tabl. Regne Anim., Mamm., p. 

158, 1842, nomen nudum. Abyssinia. 
Hippopotamus typus Duvernoy, Compt. Rend. Acad. Sci., Paris, 23: 650, 1846. 

Senegal to Abyssinia. 

Hippopotamus amphibius capensis Desmoulins. Cape Hippo 
Hippopotamus capensis Desmoulins, Dictionnaire Class. d'Hist. Nat., 8: 222, 

1825. Lower Berg River, western Cape of Good Hope. See Schwarz, E., 

Ann. Mag. Nat. Hist. (10) 14: 261, Aug. 1934. 



458 bulletin: museum of comparative zoology 

Hippopotamus australis Duvernoy, Compt. Rend. Acad. Sci. Paris, 23: 650, 
1846. Cape of Good Hope. 

Hippopotamus amphibius constrictus Miller. Angola Hippo 
Hippopotamus constrictus Miller, Smithsonian Misc. Coll., 54: no. 7, p. 1, pi. 

1-4, lower fig., 28 Mar. 1910. Angola. 
Hippopotamus constrictor Zukowsky, Arch. f. Naturgesch., 90: sect. A, pt. 1, 

p. 99, July 1924; lapsus for H. constrictus. 

Hippopotamus amphibius kiboko Heller 

Hippopotamus amphibius kiboko Heller, Smithsonian Misc. Coll., 61: no. 22, 
p. 1, 26 Jan. 1914. Lake Naivasha, Kenya Colony. 

Hippopotamus amphibius tschadensis Schwarz 

Hippopotamus amphibius tschadensis Schwarz, Ann. Mag. Nat. Hist. (8) 13: 
31, Jan. 1914. Katana, Bornu, northern Nigeria. 



Family SUID^. Pigs 

Subfamily SUIN.E 

Hylochcerus Thomas. Giant Forest Pigs 

Hylochoerus Thomas, Nature (London), 70: 577, 13 Oct. 1904; Proc. Zool. Soc. 
London, 2: 193 pis. 14, 15, 18 Apl. 1905. Genotype Hylochoerus meinertz- 
hageni Thomas. 

Hylochcerus meinertzhageni meinertzhageni Thomas 

Hylochoerus meinertzhageni Thomas, Nature (London), 70: 577, 13 Oct. 1904; 
Proc. Zool. Soc. London, 2: 193, pis. 14, 15, 18 Apl. 1905. Kakumega 
Forest, near Kaimosi, Kenya Colony (see Loveridge, A., in Allen, G. M., 
and Lawrence, B., Bull. Mus. Comp. Zool., 79: 109, Jan. 1936). 

Hylochoerus iiuriensis Matschie, Ann. Mus. du Congo, Zool., (5 =2) 1 =2: pt. 1, 
p. 9, pi. 1-5, Oct. 1906. Upper Ituri Valley, eastern Congo Beige. 

Hylochoerus gigliolii Balducci, Publ. R. 1st. Stud. Super., Firenze, p. 1-15, 2 pis., 
1909. Upper Congo. 

Hylochcerus meinertzhageni ivoriensis Bouet and Neuville 
Hylochoerus meinhertzhageni (sic) ivoriensis Bouet and Neuville, Revue Hist. 
Nat., Soc. Nat. d'Acclimat. France, 11: no. 9, p. 284, Sep. 1930. Bolobo, 
Liberia, and Georgetown, Ivory Coast, both on lower Cavalla River. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 459 

Hylochcerus meinertzhageni rimator Thomas 

Hylochaerus rimator Thomas, Abstr. Proc. Zool. Soc. London, no. 25, p. 1, 23 

Jan. 1906; Proc. Zool. Soc. London, 1: 2, text-f. 1, 7 June 1906. Ja River, 

Cameroons. 

Hylochcerus meinertzhageni schulzi Zukowsky 

Hylochaerus schulzi Zukowsky, Arch. f. Naturgesch., 87: sect. A, pt. 1, p. 181 

(text-f.), 189, Dec. 1921. Mutjek and Meru Mts., west of Kilimanjaro, 

Tanganyika Territory, 2000 m. 



Phacochcerus F. Cuvier. Wart Hogs 

Aper Pallas, Miscellanea Zool., p. 16, pi. 2; pi. 4, f. 1, 2, 4, 1766. Genotype 
Aper sethiopicus Pallas. 
Although Aper may be regarded as merely the masculine form of Sus, it is 

perhaps here used as a distinct generic term and as such would supersede 

Phacochcerus. 

Phaco-chcerus G. Cuvier, Regne Animal, ed. 1, 1: 236, footnote, 1817. Geno- 
tjrpe Sus sethiopicus Gmelin =Aper sethiopicus Pallas. 

Eureodon G. Fischer, Mem. Soc. Imp. des Nat. Moscou, 5: 373, 1817. Geno- 
type Sus sethiopicus Gmelin. 
This name should be used in preference to Phacochcerus according to M. W. 
Lyon, Jr. (Proc. Biol. Soc. Washington, 28: 141, 29 June 1915). 

Phascochoeres Cretzschmar, in Riippell's Atlas zu d. Reise im Nordlich. 
Afrika, Saugeth., p. 61, 1826, variant. 

Dinochcerus Gloger, Hand- u. Hilfsbuch d. Naturgesch., 1: xxxii, 131, 1841. 
Genotype Aper sethiopicus Pallas. 

Macrocephalus Elliot, Publ. Field Mus. Nat. Hist., zool., 8: 32, 1907, ex Frisch, 
Das Natur-System Vierfiiss. Thiere, in Tabellen, p. 3, 1775 (Frisch's 
names are not available, since he is not consistently binomial). Genotype 
Aper sethiopicus Pallas. 

For review of characters, see Lonnberg, E., Remarks on some wart-hog 
skulls in the British Museum. Proc. Zool. Soc. London, for 1908, p. 936-940, 
8 Apl. 1909. 

In addition to the living forms described, a number of subfossil fragments 
probably of no great age, have lately been named from South Africa. 

Phacochcerus ^thiopicus ^thiopicus (Pallas). Cape Wart Hog 

Aper sethiopicus Pallas, Spicilegia Zool., pt. 2, p. 2, pi. 1, 1767. South Africa, 

Cape of Good Hope. 
Phacochcerus edentatus I. Geoffroy, Dictionnaire Classique d'Hist. Nat., 13: 

320, 1828. Cape of Good Hope. 



460 bulletin: museum of comparative zoology 

Phascochcerus (sic) typicus A. Smith, South African Quart. Journ., 2: 178, 

1834. Interior of Cape Colony. 
Phacochcerus pallasi Van der Hoeven, Nova Acta Acad. Caes. Leop.-Carol., 

Breslau & Bonn, 19: 173, 1839. Cape of Good Hope. 

Phacochcerus -ethiopicus .eliani (Cretzschmar) 

Phascochoeres seliani Cretzschmar, in Riippell's Atlas zu d. Reise im Nordlich. 

Afrika, Siiugeth., p. 61, pi. 25, 26, 1826. Abyssinia, eastern slopes, and 

Kordofan. 
Phacochcerus haroia Hemprich and Ehrenberg, Symbols Physical, Mamm., 

dec. 2, folio oo, pi. 20, Nov. 1832. Arkiko, Eritrea. 
P[hacochcerus] sclateri Gray, Ann. Mag. Nat. Hist. (4) 6: 189, 263, Aug. 1870. 

Zoulla, Red Sea Coast. (Based on Sclater's account in Proc. Zool. Soc. 

London, p. 276, pi. 20 (col.), text-f., Sep. 1869.) 
Phacochcerus massaicus Lonnberg, Wiss. Ergebn. d. Schwedischen Zool. 

Exped. nach Kilimandjaro u. Leitung v. Sjostedt, 2: Mamm., p. 54-55, 

1908. Tanganyika Territory (olim "German East Africa"); probably 

Kilimanjaro-Meru region. 
Made a synonym of P. aethiopicus seliani by Hollister (Bull. U. S. Nat. 
Mus., no. 99, pt. 3, p. 51, 1924). 

Phacochcerus ^thiopicus africanus (Gmelin). Western Wart Hog 

Sus africanus Gmelin, Linnseus's Syst. Nat., ed. 13, 1: pt. 1, p. 220, 1788. 

Cape Verde, Senegambia, (to Cape of Good Hope). 
Phacochcerus incisivus I. Geoffroy, Dictionnaire Class. d'Hist. Nat., 13: 321, 

1828. Cape Verde. 

Phacochcerus .ethiopicus barkeri Rothschild 

Phacocha-rus barkeri Rothschild, Ann. Mag. Nat. Hist. (9)6: 416, Oct. 1920. 
Southwest of Bahr-el-Ghazal, Anglo-Egyptian Sudan. 

Phacochcerus ^thiopicus bufo Heller 

Phacochcerus africanus bufo Heller, Smithsonian Misc. Coll., 61 : no. 22, p. 2, 
26 Jan. 1914. Rhino Camp, Lado Enclave. 

Phacochcerus ^thiopicus centralis Lonnberg 

Phacochcerus africanus centralis Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 58: no. 2, p. 102, 1 Sep. 1917. Lake Albert Edward, 

Congo Beige. 

Phacochcerus ^thiopicus delamerei Lonnberg 

Phacochcerus delamerei Lonnberg, Proc. Zool. Soc. London, for 1908, p. 940, 

8 Apl. 1909. Northern Somaliland (see Lydekker, R., Cat. Ungulate 

Mamm. Brit. Mus., 4: 371, 1915). 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 461 

Phacochcerus ^thiopicus fossor Schwarz 

Phacochcerus sethiopicus fossor Schwarz, Ann. Mag. Nat. Hist. (8) 11: 265, 

Feb. 1913. Ketekma, east of Tschekna, Bagirmi, French Equatorial 

Africa. 

Phacochoerus .ethiopicus shortridgei St. Leger 

Phacochcerus sethiopicus shortridgei St. Leger, Ann. Mag. Nat. Hist. (10) 
10: 86, July 1932. Numkaub, Grootfontein district, Southwest Africa. 

Phacochcerus ^thiopicus sundevallii Lonnberg 

PhacochcErus sundevallii Lonnberg, Wiss. Ergebn. d. Schwedischen Zool. 

Exped. nach Kilimandjaro u. Leitung v. Sjostedt, 2: Mamm., p. 54, 

1908. Natal. 



PoTAMOCHCERUS Gray. Red River Hogs or Bush Pigs 

Potamochcerus Gray, Proc. Zool. Soc. London, for 1852, p. 129, pi. 34, 27 June 
1854. Genotype Choiropotamus pictus Gray. 
For use of Potamochcerus, see Schwarz, E., Der Gattungsname der afrikanis- 
chen Flussschweine, Zeitschr. f. Siiugetierkunde, 8: 284, 20 Dec. 1933. 

Choiropotamus Gray, List. Mamm. Brit. Mus., p. 185, 13 May 1843. Genotype 
Sus africanus Schreber = Sus koiropotamus Desmoulins. Name pre- 
occupied by Chceropotamus Desmarest, 1822, for a genus of extinct ungu- 
lates from France. 

Koiropotamus Gray, List Spec. Mamm. Brit. Mus., p. xxvii, 13 May 1843. 
Based on S^is africanus Schreber, not Gmelin, =<Sms koiropotamus Des- 
mouhns. 
According to Palmer, No. Amer. Fauna, no. 23, p. 358, 1904, this is a nomen 

nudum, but Holhster, Proc. Biol. Soc. Washington, 34: 77, 31 Mar. 1921, 

regards it as a valid name for the genus. Since both Choiropotamus and 

Koiropotamus are from the same words as Chceropotamus, it seems best to 

accept the better-known Potamochcerus. 

Nyctochaerus Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Dresden, 30: 
suppl. to sect. 2, p. 7, 1863. Genotype Nyctochoerus hassama Heuglin. 

See Lonnberg, E. Contributions to the knowledge of the genus Potamo- 
chcerus. Arkiv f. Zool., 7: no. 6, p. 1-40, 5 text-f., 28 Nov. 1910. 

de Beaux, Oscar. Beitrag zur Kenntnis der Gattung Potamochcerus Gray 
{ = Choiropotamus Graj', 1843. Thomas), in Proc. Biol. Soc. Washington, 
vol. 34, 1921, p. 135); Zool. Jahrb., Syst., 47: 379-503, pi. 4-6, 3 text-f., 25 
June 1924. 

Potamochcerus larvatus larvatus (F. Cuvier) 

Sus larvatus F. Cuvier, Mem. Mus. d'Hist. Nat. Paris, 8: 447, pi. 22, 1822. 
Madagascar. 



462 bulletin: museum of comparative zo6lo(JY 

Phacochcerus koiropotamus Lesson, Nouv. Tabl. Regne Anim., Mamm., p. 162,. 

1842. Madagascar. * 

Sus madagascariensis A. Grandidier, Revue et Mag. de Zool. (2), 19: 85, Mar. 

1867. West coast of Madagascar. 
Potamochcrrus edwardsi A. Grandidier, Revue et Mag. de Zool. (2), 19: 318, 

Sep. 1867. Morondava, southwest coast of Madagascar. 
Choir opotamus africanus Gray, List Spec. Mamm. Brit. Mus., p. 185, 13 May 

1843, in part. 

PoTAMOCHCERUS labvAtus hova Lonnberg 

Poiamochoerus larvalus hova Lonnberg, Arkiv f. Zool., 7: no. 6, p. 32, 28 Nov. 
1910. Ampitambe, eastern Madagascar. 

PoTAMOCHCERUS PORCtJS PORCus (Linnajus) 

Siis porcus Linnaeus, Syst. Nat., ed. 10, 1 : 50, 1758. Guinea, West Africa,. 

based on Marcgrave's Porcus guineensis. 
Sus penicillatus Schinz, Monogr. Saugeth., pt. 10, p. 12, 1848. 

PoTAMOCHCERUS PORCUS ALBIFRONS Du ChaiUu 

Potamochcerus albifrons Du Chaillu, Proc. Boston Soc. Nat. Hist., 7: 301, Jul}' 
1860. Between Cape Lopez and Cape St. Catherine, and headwaters of 
Fernand-Vaz River. 

Potamochcerus porcus albifrons Lonnberg, Arkiv f. Zool., 7: no. 6, p. 11, 28 Nov. 
1910. 

PoTAMOCHCERUS PORCUS ALBiNUCHALis Lonnberg 

Poiamochoerus porcus albinuchalis Lonnberg, Revue Zool. Africaine, 7: 245, 30 
Dec. 1919. Lake Leopold II, Congo Beige. 

PoTAMOCHCERUS PORCUS ARRHENii Lonnberg 

Potamochcerus intermedius arrhenii Lonnberg, Kungl. Svenska Vet.-Akad. 

Handl., Stockholm, (2) 58: no. 2, p. 104, text-f. 10-11, 1 Sep. 1917. Rut- 

shuru, eastern Congo Beige. 

PoTAMOCHCERUS PORCUS coNGicus Lonnberg 

Potamochcerus porcus congicus Lonnberg, Arkiv f. Zool., 7: no. 6, p. 14, text-f. 
2, 28 Nov. 1910. Mukimbungu, Lower Congo, Congo Beige. 

PoTAMOCHCERUS PORCUS coTTONi Pinfold 

Potamochcerus charopotamus cottoni Pinfold, Ann. Mag. Nat. Hist. (10) 2: 99, 
July 1928. Tunda, Quanza district, Angola. 

PoTAMOCHCERUS PORCUS D^MONis Major 

Potamochcerus chceropotamus dsemonis Major, Proc. Zool. Soc. London, p. 367^ 
pi. 25, f . 1 ; pi. 26, f. 3, 1 Aug. 1897. Kilimanjaro, Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 463 

POTAMOCHCERUS PORCUS HASSAMA (Heuglill) 

Nyctochcerus hassama Heuglin, Nova Acta Acad. Css. Leop.-Carol., Dresden, 
30: no. 2, suppl., p. 7, 1863. Abyssinia. 

PoTAMOCHCERUs PORCUS iNTERMEDius Lonnberg 

Potamochwrus intcrmedius Lonnberg, Arkiv f. Zool., 7: no. 6, p. 38, 28 Nov. 
1910. Victoria Nile, Uganda, 4000 ft. 
The presence of this intermediate race breaks down Lonnberg's theory of a 
western and an eastern species of bush pig, so that all seem best regarded as 
races of P. porcus Linnaeus. 

PoTAMOCHCERUs PORCUS JOHNSTONi Major 

Potamochcerus johnstoni Major, Proc. Zool. Soc. London, p. 367, pi. 25, f. 3; 
pi. 26, f. 1, 1 Aug. 1897. Ngarawi Valley, northwestern Nyasaland. 

POTAMOCHCERUS PORCUS KENi^ Lonnberg 

Potamochoerus choeropotamus keniae Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 66 
Jan. 1912. Forests near Nairobi, Kenya Colony. 

POTAMOCHCERUS PORCUS KoiROPOTAMUS (Desmoulins) 

Sus koiropotamus Desmoulins, Dictionnaire Classique d'Hist. Nat., 17: 139, pL 

10, f. 2, 1831. Probably South Africa. 
Sus africanus Gmelin, Linnaeus's Syst. Nat., ed. 13, 1 : pt. 1, p. 220, 1788, in part. 

Cape Verde to Cape of Good Hope. 
Sus capensis Gerrard, Cat. Bones of Mamm. Brit. Mus., p. 277, 1862, a nomen 

nudum. South Africa. 
PotamochcErus ch(xropotamus Major, Proc. Zool. Soc. London, p. 366, 1 Aug. 

1897. Lydekker, Cat. Ungulate Mamm. Brit. Mus., 4: 350, 1915. 

POTAMOCHCERUS PORCUS MASCHONA Lonuberg 

Potamochcerus choeropotamus maschona Lonnberg, Arkiv f. Zool., 7: no. 6, p. 20, 
28 Nov. 1910. Mashonaland, northeastern Rhodesia. 

POTAMOCHCERUS PORCUS MAWAMBICUS LorcnZ 

Potamochoerus porcus mawambicus Lorenz, Ann. Nat. Hist. Mus. Wien, 36: 
90, text-f. 1, Aug. 1923. Mawambi, Ituri district, eastern Congo Beige. 

POTAMOCHCERUS PORCUS nyasjE Major 

Potamochcerus choeropotamus nyasse Major, Proc. Zool. Soc. London, p. 367, pi. 
25, f. 3; pi. 26, f. 4, 1 Aug. 1897. Zomba, Nyasaland. 

Potamochoerus porcus pictus (Gray) 

Choiropotamus pictus Gray, Ann. Mag. Nat. Hist. (2) 10: 281, Oct. 1852. Cam- 
eroon River, Cameroons. 

Potamochcerus porcus somaliensis de Beaux 

P\otamochcErus\ larvatus somaliensis de Beaux, Zool. Jahrb., Syst., 47: 490, pi. 4, 
f. 3; pi. 6, f. 16, 25 June 1924. Bulo Burti, upper Webi Shebeli, Somaliland. 



464 bulletin: museum of comparative zoology 

PoTAMOCHCERus PORCus UBANGENSis Lonnberg 

PotamochoErus porcus ubangeni^is Lonnbcrg, Arkiv f. Zool., 7: no. 6, p. 10, 28 
Nov. 1910. Ubangi, northern Congo Beige. 

Sus Linnaeus. Common Pig; Wild Boar 

Sus Linnseus, Syst. Nat., ed. 10, 1: 49, 1758. Genotype Sus scrofa Linnseus. 
Cenluriosus Gray, Proc. Zool. Soc. London, p. 17, June 1862. As a subgenus of 

Sus, type Sus pliciceps Gray. 
Gyrosus Gerrard, ex Gray MS., Cat. Bones Mamm. Brit. Mus., p. 278, 10 

Mar. 1862; Gray, Cat. Carniv., Pachyderm., and Edentate Mamm. 

Brit. Mus., p. 347, 1869. As a subgenus of Sus, type Sus pliciceps Gray. 
Scrofa Gray, Proc. Zool. Soc. London, p. 38, May 1868. Genotype Sus domesti- 

cus Brisson=*S us scrofa Linnseus. 
Euhys Gray, Cat. Carniv., Pachyderm., and Edentate Mamm. Brit. Mus., p. 

339, 1869. As a subgenus of Sus, type Sus barbatus S. Miiller. 
Aulacochoerus Gray, Ann. Mag. Nat. Hist. (4) 11: 435, June 1873. Genotype 

Sus vittatus S. Mliller. 
Dasychoerus Gray, Ann. Mag. Nat. Hist. (4) 11 : 435, June 1873. Genotype not 

designated; included Sus verrucosus Mliller and S. celebensis Miiller. 
Sinisus Heude, Mem. concern. Hist. Nat. de I'Empire Chinois, 2: pt. 2, p. 102, 

1892. As a subgenus of Sus, type not designated, but term applied to the 

Chinese forms of Sus scrofa. 

Sus SCROFA ALGiRA Loche. Barbary Pig 

Sus scrofa var. barbarus P. L. Sclater, Proc. Zool. Soc. London, p. 443, 1860. 

"Ex Afr. bor.," =? Morocco. This is a nomen nudum. 
Sus scrofa var. algira Loche, Exploration Sci. de I'Algerie, Zool., Mammif., p. 

59, 1867. Algeria, country of Beni Sliman. 
Sus gambianus Gray, Cat. Carniv., Pachyderm, and Edentate Mamm. Brit. 

Mus., p. 346, 1869. Gambia; a nomen nudum applied to the domestic pig. 

Sus SCROFA SAHARiENSis Heim de Balsac 

Sus scrofa sahariensis Heim de Balsac, Suppl. au Bull. Biol, de France et de 

Belgique, Paris, 21: 330 (brief description), 47, 95, 237 (mention), 1936; 

Bull. Soc. Zool. de France, 62: 333, 14 Aug. 1937 (formal description). 

Jebel Guettar, northwest of Ain Sefra, Algeria. 

Sus SCROFA SENNAARENSis Gray 

Sus sennaarensis Fitzinger, Sitzb. K. Akad. Wiss., Wien, math.-nat. Kl., 29: 
365, 1858 (nomen nudum). Gray, Proc. Zool. Soc. London, p. 32, May 
1868 (description). "Sennaar; Kordofan; Sudan." 
Range: Formerly from Egypt to Sennaar and Kordofan; exterminated in the 
Nile Delta in the early '90's; persisted in Wadi Natron till about 1902, but re- 
stocked there with pigs from Hungary or Austrian Poland (see Flower, S. S., 
Proc. Zool. Soc. London, p. 442-443, 8 July 1932). 



ALLEN: CHECKLLST OF AFRICAN MAMMALS 465 

Section Tylopoda 
Family CAMELID.E 

Camelus Linnaeus. Camels 

Camelus Lixinseus, Syst. Nat., ed. 10, 1 : 65, 1758. Genotype Camelus badrianus 
Linnaeus (see Opinions of Intern. Comm. on Nomenclature, p. 37, 1910; 
notwithstanding that Palmer in 1904 designated C. dromedarius as type). 

Camelus dromedarius Linnaeus. One-humped Camel 

Camelus dromedarius Linnaeus, Syst. Nat., ed. 10, 1: 65, 1758. "Africa." 

Deserts of Libya and Arabia (see Thomas, O., Proc. Zool. Soc. London, 

p. 150, 22 Mar. 1911). 
The camel has never been known from Africa in a wQd state within historic 
times although reported as fossil in Algeria. For an account of its introduction 
by man into northeastern Africa, see Robinson, A. E., The extinct fauna of 
north-east Africa, Journ. Soc. for Preservation of the Fauna of the Empire, 
new ser., pt. 23, p. 43-48, Sep. 1934. 

Section Tragulina. Chevrotains 
Family TRAGULID.E 

Hyemoschus Gray. Water Chevrotains 

Hyemoschus Gray, Ann. Mag. Nat. Hist. (1) 16: 350, Nov. 1845. Genotype- 

Moschus aquaticus Ogilby. 
Variants: Hyeomoschus Turner, Proc. Zool. Soc. London, for 1849, p. 158, 
1850; Hyomoschus Blyth, Proc. Zool. Soc. London, for 1864, p. 483, May 1865; 
Hyaemoschus Zittel, Handb. d. Palaeontol., 4: pt. 2, p. 387, 1893. 

On the use of this name instead of Dorcatherimn, see Thomas, O., On the 
generic names applicable to the chevrotains (Tragulidae), Ann. Mag. Nat. Hist. 
(8) 18: 72-73, July 1916. 
Dorcatherium Kaup, Neues Jahrb. f. Mineral., p. 419, 1833; Descr. Ossem. Foss. 

Mamm. Mus. Darmstadt, no. 5, p. 91; Atlas, pi. 23, f. 1-16; 23 A; 23 B; 

23 C, f. 1-7, 1839. Genotype Dorcatherium naui Kaup (fossil, upper 

Miocene or lower Pliocene of Eppelsheim, Germany), in part, of Flower 

and Lydekker, and others. 

Hyemoschus aquaticus aquaticus (Ogilby) 

Moschus aquaticus Ogilby, Proc. Zool. Soc. London, for 1840, p. 35, Jan. 1841. 

Sierra Leone. 
Dorcatherium aquaticum typicum Lydekker, Proc. Zool. Soc. London, 1: 113, 

7 June 1906. 



466 bulletin: museum of comparative zoology 

Htemoschus aquaticus bates: (Lydekker) 

D[orcatherium] a[quaticum] batesi Lydekker, Proc. Zool. Soc. London, 1: 113, 
7 June 1906. Efulen, Cameroons. 

Htemoschus aquaticus cottoni (Lydekker) 

D[orcatherium] a[quaticu7n] cottoni Lydekker, Proc. Zool. Soc. London, 1: 113, 
7 June 1906. Ituri Forest, eastern Congo Beige. 



Section Pecora 
Family CERVID^. Deer 

Cervus Linnseus. Red Deer 

Cervus Linnseus, Syst. Nat., ed. 10, 1: 66, 1758. Genotype Cervus elaphus Lin- 
nseus. 

Elaphus Hamilton Smith, GrijEth's Cuvier, Animal Kingdom, 5: 307, 1827. 
Subgenus of Cervus, no type designated; included Cervus elaphus, C. cana- 
densis, C. occidentalis and C. wallichii. 

Harana Hodgson, Ann. Mag. Nat. Hist. (1) 1: 154, Apl. 1838. A subgenus of 
Cervus; type Cervus wallichii. 

Strongyloceros Owen, Hist. Brit. Fossil Mamm. and Birds, p. 469, f. 193, 195, 
1846. Asa subgenus of Cervus for C. spelieus Owen and C. elaphus Linnseus. 

Cervus elaphus barbarus Bennett. Barbary Deer 

Cervus barbarus Bennett, (List Animals in Gardens Zool. Soc. London, p. 31, 

1837); in L. Fraser, Zool. Typica, pi. 13, (about 1846). North Africa. 
Cervus corsiniacus Gervais, Ann. des Sci. Nat., zool., (3) 10: 206, 1848. 

Range: Local, and now rare in parts of Tunis, and Algeria (? Senegambia, 
apud Lydekker). 

Dama Hamilton Smith. Fallow Deer 

Dama Hamilton Smith in Griffith's Cuvier, Animal Kingdom, 5: 306, 1827. 
As a subgenus of Cervus, type Cervus dama Linnseus; raised to generic 
rank by Burnett (Quart. Journ. Sci., Lit. and Art, 28: 353, 1830). 

Platyceros Wagner, Schreber's Saugthiere, Suppl., 4: 347, 1844. As a subgenus 
of Cervus; type Cervus dama Linnaeus. 

Dactyloceros Wagner, Schreber's Saugthiere, Suppl., 5: 352, 1855. As a sub- 
genus of Cervus; type Cervus dama Linnseus. 

Dama dama dama (Linnseus). Fallow Deer 

Cervus dama Linnseus, Syst. Nat., ed. 10, 1: 67, 1758. No locality; based on 
captive deer in Sweden. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 467 

Cervus platyceros G. Cuvier, Tableau Elem. d'Hist. Nat. des Anim., p. 160, 1798. 
"Notre pays," i.e., France. 

Cervus mauricus F. Cuvier, Bull, des Sci., Soc. Philom., Paris, p. 82, 1816. No 
locality. 

Dama vulgaris Jardine, Naturalist's Library, Mamm., 3: 152, pi. 7, 1835. 
Moldavia and Lithuania. 
Range: Although attributed to Egypt on the basis of ancient Egyptian repre- 
sentations, these were probably made from introduced animals (see 
Dawson, W. R., Deer in ancient Egypt, Journ. Linn. Soc, London, zooL, 
39: 137-145, 14 text-f., 3 Oct. 1934). Rare in Algeria and Tunis, where, 
probably introduced. 



Family GIRAFFIDAE. Giraffes 

GiRAFFA Brisson 

Giraffa Brisson, Regnum Anim. in Classes IX Distrib., ed. 2, pp. 12, 37, 1762. 
Genotype Giraffa giraffa Brisson = Cervus camelopardalis Linnaeus. 

Camelopardalis Schreber, Saugthiere, 5: 1139, pi. 255*, 1784. Genotype Camelo- 
pardalis giraffa Schreber = Cervus camelopardalis Linnaus. 

Orasius Oken, Lehrbuch d. Naturgesch., pt. 3, sect. 2, p. 744, 1816. Genotype 
Cervus camelopardalis Linnaeus. 

Trachelotherium Gistel, Naturgesch. d. Thierreichs f. Hohere Schulen, p. 81, 
1848. New name for Camelopardalis Schreber. Genotype Giraffa camelo- 
pardalis (Linnaeus). 

Review: Lydekker, R. On the subspecies of Giraffa camelopardalis. Proc. 
Zool. Soc. London, 1 : 202-227, text-f. 23-37, pi. 9-16, 9 June 1904. 

Giraffa camelopardalis camelopardalis (Linnaeus). Nubian Giraffe 

Cervus camelopardalis Linnaeus, Syst. Nat., ed. 10, 1: 66, 1758. "^Ethiopia and 
Sennar," i.e., Cairo, Egypt, in captivity; assumed to be from the Abys- 
sinian, or Kassala, side of Upper Nubia, Anglo-Egyptian Sudan. 

Camelopardalis giraffa Gmelin, Linnaeus's Syst. Nat., ed. 13, 1: pt. 1, p. 181, 
1788. Sennaar between upper Egypt and Ethiopia. 

Camelopardalis sethiopica Ogilby, Prbc. Zool. Soc. London, for 1836, p. 134, 
27 June 1837. No locality. 

Giraffa camelopardalis angolensis Lydekker. Angola Giraffe. 

Giraffa camelopardalis angolensis Lydekker, Hutchinson's Animal Life, 2: 121, 

1903; Proc. Zool. Soc. London, 1: 221, pi. 14 (col.), 9 June 1904. Cunene 

River, 150 miles southwest of Humbe, Angola. 



468 bulletin: museum of comparative zoology 

GiRAFFA CAMELOPARDALis ANTiQUORUM (Jardine). Kordofan Giraffe 
Camelopardalis antiquorum Jardine, Naturalist's Library, Mamm., 3: 187, pi. 

21, 1835. "Senaar and Darfour," Anglo-Egyptian Sudan. 
Giraffa senaariensis Trouessart, Cat. Mamm. Viv. Foss., new ed., pt. 4, p. 902, 
1898. Name here credited to E. Geoffrey without further reference; it is 
evidently based on the account by I. Geoffroy, Ann. des Sci. Nat., (1) 11: 
210-223, pi. 22, 1827, of a giraffe from south of Sennaar, Anglo-Egyptian 
Sudan. 

Giraffa camelopardalis capensis (Lesson). Cape Giraffe 

Camelopardalis capensis Lesson, Nouv. Tabl. Regne Anim., Mammif., p. 168, 

1842. Cape of Good Hope. (Based on account by Levaillant, Voyage, 

pi. 8, 9, 1789.) 
Giraffa camelopardalis australis Rhoads, Proc. Acad. Nat. Sci. Philadelphia, 

p. 518, 8 Dec. 1896. South Africa (by implication). 
Giraffa camelopardalis capensis Lydekker, Proc. Zool. Soc. London, 1 : 224, 

text-f. 36, pi. 16 (col.), 9 June 1904. 

Giraffa camelopardalis congoensis Lydekker. Congo Giraffe 
Giraffa camelopardalis congoensis Lydekker, Hutchinson's Animal Life, 2: 83, 
1903; Proc. Zool. Soc. London, 1: 219, text-f. 33, 9 June 1904. Dungu, 
northeast of Uele district, Congo Beige, not Katanga as originally given 
(see Schouteden, H., Revue Zool. Africaine, 2: 134, 30 Sep. 1912). 
Range: Uele region to the Lado. 

Giraffa camelopardalis cottoni Lydekker 

Giraffa camelopardalis cottoni Lydekker, Proc. Zool. Soc. London, 1 : 207, text-f. 

25, pi. 15, f. 1, 9 June 1904. Koten plain, extreme south of the Topora 

(Doborsa) country, 3° 50' N., 34° 30' E., Uganda. 

Giraffa camelopardalis infumata Noack 

Giraffa infumata Noack, Zool. Anzeiger, 33: 356, 18 Aug. 1908. Barotse, middle 

Zambesi region, Northern Rhodesia. 
Giraffa camelopardalis thornicrofti Lydekker, Nature (London), 87: 484, 12 Oct. 

1911. Northeastern Rhodesia. 

Giraffa camelopardalis peralta Thomas. Nigerian Giraffe 
G[iraffa] c[amelopardalis] peralta Thomas, Proc. Zool. Soc. London, p. 40, 1 June 
1898. Near Lokoja, junction of Niger and Benue Rivers, Nigeria, 7° N., 

8°E. 

Giraffa camelopardalis rothschildi Lydekker 

Giraffa camelopardalis rothschildi Lydekker, Hutchinson's Animal Life, 2: 122, 

1903; Proc. Zool. Soc. London, 1 : 210, text-f. 26, 27, pis. 12, 13, 9 June 1904. 

Guas Ngishu plateau, east of Lake Baringo, Kenya Colony. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 469 

GiRAFFA CAMELOPARDALis TippELSKiRCHi Matschie. Kilimanjaro Giraffe 
Giraffa tippelskirchi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 78, 1898. 

Lake Eyassi, southeast of Victoria Nyanza, Tanganyika Territory. 
Giraffa schillingsi Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 79, 1898. 

Taveta, Kenya Colony. 
Giraffa camelopardalis tippelskirchi Lydekker, Froc. Zool. Soc. London, 1: 214, 

text-f. 28-32, 9 June 1904. 

Giraffa camelopardalis wardi Lydekker. Northern Transvaal Giraffe 
Giraffa camelopardalis wardi Lydekker, Proc. Zool. Soc. London, 1: 221, text-f. 
34, 35, pi. 15, f. 2, 9 June 1904. Northern Transvaal. 

Giraffa reticulata De Winton. Reticulated Giraffe 

Giraffa camelopardalis reticulata De Winton, Ann. Mag. Nat. Hist. (7) 4: 212, 

Sep. 1899. Loroghi Mts., Kenya Colony. 
Giraffa hagenbecki Knottnerus-Meyer, Zool. Anzeiger, 35: 800, 21 June 1910. 

Gallaland, southern Abyssinia. 
G[iraffa] reticulata nigrescens Lydekker, Nature (London), 87: 484, 12 Oct. 

1911. Probably the district north of Mt. Kenya, Kenya Colony. 



Okapia Lankester. Okapi 

Okapia Lankester, Nature (London), 64:24 7, 4 July 1901. Genotype Okapia 

johnstoni (Sclater). 
Ocapm Lankester, Science, new ser., 14: 114, 19 July 1901, ("quoting London 

Times, June 18"); variant. 

Okapia johnstoni (P. L. Sclater). Okapi 

Equus{?) johnstoni P. L. Sclater, Proc. Zool. Soc. London, 1: 50, text-f. 7, 1 
June 1901. Mundala, Semliki forest, eastern Congo Beige. 

Okapia tigrinum Johnston, Cornish's Living Animals of the World, 1 : 270, 1901. 

Okapia johnstoni Lankester, Nature (London), 64: 247, 4 July 1901; Trans. 
Zool. Soc. London, 16: 279-314, pi. 30-32, 15 text-f., Aug. 1902; Mono- 
graph of the okapi, atlas, 48 pis., 1910 (Brit. Mus). Semliki forest, Congo 
Beige near Fort Mbeni. 

Okapia erikssoni Lankester, Ann. Alag. Nat. Hist. (7) 10: 417, Nov. 1902. 
Semliki forest, Congo Beige. 

Okapia liebrechtsi Major, Proc. Zool. Soc. London, 2: 73, text-f. 7, Oct. 1902. 
Mundalah, on road from Mawambi to Beni, Congo Beige. 

Ocuapia (sic) kibalensis Gatti, Great Mother Forest, (privately printed), p. 
295, plates, 1937. Southwest of Lake Albert, eastern Congo Beige. 

On the supposed races and their identity, see Schouteden, H., Notes sur 
I'okapi, Revue Zool. Africaine, 2: 482-485, 30 May 1913. 



470 bulletin: museum of comparative zoology 

Family BOVIDyE. Cattle, Antelope, Goats 
SubfamUy ALCELAPHINiE 

Alcelaphus Blainville. Hartebeests 

Alcelaphus Blainville, Bull. des. Sci., Soc. Philom., Paris, p. 75, May 1816. 
Genotype Antilope bubalis PaWas = ArUilope buselaphus Pallas (see 
Thomas and Sclater, Book of Antelopes, 1: pt. 1, pp. 5, 7, Aug. 1894). 

Bubalis Goldfuss, Handbuch d. Zool., 2: 367, 1820. Genotype Antilope 
buselaphus Pallas. As usually quoted from Frisch, 1775, this name is not 
available since that author was non-binomial ; nor is Bubalis Lichtenstein, 
1814, valid (see Lyon, M. W., Jr., Proc. Biol. Soc. Washington, 27: 228, 
29 Dec. 1914; Hollister, N., Proc. Biol. Soc. Washington, 34: 77, 31 Mar. 
1921). 

Damalis Hamilton Smith, Griffith's Cuvier, Animal Kingdom, 4: 343, 1827. 
Genotype not designated but taken as Antilope buselaphus Pallas. 

Acronotus Hamilton Smith, Griffith's Cuvier, Animal Kingdom, 4: 346, 1827. 
As a subgenus of Damalis, type Antilope buselaphus Pallas. 

Bubalus Ogilby, Proc. Zool. Soc. London, for 1836, p. 139, 27 June 1837, not ot 
A. Smith, 1827. Genotype Bubalus mauritanicus Ogilhy = Antilope 
bubalis Pallas. 

Sigmoceros Heller, Smithsonian Misc. Coll., 60: no. 8, p. 4, 2 Nov. 1912. 
Genotype Bubalis lichtensteini Peters. 

Review: Ruxton, A. E., and Schwarz, Ernst. On hybrid hartebeests and on 
the distribution of the Alcelaphus buselaphus group. Proc. Zool. Soc. London, 
p. 567-583, map, 3 graphs, 2 pis., 16 Oct. 1929. 

Lydekker, R., and Blaine, Gilbert. Catalogue of the ungulate mammals in 
the British Museum (Natural History). Vol. 2, pp. 2-58, 1914. 

Alcelaphus buselaphus buselaphus (Pallas). Bubal Hartebeest 
Antilope buselaphus Pallas, Miscellanea Zool., p. 7, 1766. Probably Morocco. 
Antilope bubalis Pallas, Spicilegia Zool., pt. 1, p. 12, 1767; pt. 12, p. 16, 1777. 

"Africse . . . borealiori." 
Bubalus mauretanicus Ogilby, Proc. Zool. Soc. London, for 1836, p. 139, 

27 June 1837. No locality; a new name for Antilope bubalis Pallas. 
A[lcelaphus] bubalinus Flower and Lydekker, Introduct. to Study of Mamm., 

p. 335, 1891. 
Boselaphus probubalis Pomel, Carte Geol. Algerie, Paleont., Bosel., p. 27, pi. 

4, f. 1-11, 14-15; pi. 5, f.l; pi. 6, f.1-13; pi. 7, f.1-6; pi. 8, f. 1-9; pi. 9, 

f.1-4; pi. 10, f.1-5, 1894. Aboukir, Algeria. 
Boselaphus ambiguus Pomel, Carte Geol. Algerie, Paleont., Bosel., p. 52, pi. 

4, f. 12, 13; pi. 6, f. 14-19; pi. 8, f. 10-12; pi. 9, f. 5-7; pi. 10, f. 6-8, 1894. 

Ternifine, Algeria. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 471 

Boselaphus saldensis Pomel, Carte Geol. Algerie, Paleont., Bosel., p. 48, pi. 5, 
f. 3-19, 1894. Bougie, Algeria. 

Bubalis boselaphus Trouessart, Cat. Mamm. Viv. Foss., new ed., 2: pt. 4, 
p. 905, 1898; emendation. 

Bubalis bubastis Blaine, Ann. Mag. Nat. Hist. (8) 13: 335, Mar. 1914. Abadi- 
yeh, near Kena, Eg3rpt, from excavations. 
Ruxton and Schwarz (Proc. Zool. Soc. London, p. 575, 16 Oct. 1929) show 

that this is synonymous with A. buselaphus (Pallas). 

Range : Formerly the central part of eastern Morocco, at the east foot of the 
Great Atlas, in the region between high Muluya, the springs of Guir and 
the small streams that form the Uad Ziz, thence south to the South 
Oranes, between Geryville and Xott Tigri; at one time eastward probably 
to Eg3T)t. Now nearly, or quite extinct (see Cabrera, 1932). 

Alcelaphus buselaphus cokii Giinther. Coke's Hartebeest 

Alcelaphus cokii Giinther, Ann. Mag. Nat. Hist. (5) 14: 426, text-f. 1, Dec. 

1884. Mlali plains, east of Mpwapwa, Tanganyika Territory. 
Bubalis cookei Noack, Sitzb. Ges. Naturf. Freunde, Berlin, pp. 94, 95, 1905; 

misspelling of cokii. 
Bubalis cokei sabakiensis Zukowsky, Arch. f. Naturgesch., 79: sect. A, pt. 

10, p. 97, pi. 1, 1913; based on account and fig. in Wild und Hund, 16: 

225, 1910. Athi plains, south of Nairobi, Kenya Colony. 
Bubalis cokei tame Matschie and Zukowsky, Arch. f. Naturgesch., 79: sect. A, 

pt. 10, p. 97, 1913; 80: sect. A, pt. 1, p. 167, upper fig., 1914. North of 

Nairobi, Tana Region, Kenya Colony. 
Bubalis cokei wembserensis Zukowsky, Arch. f. Naturgesch., 79: sect. A, pt. 10, 

p. 98, 1913; based on account in Wild und Hund, 16: 208, 1910. Wem- 

baere plains, Lake Ushuto and Mkalamo, Tanganyika Territory. 
Bubalis cokei schillingsi Zukowsky, Arch. f. Naturgesch., 79: sect. A, pt. 10, 

p. 99, 1913; 80: sect. A, pt. 1, p. 167, lower fig., 1914. Ex Matschie in 

Schillings's "Mit BlitzUcht und Buchse," p. 384, 1910 (nomen nudum). 

Lake Jipe, southeast of Mt. Kilimanjaro, upper Pangani, Tanganyika 

Territory. 
Bubalis cokei schulzi Zukowsky, Arch. f. Naturgesch., 80: sect. A, pt. 9, p. 105, 

text-f. 104, 1914. Olossirva plateau, about 10 km. northeast of Ngoro- 

ngoro crater, Tanganyika Territory, 2500 m. 
Bubalis oscari Matschie and Zukowlsky, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. — , pi. 6, f. 3, 4, 10 Sep. 1916. Gurui, Masai country, and between 

Kapalata and Saranda, Tanganyika Territory. 
B[ubalis\ deckeni Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. — , pi. 8, f. 2, 10 Sep. 1916. Taveta plain, Kihmanjaro, Tanganyika 

Territory. 
Range: Plains of Kenya Colony and Tanganyika Territory. 



472 bulletin: museum of comparative zoology 

Alcelaphus buselaphus invadens (Schwarz) 

Bubalis major invadens Schwarz, Ann. Mag. Nat. Hist. (8) 13: 33, Jan. 1914. 
Garua, Benue River, Adamana, Cameroons. 

Alcelaphus buselaphus jacksoni (Thomas). Jackson's Hartebeest 
Bribalis jacksoni Thomas, Ann. Mag. Nat. Hist. (6) 9: 386, May 1892. Be- 
tween Lake Victoria and Lake Naivasha, Kenya Colony. 
Bubalis jacksoni insignis Thomas, Abstract Proc. Zool. Soc. London, no. 6, 
p. 22, 26 Apl. 1904; Proc. Zool. Soc. London, 1 : 455, 2 Aug. 1904. Maanja 
River, 30 miles west of Kampala, central Uganda. 

Alcelaphus buselaphus lelwel (Heuglin). Lelwel Hartebeest; Heuglin's 

Hartebeest 
A[cronotus] lelwel Heuglin, Reise in Nordost-Afrika, 2: 124, upper fig. on pi. 

facing same page, 1877. Req country, Jur, and Kosange River; type 

locality fixed as Jur River, Anglo-Egj^itian Sudan (see Schwarz, Ergebn. 

d. Zweite Deutsch. Zentr. Afr. Exped., 1 : 907, 1920; Ruxton and Schwarz, 

Proc. Zool. Soc. London, p. 577, 16 Oct. 1929). 
Bubalis niediecki Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 95, 1905. 

Jamboland, Gelo River, upper Sobat River, Abyssinia. 
Bubalis lelwel heuglini Millais, Far Away up the Nile (London), p. 244, 1924. 

Substitute for Acronotus lelwel Heuglin, 1877. 

Alcelaphus buselaphus major (Blyth). Western Hartebeest 

Boselaphus major Blyth, Proc. Zool. Soc. London, p. 52, text-f. 1, la, June 

1869. "Received from the west coast of Africa." 
Alcelaphus bubalis var. tunisianus Gray, Cat. Spec. Mamm. Brit. Mus., 

pt. 3, Ungulata Furcipeda, p. 123, 1852. "Tunis." 
Bubalis luzarchei G. Grandidier, Bull. Mus. d'Hist. Nat., Paris, 20: 316, 

2 text-f., 1914. Nieri-Ko River, near jimction with Gambia River, French 

Senegal. 
Range: Upper Gambia and Upper Senegal, French Guinea, to ?Tunis. 

Alcelaphus buselaphus matschiei (Schwarz). Matschie's Hartebeest 
Bubalis major matschiei Schwarz, Ann. Mag. Nat. Hist. (8) 13: 34, Jan. 1914. 

District of Kpandu, western Togo, Gold Coast. 
Alcelaphus buselaphus matschiei Ruxton and Schwarz,, Proc. Zool. Soc. London, 

p. 576, 16 Oct. 1929. 

Alcelaphus buselaphus modestus (Schwarz) 

Bubalis lelwel modestus Schwarz, Ann. Mag. Nat. Hist. (8) 13: 32, Jan. 1914. 

Bahr Keeta, northeast of Fort Archambault, Upper Shari district, French 

Equatorial Africa. 
Alcelaphus buselaphus modestus Ruxton and Schwarz, Proc. Zool. Soc. London, 

p. 577, 16 Oct. 1929. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 473 

Alcelaphus buselaphus roosevelti (Heller). Roosevelt's Hartebeest 
Bubalis lelwel roosevelti Heller, Smithsonian Misc. Coll., 60: no. 8, p. 7, 2 Nov. 

1912. Gondokoro, Uganda. 

Alcelaphus buselaphus swaynei (P. L. Sclater). Swayne's Hartebeest 
Bubalis sivayiiei P. L. Sclater, Proc. Zool. Soc. London, p. 98, pi. 5, text-f. 1, 

June 1892. Hand, about 100 miles from Berbera, northern Somaliland 

(Swayne, Proc. Zool. Soc. London, p. 303, 1892). 
Bubalis noacki Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 93, 1905. 

Suksuki River, south of Lake Zwai, southern Abyssinia. 
Range: Northern Somaliland and southern Abyssinia. 

Alcelaphus buselaphus tora (Gray). Tora Hartebeest 

Alcephalus (sic) tora Gray, Nature (London), 8: 364, 4 Sep. 1873; Alcelaphus 

tora Gray, Ann. Mag. Nat. Hist. (4) 12: 341, Oct. 1873. Dembelas, Bogos 

country, Abyssinia. 
Range: Eritrea to Blue Nile, Anglo-Egyptian Sudan. 

Alcelaphus buselaphus tschadensis (Schwarz) 

Bubalis lelwel tschadensis Schwarz, Ann. Mag. Nat. Hist. (8) 11: 265, Feb. 

1913. Ketekma, east of Tschekna, Bagirmi, French Equatorial Africa. 

Hybrids 

While Lydekker and Blaine (1915) regard the buselaphus group as a species 
distinct from the lelwel group, Ruxton and Schwarz (1929) consider the races 
of both as subspecies of Alcelaphus buselaphus. The latter authors believe the 
following names are based on hybrids: 

Alcelaphus buselaphus jacksoni X cokii 

Bubalis cokei kongoni Heller, Smithsonian Misc. Coll., 60: no. 8, p. 5, 2 Nov. 

1912. Loita Plains, Southern Guaso Nyiro, Kenya Colony. 

Bubalis cokei ritchiei Ruxton, Ann. Mag. Nat. Hist. (9) IS: 215, Aug. 1926. 

Southern Uasin Gishu country, south of Kisii, west of Amala (Mara) 

River, and north of Magoro River, Kenya Colony. 
Bubalis lelwel kenise Heller, Smithsonian Misc. Coll., 61: no. 17, p. 3, 21 Oct. 

1913. North Kenya plateau, 20 miles northeast of Nyeri, near Meru 
Road, Kenya Colony. 

Bubalis nakurse Heller, Smithsonian Misc. Coll., 60: no. 8, p. 6, 2 Nov. 1912. 
Nakuru, Rift Valley, Kenya Colony. 

?Alcelaphus buselaphus lelwel X swaynei 

Bubalis rothschildi Neumann, Sitzb. Ges. Naturf. Freimde, Berlin, p. 94, 1905. 
Adoshebai Valley, 80 km. north of Lake Stephanie, Abyssinia. 

Alcelaphus buselaphus lelwel X tora 

Bubalis neumanni Rothschild, Ann. Mag. Nat. Hist. (6) 20: 376, Oct. 1897. 
East shore and northeast of Lake Rudolf, Abyssinia. 



474 bulletin: museum of comparative zoology 

Bubalis torn digglei Rothschild, Ann. Mag. Nat. Hist. (8) 12: 576, Dec. 1913. 

Ofat River, near Keili, upper Blue Nile. 
Bubalis tora rahatensis Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 246, 

text-f. 2, Nov. 1906. Shunfar Ambu, western Abyssinia. 

Alcelaphus caama caama (G. Cuvier). Cape or Red Hartebeest 

Antilope caama G. Cuvier, Dictionnaire des Sci. Nat., 2: 242, 1816. Cape of 

Good Hope. 
Antilope koba Erxleben, Syst. Regni Animalis, p. 293, 1777. Description and 

fig. mentioned are of Alcelaphus caama. "ad Senegal." 
Antilope senegalensis G. Cuvier, Dictionnaire des Sci. Nat., 2: 235, 1816. 

Based on Pennant's Senegal Antelope, but the description and woodcut of 

head are of Alcelaphus caama. Senegal. 
Bubalis cama Bryden, Great and Small Game of Africa, p. 133, 1899. 
Bubalis caama obscurus Frechkop, Bull. Mus. Roy. d'Hist. Nat. Belg., Brux- 

elles, 13: no. 39, p. 11, 12, 22, pi. 1 (col.), text.f. 1, e, Oct. 1937. Locality 

unknown. 
Range: Formerly from the Cape Town district north to the Limpopo River 

and the edge of the Kalahari Desert; now extinct. 

Alcelaphus caama evalensis (Monard) 

Bubalis caama evalensis Monard, Bull. Soc. Neuchateloise des Sci. Nat., 
Neuchatel, 57: 64, figs. 9, 10, 1933. Evale, Angola. 

Alcelaphus caama selbornei (Lydekker). Selborne's Hartebeest 

Bubalis caama selbornei Lydekker, Abstract Proc. Zool. Soc. London, no. 119, 

p. 19, 15 Apl. 1913; Proc. Zool. Soc. London, p. 818, text-f. 135, 1 Sep. 

1913. Kimberley, where apparently imported from the Transvaal. 

Alcelaphus lichtensteinii lichtensteinii (Peters). Lichtenstein's Harte- 
beest 

Antilope lichtensteinii Peters, Reise nach Mossambique, Zool., Siiugeth., 
p. 190, pis. 43, 44, 1852. Tette, Mozambique. 

Bubalis leucoprymnus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 137, 
1892. Ulvami, Tanganyika Territory (see Matschie and Zukowsky, 
Sitzb. Ges. Naturf. Freunde, Berlin, p. 192, 30 Sep. 1916). 

Bubalis lichtensteini basengas Matschie and Zukowsky, Zool. Beobachter, 51: 

261, text-f., Sep. 1910; p. 377, Nov. 1910. Boundary between Shir6 
region and lower Loangwa region, Mozambique. 

Bubalis lichtensteini gombensis Matschie and Zukowsky, Zool. Beobachter, 51 : 

262, Sep. 1910. Kwa Kinga, Tabora district, Tanganyika Territory. 
Bubalis lichtensteini rukwse Matschie and Zukowsky, Zool. Beobachter, 51«: 

259, text-f., Sep. 1910; Sitzb. Ges. Naturf. Freunde, Berlin, pi. 6, f. 5, 
30 Sep. 1916; for 1917, pi. 19, f. 7; pi. 20, f. 3, 5, p. 534, 538, 28 Feb. 
1918. Kinani, southern Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 475 

Bubalis lichtensteini shirensis Matschie and Zukowsky, Zool. Beobachter, 51 : 

261, text-f., Sep. 1910; 51: 377, Nov. 1910; Sigmoceros shirensis Sitzb. 
Ges. Naturf. Freunde, Berlin, pp. 194, 201, 30 Sep. 1916. Boundary 
between Shire and lower Loangwa regions, Mozambique. 

Bubalis lichtensteini ugalse Matschie and Zukowsky, Zool. Beobachter, 51: 

262, Sep. 1910. Kwa Kinga, Tabora district, Tanganyika Territory. 
Bubalis lichtensteini ufipse Matschie and Zukowsky, Zool. Beobachter, 51: 

259, Sep. 1910. Kinani, southern Tanganyika Territory. 
See Sitzb. Ges. Naturf. Freunde, Berlin, pi. 6, 1916; for 1917, pp. 534, 

538, pi. 19, f. 1-4, 6; pi. 20, f. 2, 8, 28 Feb. 1918. 
Sigmoceros basengse Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 201, pi. 5, f. 5; pi. 8, f. 5, 30 Sep. 1916. Between Senna and 

Tette, Zambesia. 
[(Sigmoceros] heuferi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 202, 30 Sep. 1916. Substitute name for basengae, since that form 

does not occur at Basenga. 

The status of the following is questionable: — 

Sigvioceros bangx Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 206, pi. 7, f. 4, 30 Sep! 1916. Banga, southeast of Baunza, and 

north of Kafue River, Mozambique. 
Sigmoceros dieseneri Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 86, 30 Apl. 1925; figured as S. gombensis Matschie 

and Zukowsky, Sitzb. Ges. Naturf. Freunde, Berlin, pi. 7, f. 3, 30 Sep. 

1916. Rumosso, Runsewe, Tanganyika Territory. 
Sigmoceros frommi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1917, p. 532, pi. 20, f. 7, 28 Feb. 1918. Malinga Mts., southwest 

of Lake Rukwa, Tanganyika Territory. 
Sigmoceros gendagendse Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 119, pi. 7, f. 8, 30 Apl. 1925. Mt. Genda-Genda, 

Tanganyika Territory. 
Sigmoceros godonga Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 197, 30 Sep. 1916. Urema River, a branch of the Pungwe, 

Cheringoma district, Mozambique. 
Sigmoceros godowiusi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 80, pi. 5, f. 1, 3, 30 Apl. 1925. Uschingo, Tanganyika 

Territory. 
Sigmoceros gorongozse Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 196, 30 Sep. 1916. Northern Gorongoza district, 60 km. west of 

Urema River, Mozambique. 
Sigmoceros grotei Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 137, pi. 7, f. 1, 30 Apl. 1925. South of Newala on the 

Rovuma, Tanganyika Territory. 



476 bulletin: museum of compar,\tive zoology 

Sigmoceros hennigi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1922, p. 129, pi. 7, f. 2, 5, 30 Apl. 1925. Hinterland of Kilvva, 
Tanganyika Territory. 

Sigmoceros inkxdanondo Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, p. 197, 30 Sep. 1916. "Unzeilas Reich", upper Sabi, southeastern 
Mashonaland, Rhodesia. 

Sigmoceros janenschi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1922, p. 122, pi. 6, f. 1, 2; pi. 7, f. 6, 30 Apl. 1925. Makangaga, 
between Matandu and Mawudji, Tanganyika Territory. 

Sigmoceros kangosa Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1917, p. 532, 28 Feb. 1918. Makjusa's Land, northwest end of 
Lake Nyasa, near Ipiana, Nyasaland. 

Sigmoceros konzi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, Ber- 
lin, p. 203, pi. 7, f. 6, 30 Sep. 1916. North of the middle Kafue, 50 km. 
south of Broken Hill, northwestern Rhodesia. 

Sigmoceros lacrymalis Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1922, p. Ill, 30 Apl. 1925. Kangatta plains, Tanganyika Terri- 
tory. 

Sigmoceros lademanni Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, pi. 5, f. 1, 30 Sep. 1916. Kisaki, Tanganyika Territory. 

Sigmoceros leupolti Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, pi. 7, f. 4, 30 Sep. 1916; for 1922, p. 94, 30 Apl. 1925. Sikonge, 
south of Tabora, Tanganyika Territory. 

Sigmoceros lindicus Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1922, p. 132, pi. 7, f. 3, 30 Apl. 1925. Hinterland of Lindi, 
Tanganyika Territory. 

Sigmoceros milnzneri Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1917, p. 542, pi. 19, f. 5; pi. 20, f. 1. 4, 28 Feb. 1918. Mbuga 
Mpimba, west of Panji, 7° 58' S., 31° 22' E., Tanganyika Territory. 

Sigmoceros niediecki Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, p. 205, pi. 7, f. 2, 5, 30 Sep. 1916. Baunza, north of middle Kafue, 
33 km. east of where the river turns east, north of Victoria Falls, Mozam- 
bique. 

Sigmoceros niedieckianus Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 295, 
15 Dec. 1916. New name for Sigmoceros niediecki Matschie and Zukowsky; 
not Bubalis niediecki Neumann, 1905. 

Sigmoceros petersi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1917, p. 530, 28 Feb. 1918. Near Sena, Mozambique. Type 
figured in Peters, Reise nach Mossambique, Siiugeth., pi. 44, 1852. 

Sigmoceros priUivitzi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 
Berlin, for 1922, p. 96, pi. 5, f. 5, 30 Apl. 1925. Mijange district, 30 km. 
south of Itumba Qua Kiromo, Issaua-Kedete watershed, Kisigo drainage, 
Tanganyika Territory. 



ALLEN: CHECKLLST OF AFRICAN MAMMALS 477 

Sigmoceros rendalli Matschie and Zukowsky, Sitzb. Ges. Natiirf. Freunde, 

Berlin, for 1922, p. 80, 30 Apl. 1925. Kambwe, south end of Lake Nyasa, 

Nyasaland; name based on account by Rendall, in Novitates Zool., 5: 214, 

3 May 1898. 
Sigvioceros rowumae Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 139, 30 Apl. 1925. Mt. Lisoju, upper Rovuma River, 

Tanganyika Territory. 
Sigmoceros saadanicus Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 116, 30 Apl. 1925. Mt. Gendagenda, Tanganyika 

Territory. 
Sigmoceros schmitti Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 98, pi. 6, f. 5, 6, 30 Apl. 1925. Probably the Mpawagga- 

Muhindo plains, Uhehe, Tanganyika Territory. 
Sigmoceros schusteri Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 113, 30 Apl. 1925. Ruimbi Pond, north of Kinamatine 

( = Kilimatinde), Tanganyika Territory. 
Sigmoceros senganus Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. 200, pi. 6, f. 1, 2, 30 Sep. 1916. Mussenda Luz, on the Zambesi, 

between the Loangwa and Kebabassa Falls, Mozambique. 
S[igmoce.ros] stierlingi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, p. -, pi. 8, f. 7, 30 Sep. 1916; for 1922, p. 103, 30 Apl. 1925. Upper 

Ruaha River, Tanganyika Territory. 
Sigmoceros tendagurucus Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 127, pi. 7, f. 7, 30 Apl. 1925. Mchuya, near Kilwa, 

Tendaguru, Tanganyika Territory. 
Sigm.oceros idangse Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 100, pi. 6, f . 3, 4, 30 Apl. 1925. Ulanga Plains, Tangan- 
yika Territory. 
Sigmoceros imgonicus Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berhn, for 1922, p. 134, 30 Apl. 1925. Ungoni, Tanganyika Territory. 
Sigmoceros ungoniensis Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 135, 30 Apl. 1925. Ungoni, Tanganyika Territory. 
Sigmoceros uwendensis Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1917, p. 545, pi. 20, f. 6, 28 Feb. 1918. Katjantia (Katjentje), 

north of Karema, near liake Tanganyika, 6° 36' S., 30° 22' E., Tanganyika 

Territory. 
Sigmoceros wiesei Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, Berlin, 

p. 199, pi. 5, f. 6, 7, 30 Sep. 1916. West of Chifumbazi, on the Luia, tribu- 
tary of the Kapotche, Mpeseni country, Rhodesia. 
Sigmoceros wintgensi Matschie and Zukowsky, Sitzb. Ges. Naturf. Freunde, 

Berlin, for 1922, p. 92, pi. 5, f. 4, 30 Apl. 1925. Southeast of Gombe River, 

Tanganyika Territory. 



478 bulletin: museum of comparative zoology 

CoNNOCiLETES Lichtenstein. Gnus 

Connochsetes Lichtenstein, Mag. d. Ges. Naturf. Freunde, Berlin, 6: 152, 165, 
1814. As a subgenus of Antilope, type Antilope gnu Gmelin. 
Also spelled Connochoetes, Connochetes. 

Cemas Oken, Lehrbuch d. Naturgesch., pt. 3, zool., sect. 2, p. 727, 1816. As a 
subgenus of Pecws; type fixed as Cemas gnu Oken = Bos gnou Zimmer- 
mann, by P. L. Sclater and Thomas (Book of Antelopes, 1 : pt. 2, p. 93, 111, 
Jan. 1895). 

Catablepas Gray, London Medical Repository, 15: 307, 1 Apl. 1821. Genotype 
Antilope gnu Gmelin. 

Catoblepas Hamilton Smith, Griffith's Cuvier, Animal Kingdom, 4: 366, 1827. 
Genotype Antilope gnu Gmelin. 

Gorgon Gray, Gleanings from Menagerie at Knowsley Hall, 2: 20, pi. 19, f. 2, 
1850; Proc. Zool. Soc. London, for 1850, p. 139, 24 Feb. 1851; Cat. Rumi- 
nant Mamm. Brit. Mus., p. 43, 1872. As a subgenus of Catoblepas; type 
Antilope gorgon Hamilton Smith = Antilope taurina Burchell. 
Regarded by Heller (Smithsonian Misc. Coll., 61: no. 17, p. 2, 21 Oct. 1913) 

as a valid genus. 

Butragus Gray (ex Blyth, MS.), Cat. Ruminant Mamm. Brit. Mus., p. 43, 
1872. Genotype Butragus corniculatus Blyth, MS., = Antilope taurinus 
Burchell. 

CoNNOCH.«TES ANTiQUUS Broom 

Connochaetes antiquus Broom, Ann. South African Mus., 12: 14, text-f. 2, 30 

May 1913. Extinct; Haagenstad, about 30 miles north of Bloemfontein, 

Cape of Good Hope. 

CoNNOCH.ETES GNOU (Zimmermann). Black Wildebeest or White-tailed Gnu 
Bos gno\L Zimmermann, Specimen Zoologiae Geographicse, p. 372, 1777. Cape 

of Good Hope. See Allen, J. A., Bull. Amer. Mus. Nat. Hist., 16: 15, 1 Feb. 

1902. 
Antilope capensis Gatterer, Brev. Zool., 1: 80, 1780. 
Antilope gmi Gmelin, Linnseus's Syst. Nat., ed. 13, 1: pt. 1, p. 189, 1788. Great 

Namaqualand, Cape of Good Hope. 
C[atoblepas] brooksii Hamilton Smith in Griffith's Cuvier, Animal Kingdom, 5: 

369, footnote, 1827. No locality, based on a horn of uncertain identity. 
Catoblepas operculatus Brookes, Cat. Brookes Mus., p. 64, 1828. 
Bos connochoetes Forster, Descrip. Animahum, p. 392, 1844. Cape of Good Hope. 

Connochetes taurinus taurinus (Burchell). Brindled Gnu; Blue Wilde- 
beest 

Antilope taurina Burchell, Travels in Interior of South Africa, 2: 278, foot- 
note, 1824. Bechuanaland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 479 

Catoblepas gorgon Hamilton Smith, Griffith's Cuvier, Animal Kingdom, 4: 371, 

pi., 1827; 5: 369, 1827. Interior of South Africa. 
Gorgon fasciatus Gray, Cat. Ruminant Mamm. Brit. Mus., p. 43, 1872. South 

Africa. 
Butragus corniculatus Gray (ex Blyth MS.), Cat. Ruminant Mamm. Brit. Mus., 

p. 43, 1872. South Africa. 
Catoblepas reichci Noack, Zool. Anzeiger, 16: 154, 1 May 1893. Limpopo, 

northern Transvaal. 

CoNNOCH^TES TAtTRiNus ALBOJUBATUS Thomas. White-bearded Wildebeest 
Connochsetes taurinus alhojuhatus Thomas, Ann. Mag. Nat. Hist. (6) 9: 388, 

May 1892. "Uganda" = Athi plains, Kenya Colony. 
Connochwtes (sic) hecki Neumann, Sitzb. Ges. Naturf. Freunde, Berlin, p. 96, 

1905. Tanganyika Territory, specimens described from Gunii. 
Gorgon alhojuhatns mearnsi Heller, Smithsonian Misc. Coll., 61: no. 17, p. 1, 

21 Oct. 1913. Loita plains, southern Kenya Colony. 
Connochsetus (sic) alhojuhatus henrici Zukowsky, Arch. f. Naturgesch, 79: sect. 

A, pt. 12, p. 83, pi. 3, upper f.; pi. 4, 1913. Seringeti steppes, Tanganyika 

Territory. 
Connochsetus (sic) alhojuhatus lorenzi Zukowsky, Arch. f. Naturgesch., 79: sect. 

A, pt. 12, p. 81, pi. 1, 1913. Ngorongoro crater, Tanganyika Territory. 
Connochsetus (sic) alhojuhatus schulzi Zukowsky, Arch. f. Naturgesch., 79: sect. 

A, pt. 12, p. 82, pi. 2: pi. 3, lower f., 1913. Ngorongoro basin, Tanganyika 

Territory. 

Connochwtes taurinus cooksoni Blaine. Cookson's Wildebeest 
Connochsetes taurinus cooksoni Blaine, Ann. Mag. Nat. Hist. (8) 13: 377, Mar, 
1914. East bank of Loangwa River, northeastern Rhodesia. 

Connochwtes taurinus johnstoni P. L. Sclater. Nyasa Wildebeest 
Connochsetes taurinus johnstoni P. li. Sclater, Proc. Zool. Soc. London, p. 606, 

pi. 28, Oct. 1896. Mlanje plain, south end of Lake Chilwa, Nyasaland. 
Connochsetes taurinus (vel johnstoni) rufijianus de Beaux, Zool. Anzeiger, 38 : 

579, text-f. 1, 2, 19 Dec. 1911. Rufigi valley, Tanganyika Territory. 

Connochwtes taurinus mattosi Blaine. Angola Wildebeest 

Connochsetes taurinus liiattosi Blaine; Ann. Mag. Nat. Hist. (9) 15: 129, Jan. 

1925. Cotypes from (1) Chiacusse, Huilla district, Cunene River; (2) 

Bamber River, a tributary of the Cunene; and (3) Chicoonga, northeast 

of Dongo, Angola. 
Connochsetes taurinus borlei Monard, Bull. Soc. Neuchateloise des Sci. Nat., 

Neuchatel, 57: 64, 1933. Angola, Rio Mbale (female) and Chimporo 

(male)— see ibid., 54: 75-76, 1930. 



480 bulletin: museum of comparative zoology 



Damaliscus p. L. Sclater and Thomas 

Damaliscus P. L. Sclater and Thomas, Book of Antelopes, 1: pt. 1, p. 3, 51, 
Aug. 1894. Genotype Antilope pygarga Pallas. 

Damalis Gray, Ann. Mag. Nat. Hist. (1) 18: 233, Oct. 1846. Genotype Damalis 
lunatus (Burchell), see Sclater and Thomas, Book of Antelopes 1: 51, Aug. 
1894. Preoccupied by Damalis H. Smith. 

Beatragus Heller, Smithsonian Misc. Coll., 60: no. 8, p. 8, 2 Nov. 1912. Geno- 
type Damaliscus hunteri (P. L. Sclater). 

Ajitilope, Alcelaphus, Buhalis, Capra, Cerophorus, Gazella, in part of older 
authors. 

Antilopa Blainville, in Gervais's Dictionnaire d. Sci. Nat., suppl. I, pt. 2, p. 
260, 1841. Emendation oi Antilope. 

Damaliscus albifrons (Burchell). Blesbok 

[Antilope] albifrons Burchell. Travels in Interior of South Africa, 2:335, 1824. 

Northern Cape Colony (Lydekker and Blaine). 
Range : Formerly South Africa to the northern plains of Cape Colony, Orange 

Free State, Transvaal, and Bechuanaland; now much restricted. 

Damaliscus hunteri (P. L. Sclater). Hvmter's Hartebeest or Hirola 

Cobus hunteri P. L. Sclater, Field (London), 73:260, 23 Feb. 1889 (nomen 

nudum). "Eastern Africa." North side of Tana Valley, Jubaland, eastern 

Kenya Colony. 
Damalis hunteri P. L. Sclater, Proc. Zool. Soc. London, p. 58, 1 text-f., June 

1889; p. 372, text-f. A-C, pi. 42 (col.), Oct. 1889. North side of Tana 

River, Kenya Colony. 
Range: Jubaland. 

Damaliscus korrigum korrigum (Ogilby). Korrigum or Senegal Hartebeest 
Antilope senegalensis Children, in Denham and Clapperton's Travels, append., 

p. 192, 1826. Plains of central Africa. 
Not of Cuvier, 1804, which is a description of Alcelaphus caama. 
A[ntilope] korrigum Ogilby, Proc. Zool. Soc. London, for 1836, p. 103, 20 Feb. 

1837. Bornu, northern Nigeria. 
Damaliscus corrigum H. C. V. Hunter, Great and Small Game of Africa, p. 166, 

1899. 
Damaliscus koba Lydekker and Blaine, Cat. Ungulate Mamm. Brit. Mus., 2: 

40, 1914; not Antilope koba Erxleben, 1777, which is Alcelaphus caama. 

Damaliscus korrigum eurus Blaine 

Damaliscus korrigum eurus Blaine, Ann. Mag. Nat. Hist. (8) 13:335, Mar. 1914. 
Upper Ruaha River, Tanganyika Territory. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 481 

Damaliscus korrigum jimela (Matschie). Topi 

Damalis jimela Matschie, Sitzb. Ges. Naturf. Freunde, Berlin, p. 135, 1892. 

Unyamwesi, southeast of Lake Victoria, Tanganyika Territory. 
Buhalis jimeru Lydekker, Horns and Hoofs, p. 203, 1893 (errorim). 
Damaliscus phalius Cabrera, Proc. Zool. Soc. London, for 1910, p. 998, text-f. 

149,. 11 Jan. 1911. Guas Ngishu plateau, northwestern Kenya Colony. 

Damaliscus korrigum lyra Schwarz 

Damaliscus koba lyra Schwarz, Ann. Mag. Nat. Hist. (8) 13: 34, Jan. 1914. 
Ndioko, Gribingi River, upper Shari district, French Equatorial Africa. 

Damaliscus kobrigum purpurescens Blaine 

Damaliscus korrigum purpurescens Blaine, Ann. Mag. Nat. Hist. (8) 13: 333, 
Mar. 1914. Ibi, Benue River, northern Nigeria. 

Damaliscus korrigum selousi Lydekker. Selous's Topi 
Damaliscus corrigum selousi Lydekker, Field (London), 110: 250, 10 Aug. 1907. 
Near Londiani plateau, Kenya Colony. 

Damaliscus korrigum tiang (Heuglin). Tiang 

Buhalis koba Sundevall, Kongl. Svenska Vet.-Akad. Handl., Stockholm, for 
1844, p. 209, 1846; not Antilope koba Erxleben, 1777. Sennaar, Senegal. 

Damalis tiang Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 30: no. 2, 
p. 22, pi. 1, f. 1, a, b, 1863. Sobat Valley, Anglo-Egyptian Sudan. 

Damalis tiang-riel Heuglin, Nova Acta Acad. Cses. Leop.-Carol., Halle, 30: 
no. 2, p. 23, pi. 2, f. 9, 1863. Bahr-el-Abiad, Anglo-Egyptian Sudan. 

Damaliscus corrigum jonesi Lydekker, Field (London), 110: 250, 10 Aug. 1907. 
Kordofan. 
Synonym of Damaliscus koba tiang according to Schwarz, Ann. Mag. Nat. 

Hist. (8) 13:35, Jan. 1914. 

Damaliscus floweri Matschie, Jahrbuch des Inst. f. Jagdkunde, 2: 168, 1913. 
Near Sherif Harrabulla, between Karkoj and Roseires, Blue Nile, Anglo- 
Egyptian Sudan. 

Damaliscus korrigum topi Blaine 

Damaliscus korrigum topi Blaine, Ann. Mag. Nat. Hist. (8) 13: 333, Mar. 1914. 
Near Malindi, Kenya Colony. 

Damaliscus korrigum ugand^ Blaine 

Damaliscus korrigum ugandx Blaine, Ann. Mag. Nat. Hist. (8) 13: 334, Mar. 
1914. Southwestern Ankole, Uganda. 



482 bulletin: museum of comparative zoology 

Damaliscus lunatus lunatus (Burchell). Sassaby or Bastard Hartebeest 
Antilope lunata Burchell, Travels in Interior of South Africa. 2: 334, 2 text- 
figs., 1824. Bechuanaland. 
Range: Southeastern Africa from north of the Orange River as far westward 
as Lake Ngami, north to northern Rhodesia. 

Damaliscus lunatus reclinis Matschie 

Damaliscus lunatus redinis'M.atscMe, Deutsche Jager-Zeitung, 59: 119 (77 of 
reprint), 1912. Caprivi Peak, Southwest Africa. 

Damaliscus pygargus (Pallas\ Bontebok 

Antilope dorcas Pallas, Miscellanea Zool., p. 6, 1766, in part. No locality. 

Antilope pygargus and pygarga Pallas, Spicilegia Zool., pt. 1, p.*0, 1767; pt. 12, 

p. 15, 1777. Cape of Good Hope implied. 
Antihpe grisea Boddsert, Elenchus Animalium, p. 139, 1785. Cape of Good 

Hope. 
Capra scripta Thunberg, Voyages au Japon, etc., 2: 47, 1796. Caffrerie. 
Antilope maculata Thunberg, Mem. Acad. Sci. St. Petersbourg, 3: 315, 1811. 
Antilope personala Woods, Zool. Journ., 5: 2, pi. 1, 1829. Vicinity of the Cape 

of Good Hope. 
Range: Formerly from Cape of Good Hope north to Orange River; now 

"existing only in a semi-domesticated condition on some farms in the 

Bredasdoop district of the Cape Province"' (Haagner, 1920). 



Subfamily CEPHALOPHIN^. Duikers 

See: Lonnberg, E. Notes on the members of the genera "Cephalophus" and 
"Sylvicapra" in the Congo Museum. Revue Zool. Africaine, 7: 162-185, 1 Oct. 
1919. 

Review: St. Leger, J. A kej^ to the species and subspecies of the subgenus 
Cephalophus. Proc. Zool. Soc. London, p. 209-228, 16 Apl. 1936. 



Cephalophus Hamilton Smith 

Cephalophus Hamilton Smith, Griffith's Cuvier, Animal Kingdom, 5: 344, 1827. 
As a subgenus of Antilope, including ten species; type fixed as Antilope 
silvicultrix Afzelius (see P. L. Sclater and Thomas, Book of Antelopes, 1 : 
pt. 3, p. 121, May 1895). 

Philantomba Blyth, in Cuvier's Animal Kingdom, p. 140, 1840. No type speci- 
fied. 

Cephalophora Gray, Ann. Mag. Nat. Hist. (1) 10: 266, Dec. 1842. Genotype 
Cephalophora coronata Gray. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 483 

Cephalophorus Gray, Ann. Mag. Nat. Hist. (1) 10: 267, Dec. 1842. Genotype 
Antilope ogilbyi Waterhouse. Intended as an emendation of Cephalophus. 

Cephalolophus Wagner, Schreber's Saugthiere, Suppl., 4: 445, 1844; emenda- 
tion. 

Terpone Gray, Proc. Zool. Soc. London, p. 592, Aug. 1871. Genotype Cephalo- 
phus longiceps Gray = Antilope silvicuUrix Afzelius. 

Potamotragns Gray, Cat. Ruminant Mamm. Brit. Mus., p. 25, 1872. Genotype 
Cephalophus melanoprymnus Greiy = Antilope silvicuUrix Afzelius. 

Cephalopia Knottnerus-Meyer, Arch. f. Naturgesch. 73: sect. A, pt. 1, p. 44, 
Jan. 1907. As a subgenus of Cephalophus; type not specified, but includes 
C. ogilbyi (Waterhouse) and C. leucogastra Gray. 

Cephalophella Knottnerus-Meyer, Arch. f. Naturgesch., 73: sect. A, pt. 1, p. 45, 
Jan. 1907. As a subgenus of Cephalophus; type Cephalophus callipygus 
Peters. 

Cephalophidium Knottnerus-Meyer, Arch. f. Naturgesch., 73: sect. A, pt. 1, 
p. 45, Jan. 1907. As a subgenus of Cephalophus; type Cephalophus niger 
Gray. 

Cephalophops Knottnerus-Meyer, Arch. f. Naturgesch., 73: sect. A, pt. 1, p. 46, 
Jan. 1907. As a subgenus of Cephalophus; type Cephalophus dorsalis Gray. 

Cephalophula Knottnenis-Meyer, Arch. f. Naturgesch., 73: sect. A, pt. 1, p. 46, 
Jan. 1907. As a subgenus of Cephalophus; type Cephalophus doria Ogilby. 



Subgenus Cephalophus Hamilton Smith 

Cephalophus adersi Thomas. Zanzibar Duiker 

Cephalophus adersi Thomas, Ann. Mag. Nat. Hist. (9) 2: 151, Aug. 1918. 
Island of Zanzibar. 



Cephalophus callipygus Peters. Peters's Duiker 

Cephalophus callipygus Peters, Monatsb. K. Preuss. Akad. Wiss., Berlin, p. 483, 
pi. 3, 4, 1876. Gaboon. 
Range: Gaboon and Cameroons. 

Cephalophus dorsalis dorsalis Gray. Bay Duiker 

Cephalophus dorsalis Gray, Ann. Mag. Nat. Hist. (1) 18: 165, Sep. 1846. 

Sierra Leone. 
Cephalophus badius Graj', Cat. Mamm. Brit. Mus., pt. 3, Ungulata Furcipeda, 

p. 84, 1852. Sierra Leone. 
Cephalophus breviceps Gray, Proc. Zool. Soc. London, p. 202, pi. 20, Sep. 1866. 

"West coast of Africa." 
Cephalophus dorsalis, subsp. typicus Thomas, Proc. Zool. Soc. London, p. 420, 

Oct. 1892. 



484 bulletin: museum of comparative zoology 

Cephalophus dorsalis arrhenii Lonnberg. Arrhenius's Bay Duiker 
Cephalophus castaneiis arrhenii Lonnberg, Kuiigl. Svenska Vet.-Akad. Handl., 
Stockholm, (2) 58: no. 2, p. 92, pi. 11, f. 3, 1 Sep. 1917. Beni, Congo Beige. 

Cephalophus dorsalis castaneus Thomas 

Cephalophus dorsalis castaneiis Thomas, Proc. Zool. Soc. London, p. 421, Oct. 

1892. Cameroons. 
Cephalophus dorsalis orientalis Schwarz, Ann. Mag. Nat. Hist. (8) 13: 34, Jan. 
1914. Koloka, near Angu, Uele River, Congo Beige. 
For comments on the possible invalidity of this race see Lonnberg, E., Revue 
Zool. Africaine, 7: 173, 1919; also St. Leger, J., Proc. Zool. Soc. London, p. 220, 
16 Apl. 1936, who considers it the same as C. d. castaneus Thomas. 
Cephalophus dorsalis kuha Lorenz, Ann. Naturhist. Mus. Wien, 36: 112, text-f. 
7, Oct. 1923. Ukaiko, Ituri district, Congo Beige. 
A synonym of C. d. castaneus according to St. Leger, Proc. Zool. Soc. London, 
p. 220, 16 Apl. 1936. 

Cephalophus dorsalis leucochilus Jentink 

Cephalophus lexicochilus Jentink, Notes Leyden Mus., 23: 21, July 1901. 

Loango, Angola. 
"Very probably" a race of C. dorsalis (fide Lydekker and Blaine). 
Cephalophus dorsalis leucochilus Schwarz, Ann. Mag. Nat. Hist. (8) 13: 493, 

May 1914. 

Cephalophus harveyi harveyi Thomas. Harvey's Duiker 
Cephalolophus (sic) harveyi Thomas, Ann. Mag. Nat. Hist. (6) 11 : 48, Jan. 1893. 
Kilimanjaro district, Kaha forest, west of Taveta, Tanganyika Territory. 
This is the oldest name applied to a member of the group in which St. Leger 
includes iveynsi and related suVjspecies. The latter therefore should stand as 
subspecies of harveyi rather than of weynsi. 

Cephalophus harveyi keniae Lonnberg, Ann. Mag. Nat. Hist. (8) 9: 65, Jan. 
1912; Kongl. Svenska Vet.-Akad. Handl., Stockholm, (2) 48: no. 5, p. 150, 
26 June 1912. Near Nairobi, Kenya Colony. 
Identical with C. h. harveyi according to St. Leger (Proc. Zool. Soc. London, 
p. 218, 16 Apl. 1936). 

Cephalophus harveyi barbertoni Kershaw 

Cephalophus barbertoni Kershaw, Ann. Mag. Nat. Hist. (9) 11: 596, May 1923. 

Mt. Elgon, northeast of Lake Victoria, Kenya Colony, 7-8000 ft. 
Cephalophus weynsi barbertoni St. Leger, Proc. Zool. Soc. London, p. 217, 16 

Apl. 1936. 

Cephalophus harveyi bottegoi de Beaux 

Cephalophus harveyi bottegoi de Beaux, Atti Soc. Ital. Sci. Nat. e Mus. Civ. Stor. 
Nat. Milano, 63: 6, July 1924. Gourar Ganana, Italian Somaliland. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 485 

Cephalophtjs harveyi ignifer Thomas 

Cephalophus ignifer Thomas, Proc. Zool. Soc. London, 1: 226, Aug. 1903. 

Eldoma Ravine, Kenya Colony. 
Cephalophus weynsi ignifer St. Leger, Proc. Zool. Soc. London, p. 218, 16 Apl. 

1936. 

Cephalophus harveyi rutshuricus Lonnberg 

Cephalophus weynsi rutshuricus Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, (2) 58: no. 2, p. 86, pi. 10, f. 2, 1 Sep. 1917. Kabwe, eastern 

Congo Beige. 

Cephalophus harveyi weynsi Thomas. Weyns's Duiker 
Cephalophus weynsi Thomas, Ann. Mus. Congo, Zool., 2: pt. 1, p. 15, pi. 6, Aug. 
1901. Near Stanley Falls, Congo Beige. 
In her review of Cephalophus, Miss St. Leger makes various described forms 
related to weynsi subspecies of it, but the form harveyi was first described and 
hence becomes the typical subspecies. 

Cephalophus johnstoni Thomas, Proc. Zool. Soc. London, 2: 89, 1 Oct. 1901. 
Toro district, east of Mt. Ruwenzori, Uganda. 
A synonym of C. h. weynsi according to St. Leger (Proc. Zool. Soc. London, 
p. 216, 16 Apl. 1936). 

Cephalophus centralis M. de Rothschild and Neuville, Compt. Rend. Acad. Sci. 
Paris, 144: 217, 1907. Ituri valley, eastern Congo Beige. 
A synonym of C. h. weynsi according to St. Leger (Proc. Zool. Soc. London, 
p. 216, 16 Apl. 1936). 

Cephalophus leopoldi M. de Rothschild and Neuville, Compt. Rend. Acad. Sci. 
Paris, 143: 1257, 1906 (? = 1907). Ituri valley, near Irumu, eastern Congo 
Beige. 
A synonym of C. h. weynsi according to St. Leger (Proc. Zool. Soc. London, 
p. 216, 16 Apl. 1936). 

Range: From lower Uele (Bambili) and the Ituri Forest (Beni) south to the 
Stanleyville district and vicinity of Lake Leopold II, Congo Beige (see 
Lonnberg, E., Revue Zool. Africaine, 7: 164, 1 Oct. 1919, for comments). 

Cephalophus jentinki Thomas. Jentink's Duiker 

Antilope (Terpone) longiceps Jentink, Notes Leyden Mus., 7: 272, pi. 10, Oct. 

1885, not Cephalophus longiceps Gray. Liberia. 
Cephalophus jentinki Thomas, Pro>.. Zool. Soc. London, p. 417, Oct. 1892. 

Liberia. 
Range: Apparently confined to Liberia. 

Cephalophus leucogaster Gray. Gaboon Duiker 

Cephalophus leucogaster Gray, Ann. Mag. Nat. Hist. (4) 12: 43, July 1873. 
Gaboon. 



486 bulletin: museum of comparative zoology 

Cephalophus leucogaster seke Lorenz, Ann. Naturhist. Mus. Wien, 36: 104, text-f. 
5, 8, Oct. 1923. Mawambi and Ukaika, Ituri district, Congo Beige. 
Identical with typical race according to St. Leger, J., Proc. Zool. Soc. London, 
p. 222, 16 Apl. 1936. 

Cephalophus natalensis natalensis A. Smith. Red or Natal Duiker 
Cephalophus natalensis A. Smith, South African Quart. Journ., 2: 217, 1834; 

Illustr. Zool. South Africa, Mamm., pi. 32 and text, Jan. 1841. Port Natal, 

Natal. 

Cephalophus natalensis amcenus Wroughton 

Cephalophus natalensis amcenus Wroughton, Ann. Mag. Nat. Hist. (8) 8: 277, 

278, Aug. 1911. Legogot, Transvaal. 

Cephalophus natalensis bradshawi Wroughton. Bradshaw's Duiker 
Cephalophus natalensis hradshawi Wroughton, Ann. Mag. Nat. Hist. (8) 8: 277, 

279, Aug. 1911. Chiromo, Shire valley, Nyasaland. 

May be merely a melanistic form of C. natalensis walkeri Thomas (St. Leger, 
J., Proc. Zool. Soc. London, p. 219, 16 Apl. 1936). 

Cephalophus natalensis lebombo Roberts 

Cephalophus natalensis lebombo Roberts, Ann. Transvaal Mus., 18: 248, 21 Feb. 
1936. Mkusi River, near Ubombo, Mozambique. 

Cephalophus natalensis robertsi W. Rothschild. Roberts's Duiker 
Cephalophus robertsi W. Rothschild, Proc. Zool. Soc. London, p. 691, 10 Oct. 

1906. Mozambique. 
Cephalophus natalensis vassei Trouessart, Bull. Mus. d'Hist. Nat., Paris, p. 445, 

1906. Guenguere, Mozambique. 
Synonym of robertsi (see Wroughton, Ann. Mag. Nat. Hist. (8) 8: 277, Aug. 
1911). 

Cephalophus natalensis walkeri Thomas 

Cephalophus walkeri Thomas, Abstract Proc. Zool. Soc. London, no. 31, p. 1, 

8 May 1906; Proc. Zool. Soc. London, p. 464, pi. 35, 10 Oct. 1906. Tuchihi 

River, about 25 miles from Blantyre, Nyasaland. 
Cephalophus natalensis walkeri Schwarz, Ann. Mag. Nat. Hist. (8) 13: 493, 

May 1914. 

Cephalophus niger Gray. Black Duiker 

Cephalophus niger Gray, Ann. Mag. Nat. Hist. (1) 18: 165, Sep. 1846. Guinea. 
Antilope pluto Temminck, Esquisses Zool. sur la Cote de Guin6, p. 214, 1853. 
Guinea coast, especially near Chama and Dabocrom. 



ALLEN: CHECKLIST OF AFRICAN MAMMALS 487 

Cephalophtjs nigrifrons nigrifrons Gray. Black-fronted Duiker 
Cephalophus nigrifrons Gray, Proc. Zool. Soc. London, p. 598, pi. 46, text-f. 6, 

Aug. 1871. Gaboon. 
Cephalophus aureus Gray, Ann. Mag. Nat. Hist. (4) 12: 42, July 1873. Gaboon. 
Cephalophus emini Noack, Zool. Anzeiger, 27:405, 22 Mar. 1904. Probably 

from west of Lake Victoria, beyond Bukoba, east-central Africa. 
Based on a headless skin; regarded as a synonym of C. nigrifrons by St. Leger 
(Proc. Zool. Soc. London, p. 211. 16 Apl. 1936). 
Cephalophus claudi Thomas and Wroughton, Ann. Mag. Nat. Hist. (7) 19: 

386, May 1907. Bambili, Uele River, Congo Beige. 
A subspecies of C. nigrifrons (fide Lydekker and Blaine), and so accepted by 
Lonnberg (Revue Zool. Africaine, 7: 167, 1919), but by St. Leger made a 
synonym of nigrifrons. 
Cephalophus claudi mixtus Lonnberg, Kungl. Svenska Vet.-Akad. Handl., 

Stockholm, 58: no. 2, p. 89, pi. 10, f. 3, 1 Sep. 1917. Beni, Congo Beige. 
Cephalophus nigrifrons lusumbi Lonnberg, Revue Zool. Africaine, 7: 168, 1 Oct. 

1919. Kinda, Lulua district, western Katanga, Congo Beige. 
Cephalophus nigrifrons apanbanga Lorenz-Liburnau, Ann. Naturhist. Mus. 

Wien, 36: 108, text-f. 6, Oct. 1923. Ukaika, Ituri district, Congo Beige. 
A synonymi of C. nigrifrons according to St. Leger (Proc. Zool. Soc. London, 
p. 211, 16 Apl. 1936). 

Cephalophus nigrifrons fosteri St. Leger. Foster's Black-fronted Duiker 
Cephalophus fosteri St. Leger, Ann. Mag. Nat. Hist. (10) 14: 383, Sep. 1934. 

Mt. Elgon, Kenya Colony, 9300 ft. 
Cephalophus nigrifrons fosteri St. Leger, Proc. Zool. Soc. London, p. 214, 16 

Apl. 1936. 

Cephalophus nigrifrons hooki St. Leger. Hook's Black-fronted Duiker 
Cephalophus hooki St. Leger, Ann. Mag. Nat. Hist. (10) 14: 384, Sep. 1934. 

Southwest face of Mt. Kenya, Kenya Colony, 8500-10,000 ft. 
Cephalophus nigrifrons hooki St. Leger, Proc. Zool. Soc. London, p. 214, 16 Apl. 

1936. 
For Mr. Raymond Hook. 

Cephalophus nigrifrons kivuensis Lonnberg. Kivu Black-fronted Duiker 
Cephalophus nigrifrons kivuensis Lonnberg, Revue Zool. Africaine, 7: 165, 1 
Oct. 1919. Lake Kivu region, XlJongo Beige. 

Cephalophus nigrifrons rubious Thomas 

Cephalophus rubidus Thomas, Proc. Zool. Soc. London, 2: 89, 1 Oct. 1901. 

"Ruwenzori district." 
Cephalophus yiigrifrons rubidus St. Leger, Prop. Zool. Soc. London, p. 213, 16 

Apl. 1936. 



488 bulletin: museum of comparative zoology 

Cephalophus ogilbyi ogilbyi (Waterhoiise). Ogilby's Duiker 

Antilope ogilbyi Waterhouse, Proc. Zool. Soc. London, p. 60, 1838. Island of 

Fernando Po. 
Cephalophus ogilbii Gray, Ann. Mag. Nat. Hist. (1) 18: 165, Sep. 1846. 

Cephalophus ogilbyi brookei Thomas. Brooke's Duiker 
Cephalophtis brookei Thomas, Ann. Mag. Nat. Hist. (7) 11: 290, Mar. 1903. 
Fanti, Gold Coast. 
The "mainland representative of ogilbyi" (Lydekker and Blaine). 

Cephalophus rufilatus rufilatus Gray. Red-flanked Duiker 

Antilope griimnia Desmarest, Nouv. Dictionnaire d'Hist. Nat., ed. 2, 2: 191, 

1816, not of Pallas. Coast of Guinea. 
Cephalophus rufilatus Gray, Ann. Mag. Nat. Hist. (1) 18: 166, Sep. 1846. 

Waterloo Village, Sierra Leone. 
Cephalophus rufilatus cuvieri Fitzinger, Sitzb. K. K. Akad. Wiss., Wien, 59: 

sect. 1, p. 166, 1869; based on Cephalophus rufilatus, var. Gray. "Sierra 

Leone, Ashantee." 

Cephalophus rufilatus rubidior Thomas and Wroughton 
Cephalophus rufilatus rubidior Thomas and Wroughton, Ann. Mag. Nat. Hist. 
(7) 19: 385, May 1907. Upper Uele Valley, Congo Beige. 

Cephalophus silvicultor silvicultor (Afzelius). Yellow-backed Duiker 
Antilope silvicultrix Afzelius, Nova Acta Reg. Soc. Sci., Upsala, 7: 265, pi. 8, 

f. 1, 1815. Sierra Leone, and regions adjacent to the Pongas and Quia 

Rivers. 
Cephalophus punctulatus Gray, Gleanings from the Menagerie at Knowsley 

Hall, 2: 11, pi. 11, f. 1, 1850. Sierra Leone. 
Cephalophus longiceps Gray, Proc. Zool. Soc. London, p. 204, text-f. 1, June 

1865. Gaboon. 
Cephalophus r